391 lines
52 KiB
XML
391 lines
52 KiB
XML
<document id="060285B7098615C597E12C8370D897FD" ID-CLB-Dataset="22253" ID-DOI="10.11646/zootaxa.4772.3.2" ID-GBIF-Dataset="983953d9-e7f1-48e8-b24d-b004ada2d986" ID-ISSN="1175-5326" ID-Zenodo-Dep="3819706" ID-ZooBank="403F169F-DD66-4A64-8887-571D55066E33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589175668640" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2020" docId="038887EDFFFEFFCDFF081CA306BBFDB3" docLanguage="en" docName="zootaxa.4772.3.2.pdf" docOrigin="Zootaxa 4772 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Triacanthoneus Anker 2010" docType="treatment" docVersion="10" lastPageNumber="456" masterDocId="FFB1FF95FFFDFFCAFF9F1A53077EFFA8" masterDocTitle="Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010 with description of a second eastern Pacific species (Malacostraca: Decapoda)" masterLastPageNumber="468" masterPageNumber="450" pageNumber="453" updateTime="1698839100595" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F56C61CA593A618D0FF5817355C8B573">Taxonomic remarks on the alpheid shrimp genus Triacanthoneus Anker, 2010 with description of a second eastern Pacific species (Malacostraca: Decapoda)</mods:title>
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<mods:namePart id="2740A642A44882A791D6820B6C91A26E">Anker, Arthur</mods:namePart>
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<mods:title id="D3E449DC74974B2A648095C7F50943F5">Zootaxa</mods:title>
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<mods:part id="1843E6957B0244B1DA26133A909156E5">
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<mods:date id="0B0591F67A142335765DB219DD5D88E0">2020</mods:date>
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<mods:number id="D3F031E21E3BEE5CA137323FFA88BCBA">2020-05-11</mods:number>
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<mods:number id="EF4A5E2B32C6331BDF3A98241443F488">4772</mods:number>
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<mods:start id="9B8011C0579A3D00F856FF6FA6C5913C">450</mods:start>
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<treatment id="038887EDFFFEFFCDFF081CA306BBFDB3" ID-DOI="http://doi.org/10.5281/zenodo.3843769" ID-GBIF-Taxon="164035963" ID-Zenodo-Dep="3843769" LSID="urn:lsid:plazi:treatment:038887EDFFFEFFCDFF081CA306BBFDB3" httpUri="http://treatment.plazi.org/id/038887EDFFFEFFCDFF081CA306BBFDB3" lastPageId="7" lastPageNumber="456" pageId="3" pageNumber="453">
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<subSubSection id="C33B6570FFFEFFC9FF081CA30546F8A3" box="[151,568,1776,1803]" pageId="3" pageNumber="453" type="nomenclature">
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<paragraph id="8B9E36FBFFFEFFC9FF081CA30546F8A3" blockId="3.[151,568,1776,1803]" box="[151,568,1776,1803]" pageId="3" pageNumber="453">
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<heading id="D0D68197FFFEFFC9FF081CA30546F8A3" bold="true" box="[151,568,1776,1803]" fontSize="11" level="1" pageId="3" pageNumber="453" reason="1">
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<emphasis id="B955EAE9FFFEFFC9FF081CA30546F8A3" bold="true" box="[151,568,1776,1803]" pageId="3" pageNumber="453">
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Genus
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<taxonomicName id="4C214D78FFFEFFC9FF751CA30546F8A3" ID-CoL="85JMG" authority="Anker, 2010" authorityName="Anker" authorityYear="2010" box="[234,568,1776,1803]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="453" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EAE9FFFEFFC9FF751CA306DFF8A3" bold="true" box="[234,417,1776,1803]" italics="true" pageId="3" pageNumber="453">Triacanthoneus</emphasis>
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Anker, 2010
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6570FFFEFFC9FF081D660692F8E5" box="[151,492,1844,1869]" pageId="3" pageNumber="453" type="reference_group">
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<paragraph id="8B9E36FBFFFEFFC9FF081D660692F8E5" blockId="3.[151,492,1844,1869]" box="[151,492,1844,1869]" pageId="3" pageNumber="453">
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<treatmentCitationGroup id="AB3111D5FFFEFFC9FF081D660692F8E5" box="[151,492,1844,1869]" pageId="3" pageNumber="453">
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<treatmentCitation id="0A8010EAFFFEFFC9FF081D660696F8E5" ID-DOI="http://doi.org/10.5281/zenodo.6206923" ID-Zenodo-Dep="6206923" author="Anker, A." box="[151,488,1844,1869]" httpUri="http://treatment.plazi.org/id/039F5848FF97FFEDE5F3FD3FFA5DCA62" page="48" pageId="3" pageNumber="453" year="2010">
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<taxonomicName id="4C214D78FFFEFFC9FF081D660696F8E5" ID-CoL="85JMG" authority="Anker 2010 a: 48" authorityName="Anker" authorityPageNumber="48" authorityYear="2010" box="[151,488,1844,1869]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="453" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EAE9FFFEFFC9FF081D66064BF8E5" box="[151,309,1845,1869]" italics="true" pageId="3" pageNumber="453">Triacanthoneus</emphasis>
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<bibRefCitation id="EFB04B0AFFFEFFC9FEA41D670696F8E5" author="Anker, A." box="[315,488,1844,1869]" pageId="3" pageNumber="453" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a: 48</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6570FFFEFFCEFF581D2F053CFB43" lastPageId="4" lastPageNumber="454" pageId="3" pageNumber="453" type="diagnosis">
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<paragraph id="8B9E36FBFFFEFFCEFF581D2F053CFB43" blockId="3.[151,1437,1915,2014]" lastBlockId="4.[151,1437,153,2015]" lastPageId="4" lastPageNumber="454" pageId="3" pageNumber="453">
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<emphasis id="B955EAE9FFFEFFC9FF581D2F06D1F83E" bold="true" box="[199,431,1915,1942]" pageId="3" pageNumber="453">Emended diagnosis</emphasis>
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. Body slender, not compressed. Dorsal mid-line of carapace with one strong, sharp, anteriorly directed tooth arising at carapace mid-length or more or less posterior/anterior to it; dorsal carina typically present as continuation of rostral carina, rarely absent (only in
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<taxonomicName id="4C214D78FFFEFFC9FCC01D9704A9F875" authorityName="Anker" authorityYear="2010" box="[863,983,1988,2013]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="453" phylum="Arthropoda" rank="species" species="armatus">
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<emphasis id="B955EAE9FFFEFFC9FCC01D9704A9F875" box="[863,983,1988,2013]" italics="true" pageId="3" pageNumber="453">T. armatus</emphasis>
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</taxonomicName>
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). Carapace flanks with or without simi- larly strong, sharp, anteriorly directed teeth (one on each side) in hepatic or post-hepatic position. Anterolateral margin of carapace with or without sharp teeth arising directly from postorbital margin or slightly posterior to it (possibly homologous to orbital teeth). Rostrum well-developed, distally acute, ventral margin with small sharp tooth typically in subdistal position, rarely near rostral mid-length (only in
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<taxonomicName id="4C214D78FFF9FFCEFC711B5503EBFEB7" authorityName="Alvarez, Iliffe, Gonzalez & Villalobos" authorityYear="2012" box="[1006,1173,261,287]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="akumalensis">
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<emphasis id="B955EAE9FFF9FFCEFC711B5503EBFEB7" box="[1006,1173,261,287]" italics="true" pageId="4" pageNumber="454">T. akumalensis</emphasis>
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</taxonomicName>
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). Pterygostomial angle not produced anteriorly. Cardiac notch variously developed, either deep or reduced to small incision. First to fourth pleonites posteroventrally rounded; fifth pleonite posteroventrally angular or more or less projecting; sixth pleonite without articulated plate. Telson typically slender, with two pairs of cuspidate setae on dorsal surface; posterior margin straight or with variously developed median incision, with two pairs of stout spiniform setae; anal tubercles absent. Eyes typically partly or largely exposed in dorsal view, largely exposed in lateral view, rarely completely concealed in dorsal view and partly visible in lateral view. Antennular peduncle with small sharp tooth on mesioventral carina of first article; stylocerite well developed, with acute tip reaching well beyond distal margin of first article, sometimes with thin dorsal longitudinal carina terminating anteriorly in acute or subacute, dorsally pointing tooth; lateral flagellum with well-developed accessory ramus. Antenna with well-developed, ovate scaphocerite, blade broad and reaching beyond distolateral tooth; carpocerite short, reaching at most half-length of scaphocerite. Mouthparts typical for family; mandible with palp subdivided into two articles; maxillular palp with both lobes furnished with setae. Third maxilliped with lateral plate rounded or subacutely produced; ultimate article tapering into corneous tip, without robust spiniform setae. Chelipeds strongly unequal and asymmetrical, carried flexed when not in use. Major cheliped elongate, slender; ischium with two or three cuspidate setae; merus flattened ventrally; carpus subcylindrical, slender, widening distally; chela smooth, finger cutting edges typically serrated with small subtriangular teeth, rarely (only in
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<taxonomicName id="4C214D78FFF9FFCEFDB8191505B0FCF7" authorityName="Alvarez, Iliffe, Gonzalez & Villalobos" authorityYear="2012" box="[551,718,837,863]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="akumalensis">
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<emphasis id="B955EAE9FFF9FFCEFDB8191505B0FCF7" box="[551,718,837,863]" italics="true" pageId="4" pageNumber="454">T. akumalensis</emphasis>
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</taxonomicName>
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) with raptorial teeth intercalated between smaller teeth. Minor cheliped slender, slightly elongate, much shorter than major cheliped; ischium with two or three cuspidate setae; chela simple, finger cutting edges unarmed. Second pereiopod slender; ischium armed with one to three cuspidate seta(e); carpus with five subdivisions, first always longest. Third and fourth pereiopods slender; ischium with two or three cuspidate setae; merus and carpus unarmed; propodus with small slender spiniform setae; dactylus simple, conical, slender. Fifth pereiopod with ischium unarmed; propodal brush either well developed or reduced. Second pleopod with appendix masculina and appendix interna in all specimens, including ovigerous ones. Uropod with slender exopod and endopod; exopod with small distolateral tooth and slender distolateral spiniform seta, diaeresis present, more or less sinuous. Gill/exopod formula: 5 pleurobranchs (above P1–5) [with spiny process above P
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<taxonomicName id="4C214D78FFF9FFCEFF2A1ED90625FB0B" authorityName="Alvarez, Iliffe, Gonzalez & Villalobos" authorityYear="2012" box="[181,347,1161,1187]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="akumalensis">
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<emphasis id="B955EAE9FFF9FFCEFF2A1ED90625FB0B" box="[181,347,1161,1187]" italics="true" pageId="4" pageNumber="454">T. akumalensis</emphasis>
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</taxonomicName>
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]; 1 arthrobranch (at Mxp3); 0–1 podobranchs (at Mxp2) [typically 0, but with 1 well-developed podobranch in
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<taxonomicName id="4C214D78FFF9FFCEFEDE1EFD0699FB6F" authorityName="Alvarez, Iliffe, Gonzalez & Villalobos" authorityYear="2012" box="[321,487,1197,1223]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="akumalensis">
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<emphasis id="B955EAE9FFF9FFCEFEDE1EFD0699FB6F" box="[321,487,1197,1223]" italics="true" pageId="4" pageNumber="454">T. akumalensis</emphasis>
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</taxonomicName>
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]; 2 lobe-shaped epipods (Mxp1–2); 5 mastigobranchs (Mxp3, P1–4), 5 sets of setobranchs (P1–5); 3 exopods (Mxp1–3).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6570FFF9FFCEFF581EA605C6FAD4" pageId="4" pageNumber="454" type="type_taxon">
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<paragraph id="8B9E36FBFFF9FFCEFF581EA603E1FAA7" blockId="4.[151,1437,153,2015]" box="[199,1183,1268,1295]" pageId="4" pageNumber="454">
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<emphasis id="B955EAE9FFF9FFCEFF581EA60625FAA7" bold="true" box="[199,347,1268,1295]" pageId="4" pageNumber="454">
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<typeStatus id="549A8859FFF9FFCEFF581EA6067CFAA7" box="[199,258,1269,1295]" pageId="4" pageNumber="454">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="4C214D78FFF9FFCEFEF61EA605A1FAA7" ID-CoL="85ZFV" authority="Anker, 2010" authorityName="in Anker" authorityYear="2010" box="[361,735,1269,1295]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="toro">
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<emphasis id="B955EAE9FFF9FFCEFEF61EA60532FAA7" box="[361,588,1269,1295]" italics="true" pageId="4" pageNumber="454">Triacanthoneus toro</emphasis>
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Anker, 2010
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</taxonomicName>
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, by original designation (
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<bibRefCitation id="EFB04B0AFFF9FFCEFC641EA603EAFAA7" author="Anker, A." box="[1019,1172,1269,1295]" pageId="4" pageNumber="454" pagination="47 - 63" refId="ref10850" refString="Anker, A. (2010 a) A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts. Zootaxa, 2652 (1), 47 - 63. https: // doi. org / 10.11646 / zootaxa. 2652.1.4" type="journal article" year="2010">Anker 2010a</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8B9E36FBFFF9FFCEFF581F4B05C6FAD4" blockId="4.[151,1437,153,2015]" pageId="4" pageNumber="454">
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<emphasis id="B955EAE9FFF9FFCEFF581F4B06A8FA9B" bold="true" box="[199,470,1304,1331]" pageId="4" pageNumber="454">Other species included</emphasis>
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.
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<taxonomicName id="4C214D78FFF9FFCEFE7B1F4A04E3FA9B" ID-CoL="85ZFR" authority="Anker, 2010" authorityName="Anker" authorityYear="2010" box="[484,925,1305,1331]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="alacranes">
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<emphasis id="B955EAE9FFF9FFCEFE7B1F4A0479FA9B" box="[484,775,1305,1331]" italics="true" pageId="4" pageNumber="454">Triacanthoneus alacranes</emphasis>
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Anker, 2010
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</taxonomicName>
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;
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<taxonomicName id="4C214D78FFF9FFCEFC371F4903C3FA9B" ID-CoL="85ZFT" authority="Anker, 2010" authorityName="Anker" authorityYear="2010" box="[936,1213,1305,1331]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="pacificus">
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<emphasis id="B955EAE9FFF9FFCEFC371F490356FA9B" box="[936,1064,1305,1331]" italics="true" pageId="4" pageNumber="454">T. pacificus</emphasis>
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Anker, 2010
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</taxonomicName>
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;
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<taxonomicName id="4C214D78FFF9FFCEFB571F4907A5FAFF" ID-CoL="85XZP" authority="(Anker, 2010)" authorityName="Anker" authorityYear="2020" baseAuthorityName="Anker" baseAuthorityYear="2010" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="armatus" status="comb. nov.">
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<emphasis id="B955EAE9FFF9FFCEFB571F49023EFA9B" box="[1224,1344,1306,1331]" italics="true" pageId="4" pageNumber="454">T. armatus</emphasis>
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(Anker, 2010)
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</taxonomicName>
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,
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<emphasis id="B955EAE9FFF9FFCEFF771F6E0614FAFF" bold="true" box="[232,362,1341,1367]" pageId="4" pageNumber="454">
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<taxonomicNameLabel id="A2665792FFF9FFCEFF771F6E0614FAFF" box="[232,362,1341,1367]" pageId="4" pageNumber="454" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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;
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<taxonomicName id="4C214D78FFF9FFCEFEE51F6D0359FAFF" ID-CoL="85ZFQ" authority="Alvarez, Iliffe, Gonzalez & Villalobos, 2012" authorityName="Alvarez, Iliffe, Gonzalez & Villalobos" authorityYear="2012" box="[378,1063,1341,1367]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="akumalensis">
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<emphasis id="B955EAE9FFF9FFCEFEE51F6D055FFAFF" box="[378,545,1341,1367]" italics="true" pageId="4" pageNumber="454">T. akumalensis</emphasis>
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Alvarez, Iliffe, Gonzalez & Villalobos, 2012
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</taxonomicName>
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;
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<taxonomicName id="4C214D78FFF9FFCEFBAC1F6D062AFAD3" ID-CoL="85ZFS" authority="Alvarez, Iliffe & Villalobos, 2014" authorityName="Alvarez, Iliffe & Villalobos" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="chapelianus">
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<emphasis id="B955EAE9FFF9FFCEFBAC1F6D03A9FAFF" box="[1075,1239,1341,1367]" italics="true" pageId="4" pageNumber="454">T. chapelianus</emphasis>
|
||
<bibRefCitation id="EFB04B0AFFF9FFCEFB411F6E062AFAD3" author="Alvarez, F. & Iliffe, T. M. & Villalobos, J. L." pageId="4" pageNumber="454" pagination="88 - 94" refId="ref10678" refString="Alvarez, F., Iliffe, T. M. & Villalobos, J. L. (2014) A new species of the alpheid shrimp genus Triacanthoneus Anker, 2010 (Crustacea: Alpheidae) from Belize. Zootaxa, 3768 (1), 88 - 94. https: // doi. org / 10.11646 / zootaxa. 3768.1.6" type="journal article" year="2014">Alvarez, Iliffe & Villalobos, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C214D78FFF9FFCEFEC01F3106B8FAD3" ID-CoL="874KG" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[351,454,1377,1403]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="blanca" status="sp. nov.">
|
||
<emphasis id="B955EAE9FFF9FFCEFEC01F3106B8FAD3" box="[351,454,1377,1403]" italics="true" pageId="4" pageNumber="454">T. blanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B955EAE9FFF9FFCEFE511F320559FAD3" bold="true" box="[462,551,1377,1403]" pageId="4" pageNumber="454">
|
||
<taxonomicNameLabel id="A2665792FFF9FFCEFE511F320559FAD3" box="[462,551,1377,1403]" pageId="4" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33B6570FFF9FFCEFF581FD7057AFA6C" pageId="4" pageNumber="454" type="distribution">
|
||
<paragraph id="8B9E36FBFFF9FFCEFF581FD7057AFA6C" blockId="4.[151,1437,153,2015]" pageId="4" pageNumber="454">
|
||
<emphasis id="B955EAE9FFF9FFCEFF581FD70627FA37" bold="true" box="[199,345,1412,1439]" pageId="4" pageNumber="454">Distribution</emphasis>
|
||
. Western Atlantic (southern Gulf of Mexico and western and central Caribbean Sea) and eastern Pacific (
|
||
<collectingCountry id="F336766BFFF9FFCEFF6C1FFA0630FA6B" box="[243,334,1449,1475]" name="Panama" pageId="4" pageNumber="454">Panama</collectingCountry>
|
||
and
|
||
<collectingCountry id="F336766BFFF9FFCEFE1B1FFA0686FA6C" box="[388,504,1449,1476]" name="Colombia" pageId="4" pageNumber="454">Colombia</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33B6570FFF9FFCEFF581F9E03EDF9FB" pageId="4" pageNumber="454" type="biology_ecology">
|
||
<paragraph id="8B9E36FBFFF9FFCEFF581F9E03EDF9FB" blockId="4.[151,1437,153,2015]" pageId="4" pageNumber="454">
|
||
<emphasis id="B955EAE9FFF9FFCEFF581F9E0658FA4F" bold="true" box="[199,294,1485,1511]" pageId="4" pageNumber="454">Ecology</emphasis>
|
||
.
|
||
<materialsCitation id="3B493CA6FFF9FFCEFEAB1F9E04E8F987" ID-GBIF-Occurrence="2611567311" elevation="8" location="Western Atlantic" municipality="Coral" pageId="4" pageNumber="454" specimenCount="1">
|
||
<location id="8EFE6020FFF9FFCEFEAB1F9E068AFA4F" LSID="urn:lsid:plazi:treatment:038887EDFFFEFFCDFF081CA306BBFDB3:8EFE6020FFF9FFCEFEAB1F9E068AFA4F" box="[308,500,1485,1511]" municipality="Coral" name="Western Atlantic" pageId="4" pageNumber="454">Western Atlantic</location>
|
||
:
|
||
<collectingMunicipality id="6BFAAC81FFF9FFCEFE611F9E0542FA4F" box="[510,572,1485,1511]" pageId="4" pageNumber="454">Coral</collectingMunicipality>
|
||
reefs and associated rubble-rich habitats; sand flats with abundant coral rubble and sea grass, often close to mangroves; two taxa recorded from anchialine and marine caves, although only one of them appears to be restricted to caves; depth range:
|
||
<quantity id="4CD99B1EFFF9FFCEFCBA1C4704ECF987" box="[805,914,1556,1583]" metricMagnitude="0" metricUnit="m" metricValue="7.75" metricValueMax="15.0" metricValueMin="0.5" pageId="4" pageNumber="454" unit="m" value="7.75" valueMax="15.0" valueMin="0.5">
|
||
<elevation id="000CD1C8FFF9FFCEFCBA1C4704ECF987" box="[805,914,1556,1583]" metricMagnitude="0" metricUnit="m" metricValue="7.75" metricValueMax="15.0" metricValueMin="0.5" pageId="4" pageNumber="454" unit="m" value="7.75" valueMax="15.0" valueMin="0.5">0.5–15 m</elevation>
|
||
</quantity>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B493CA6FFF9FFCEFC031C4603EDF9FB" ID-GBIF-Occurrence="2611567310" elevation="4" location="Eastern Pacific" pageId="4" pageNumber="454" specimenCount="1">
|
||
<location id="8EFE6020FFF9FFCEFC031C460337F987" LSID="urn:lsid:plazi:treatment:038887EDFFFEFFCDFF081CA306BBFDB3:8EFE6020FFF9FFCEFC031C460337F987" box="[924,1097,1557,1583]" name="Eastern Pacific" pageId="4" pageNumber="454">Eastern Pacific</location>
|
||
: intertidal mud-sand flats with abundant rocks, subtidal sand-rubble flats with rocks and coral rubble; depth range:
|
||
<quantity id="4CD99B1EFFF9FFCEFBDA1C6A03F1F9FB" box="[1093,1167,1593,1619]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="4" pageNumber="454" unit="m" value="4.0" valueMax="8.0" valueMin="0.0">
|
||
<elevation id="000CD1C8FFF9FFCEFBDA1C6A03F1F9FB" box="[1093,1167,1593,1619]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="4" pageNumber="454" unit="m" value="4.0" valueMax="8.0" valueMin="0.0">0–8 m</elevation>
|
||
</quantity>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33B6570FFF9FFCDFF581C0E06BBFDB3" lastPageId="7" lastPageNumber="457" pageId="4" pageNumber="454" type="discussion">
|
||
<paragraph id="8B9E36FBFFF9FFCFFF581C0E023AFF7F" blockId="4.[151,1437,153,2015]" lastBlockId="5.[151,1437,153,611]" lastPageId="5" lastPageNumber="455" pageId="4" pageNumber="454">
|
||
<emphasis id="B955EAE9FFF9FFCEFF581C0E064AF9DF" bold="true" box="[199,308,1629,1655]" pageId="4" pageNumber="454">Remarks</emphasis>
|
||
. The inclusion of
|
||
<taxonomicName id="4C214D78FFF9FFCEFD991C0D05D4F9DF" authorityName="Alvarez, Iliffe & Villalobos" authorityYear="2014" box="[518,682,1629,1655]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="chapelianus">
|
||
<emphasis id="B955EAE9FFF9FFCEFD991C0D05D4F9DF" box="[518,682,1629,1655]" italics="true" pageId="4" pageNumber="454">T. chapelianus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C214D78FFF9FFCEFD4D1C0E04FEF9DF" authorityName="Anker" authorityYear="2010" box="[722,896,1629,1655]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF9FFCEFD4D1C0E04FEF9DF" box="[722,896,1629,1655]" italics="true" pageId="4" pageNumber="454">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
by Alvarez
|
||
<emphasis id="B955EAE9FFF9FFCEFB921C0D0338F9DF" box="[1037,1094,1629,1655]" italics="true" pageId="4" pageNumber="454">et al.</emphasis>
|
||
(2014), followed by the present inclusion of
|
||
<taxonomicName id="4C214D78FFF9FFCEFED01CD106B6F933" authorityName="Anker" authorityYear="2020" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[335,456,1666,1691]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis id="B955EAE9FFF9FFCEFED01CD106B6F933" box="[335,456,1666,1691]" italics="true" pageId="4" pageNumber="454">T. armatus</emphasis>
|
||
</taxonomicName>
|
||
, result in a certain heterogeneity in the genus, especially in the presence or absence of the strong, sharp, anteriorly directed teeth on the hepatic or post-hepatic area of the carapace. Based on this feature alone,
|
||
<taxonomicName id="4C214D78FFF9FFCEFF7C1C9A06EFF94B" authorityName="Anker" authorityYear="2010" box="[227,401,1737,1763]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF9FFCEFF7C1C9A06EFF94B" box="[227,401,1737,1763]" italics="true" pageId="4" pageNumber="454">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
can be subdivided into two distinct species groups. The first group includes all species, in which these teeth arise from the hepatic or post-hepatic area of the carapace, viz.
|
||
<taxonomicName id="4C214D78FFF9FFCEFB811CBD0319F8AF" authorityName="in Anker" authorityYear="2010" box="[1054,1127,1774,1799]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="toro">
|
||
<emphasis id="B955EAE9FFF9FFCEFB811CBD0319F8AF" box="[1054,1127,1774,1799]" italics="true" pageId="4" pageNumber="454">T. toro</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D78FFF9FFCEFBEB1CBD0383F8AF" authorityName="Anker" authorityYear="2010" box="[1140,1277,1773,1799]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFF9FFCEFBEB1CBD0383F8AF" box="[1140,1277,1773,1799]" italics="true" pageId="4" pageNumber="454">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D78FFF9FFCEFA941CBD07CBF883" authorityName="Alvarez, Iliffe, Gonzalez & Villalobos" authorityYear="2012" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="akumalensis">
|
||
<emphasis id="B955EAE9FFF9FFCEFA941CBD07CBF883" italics="true" pageId="4" pageNumber="454">T. akumalensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D78FFF9FFCEFF5C1D41063BF883" authorityName="Anker" authorityYear="2010" box="[195,325,1809,1835]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="pacificus">
|
||
<emphasis id="B955EAE9FFF9FFCEFF5C1D41063BF883" box="[195,325,1809,1835]" italics="true" pageId="4" pageNumber="454">T. pacificus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D78FFF9FFCEFEE21D41069BF883" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[381,485,1809,1835]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="blanca" status="sp. nov.">
|
||
<emphasis id="B955EAE9FFF9FFCEFEE21D41069BF883" box="[381,485,1809,1835]" italics="true" pageId="4" pageNumber="454">T. blanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B955EAE9FFF9FFCEFE721D420539F883" bold="true" box="[493,583,1809,1835]" pageId="4" pageNumber="454">
|
||
<taxonomicNameLabel id="A2665792FFF9FFCEFE721D420539F883" box="[493,583,1809,1835]" pageId="4" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see below). The second group includes two species, viz.
|
||
<taxonomicName id="4C214D78FFF9FFCEFB471D41022EF883" authorityName="Anker" authorityYear="2020" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[1240,1360,1810,1835]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis id="B955EAE9FFF9FFCEFB471D41022EF883" box="[1240,1360,1810,1835]" italics="true" pageId="4" pageNumber="454">T. armatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D78FFF9FFCEFA171D410660F8E7" authorityName="Alvarez, Iliffe & Villalobos" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="chapelianus">
|
||
<emphasis id="B955EAE9FFF9FFCEFA171D410660F8E7" italics="true" pageId="4" pageNumber="454">T. chapelianus</emphasis>
|
||
</taxonomicName>
|
||
, which do not have strong teeth on the hepatic or post-hepatic area of the carapace, but have smaller sharp teeth arising either directly from the anterolateral margin of the carapace (in the former species) or just posterior to it (in the latter species), i.e., in post-orbital position. The homology of these teeth with the orbital teeth of other alpheid shrimp genera, including
|
||
<taxonomicName id="4C214D78FFF9FFCEFDC71DF205AFF813" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[600,721,1953,1979]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF9FFCEFDC71DF205AFF813" box="[600,721,1953,1979]" italics="true" pageId="4" pageNumber="454">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
, appears to be more likely than the hypothesis of their hepatic origin and posterior “migration” towards the post-orbital area (Alvarez
|
||
<emphasis id="B955EAE9FFF9FFCEFC511D950376F877" box="[974,1032,1989,2015]" italics="true" pageId="4" pageNumber="454">et al.</emphasis>
|
||
2014), especially in
|
||
<taxonomicName id="4C214D78FFF9FFCEFB631D950208F877" authorityName="Anker" authorityYear="2020" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[1276,1398,1990,2015]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="454" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis id="B955EAE9FFF9FFCEFB631D950208F877" box="[1276,1398,1990,2015]" italics="true" pageId="4" pageNumber="454">T. armatus</emphasis>
|
||
</taxonomicName>
|
||
, in which they truly arise from the anterolateral margin of the carapace. However, only a detailed study of the presently unknown larvae and post-larvae, combined with Hox genes analyses, could shed more light of this subject.
|
||
</paragraph>
|
||
<paragraph id="8B9E36FBFFF8FFCFFF581AB206C2FDCB" blockId="5.[151,1437,153,611]" pageId="5" pageNumber="455">
|
||
At least two evolutionary scenarios are possible to explain the general similarity of
|
||
<taxonomicName id="4C214D78FFF8FFCFFB1E1AB10384FF53" authorityName="Anker" authorityYear="2020" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[1153,1274,226,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis id="B955EAE9FFF8FFCFFB1E1AB10384FF53" box="[1153,1274,226,251]" italics="true" pageId="5" pageNumber="455">T. armatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D78FFF8FFCFFAAC1AB1067CFEB7" authority=", In" authorityName="In" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="species" species="chapelianus">
|
||
<emphasis id="B955EAE9FFF8FFCFFAAC1AB107A5FEB7" italics="true" pageId="5" pageNumber="455">T. chapelianus</emphasis>
|
||
, In
|
||
</taxonomicName>
|
||
the first scenario, their great resemblance could be result of a convergent evolution, i.e. these two taxa may not be related at all or only distantly so (in which case, the herein proposed species transfer would obviously be unnecessary). In the second scenario,
|
||
<taxonomicName id="4C214D78FFF8FFCFFDF31B1D0598FECF" authorityName="Anker" authorityYear="2020" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[620,742,334,359]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis id="B955EAE9FFF8FFCFFDF31B1D0598FECF" box="[620,742,334,359]" italics="true" pageId="5" pageNumber="455">T. armatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D78FFF8FFCFFCBF1B1D04BAFECF" box="[800,964,333,359]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="species" species="chapelianus">
|
||
<emphasis id="B955EAE9FFF8FFCFFCBF1B1D04BAFECF" box="[800,964,333,359]" italics="true" pageId="5" pageNumber="455">T. chapelianus</emphasis>
|
||
</taxonomicName>
|
||
could form a small clade on its own, for instance, in an intermediate position between
|
||
<taxonomicName id="4C214D78FFF8FFCFFD041B22046AFE23" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[667,788,369,395]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFD041B22046AFE23" box="[667,788,369,395]" italics="true" pageId="5" pageNumber="455">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
and the other species of
|
||
<taxonomicName id="4C214D78FFF8FFCFFBB21B2203A5FE23" authorityName="Anker" authorityYear="2010" box="[1069,1243,369,395]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFBB21B2203A5FE23" box="[1069,1243,369,395]" italics="true" pageId="5" pageNumber="455">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
, or nested somewhere else within
|
||
<taxonomicName id="4C214D78FFF8FFCFFEF61BC6069CFE07" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[361,482,405,431]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFEF61BC6069CFE07" box="[361,482,405,431]" italics="true" pageId="5" pageNumber="455">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
. The latter genus is becoming large and increasingly complex morphologically and may well paraphyletic in respect to at least some of the allied genera (including
|
||
<taxonomicName id="4C214D78FFF8FFCFFBA91BEA039AFE7B" authorityName="Anker" authorityYear="2010" box="[1078,1252,441,467]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFBA91BEA039AFE7B" box="[1078,1252,441,467]" italics="true" pageId="5" pageNumber="455">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
). However, this topic cannot be discussed further without a comprehensive morphological and molecular analysis of several alpheid genera, focusing specifically on
|
||
<taxonomicName id="4C214D78FFF8FFCFFE6118520509FDB3" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[510,631,513,539]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFE6118520509FDB3" box="[510,631,513,539]" italics="true" pageId="5" pageNumber="455">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D78FFF8FFCFFD1B1852044CFDB3" authorityName="Anker" authorityYear="2010" box="[644,818,513,539]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFD1B1852044CFDB3" box="[644,818,513,539]" italics="true" pageId="5" pageNumber="455">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D78FFF8FFCFFCDF185104D4FDB3" authorityName="Dworschak, Anker & Abed-Navandi" authorityYear="2000" box="[832,938,514,539]" class="Malacostraca" family="Alpheidae" genus="Deioneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFCDF185104D4FDB3" box="[832,938,514,539]" italics="true" pageId="5" pageNumber="455">Deioneus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D78FFF8FFCFFC4118520320FDB3" authorityName="Komai & Fujita" authorityYear="2018" box="[990,1118,513,539]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFC4118520320FDB3" box="[990,1118,513,539]" italics="true" pageId="5" pageNumber="455">Caligoneus</emphasis>
|
||
</taxonomicName>
|
||
. For the time being, it seems most reasonable and taxonomically convenient to retain
|
||
<taxonomicName id="4C214D78FFF8FFCFFC8A187604BDFD97" authorityName="Anker" authorityYear="2010" box="[789,963,549,575]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFF8FFCFFC8A187604BDFD97" box="[789,963,549,575]" italics="true" pageId="5" pageNumber="455">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
as a valid genus with a new composition, awaiting such an analysis.
|
||
</paragraph>
|
||
<caption id="DF5E6673FFF8FFCFFF081D09022BF877" ID-DOI="http://doi.org/10.5281/zenodo.3819712" ID-Zenodo-Dep="3819712" httpUri="https://zenodo.org/record/3819712/files/figure.png" pageId="5" pageNumber="455" startId="5.[151,250,1882,1907]" targetBox="[160,1428,643,1851]" targetPageId="5">
|
||
<paragraph id="8B9E36FBFFF8FFCFFF081D09022BF877" blockId="5.[151,1437,1882,2015]" pageId="5" pageNumber="455">
|
||
<emphasis id="B955EAE9FFF8FFCFFF081D090672F8DB" bold="true" box="[151,268,1882,1907]" pageId="5" pageNumber="455">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C214D78FFF8FFCFFE871D08057EF8DB" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[280,512,1883,1907]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="455" phylum="Arthropoda" rank="species" species="blanca" status="sp. nov.">
|
||
<emphasis id="B955EAE9FFF8FFCFFE871D08057EF8DB" box="[280,512,1883,1907]" italics="true" pageId="5" pageNumber="455">Triacanthoneus blanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B955EAE9FFF8FFCFFD9A1D08052BF8DB" bold="true" box="[517,597,1883,1907]" pageId="5" pageNumber="455">
|
||
<taxonomicNameLabel id="A2665792FFF8FFCFFD9A1D08052BF8DB" box="[517,597,1883,1907]" pageId="5" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, non-ovigerous specimen (cl 3.8 mm) from Isla Coiba, Panama (FLMNH UF 51919): A—second pereiopod, lateral view; B—same, chela, mesial view; C—third pereiopod, lateral view; D—same, distal portion of propodus and dactylus, mesial view; E—fourth pereiopod, lateral view; F—same, distal portion of propodus and dactylus, mesial view; G—fifth pereiopod, lateral view; H—same, distal portion of propodus and dactylus, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5E6673FFFBFFCCFF081CB00364F888" ID-DOI="http://doi.org/10.5281/zenodo.3819716" ID-Zenodo-Dep="3819716" httpUri="https://zenodo.org/record/3819716/files/figure.png" pageId="6" pageNumber="456" startId="6.[151,250,1763,1788]" targetBox="[161,1425,181,1738]" targetPageId="6">
|
||
<paragraph id="8B9E36FBFFFBFFCCFF081CB00364F888" blockId="6.[151,1436,1763,1824]" pageId="6" pageNumber="456">
|
||
<emphasis id="B955EAE9FFFBFFCCFF081CB00672F954" bold="true" box="[151,268,1763,1788]" pageId="6" pageNumber="456">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C214D78FFFBFFCCFE871CB7057EF954" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[280,512,1764,1788]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="456" phylum="Arthropoda" rank="species" species="blanca" status="sp. nov.">
|
||
<emphasis id="B955EAE9FFFBFFCCFE871CB7057EF954" box="[280,512,1764,1788]" italics="true" pageId="6" pageNumber="456">Triacanthoneus blanca</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B955EAE9FFFBFFCCFD9A1CB7052BF954" bold="true" box="[517,597,1764,1788]" pageId="6" pageNumber="456">
|
||
<taxonomicNameLabel id="A2665792FFFBFFCCFD9A1CB7052BF954" box="[517,597,1764,1788]" pageId="6" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, non-ovigerous specimen (cl 3.8 mm) from Isla Coiba, Panama (FLMNH UF 51919): living shrimp in dorsal [A] and lateral [B] views. Photographs by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9E36FBFFFBFFCDFF581D2106BBFDB3" blockId="6.[151,1436,1906,2041]" lastBlockId="7.[151,1437,153,539]" lastPageId="7" lastPageNumber="457" pageId="6" pageNumber="456">
|
||
All members of
|
||
<taxonomicName id="4C214D78FFFBFFCCFE191D21054AF824" authorityName="Anker" authorityYear="2010" box="[390,564,1906,1932]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="456" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFBFFCCFE191D21054AF824" box="[390,564,1906,1932]" italics="true" pageId="6" pageNumber="456">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
, as redefined above, are morphologically distinct from all species currently assigned to
|
||
<taxonomicName id="4C214D78FFFBFFCCFE831DC506EBF818" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[284,405,1942,1968]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="456" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFBFFCCFE831DC506EBF818" box="[284,405,1942,1968]" italics="true" pageId="6" pageNumber="456">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a strong sharp tooth on the dorsal mid-line of the carapace, situated well posterior to the rostral base (ranging from 0.3 length of the carapace in
|
||
<taxonomicName id="4C214D78FFFBFFCCFC271DE8034EF87C" authorityName="Anker" authorityYear="2020" baseAuthorityName="Anker" baseAuthorityYear="2010" box="[952,1072,1979,2004]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="456" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis id="B955EAE9FFFBFFCCFC271DE8034EF87C" box="[952,1072,1979,2004]" italics="true" pageId="6" pageNumber="456">T. armatus</emphasis>
|
||
</taxonomicName>
|
||
to almost 0.7 length of the carapace in
|
||
<taxonomicName id="4C214D78FFFBFFCCFF6E1D8C0602F850" authorityName="Anker" authorityYear="2010" box="[241,380,2014,2040]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="456" phylum="Arthropoda" rank="species" species="alacranes">
|
||
<emphasis id="B955EAE9FFFBFFCCFF6E1D8C0602F850" box="[241,380,2014,2040]" italics="true" pageId="6" pageNumber="456">T. alacranes</emphasis>
|
||
</taxonomicName>
|
||
). However, it must be noted that in the presently monotypic
|
||
<taxonomicName id="4C214D78FFFBFFCCFBAA1D8D03CBF850" authorityName="Komai & Fujita" authorityYear="2018" box="[1077,1205,2014,2040]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="456" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFBFFCCFBAA1D8D03CBF850" box="[1077,1205,2014,2040]" italics="true" pageId="6" pageNumber="456">Caligoneus</emphasis>
|
||
</taxonomicName>
|
||
, one strong tooth or two strong teeth is/are present on the carapace mid-line, adjacent to the rostral base (Komai & Fujita 2018: 2A–C), i.e. in a rather different position (much more anterior and also more elevated relative to the rostrum) from the more posterior mid-dorsal tooth in the species of
|
||
<taxonomicName id="4C214D78FFFAFFCDFD1C1AB2044FFF53" authorityName="Anker" authorityYear="2010" box="[643,817,225,251]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="457" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFAFFCDFD1C1AB2044FFF53" box="[643,817,225,251]" italics="true" pageId="7" pageNumber="457">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
. Additionally,
|
||
<taxonomicName id="4C214D78FFFAFFCDFC461AB20327FF53" authorityName="Komai & Fujita" authorityYear="2018" box="[985,1113,225,251]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="457" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFAFFCDFC461AB20327FF53" box="[985,1113,225,251]" italics="true" pageId="7" pageNumber="457">Caligoneus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C214D78FFFAFFCDFB711AB202E2FF53" authorityName="Anker" authorityYear="2010" box="[1262,1436,225,251]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="457" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFAFFCDFB711AB202E2FF53" box="[1262,1436,225,251]" italics="true" pageId="7" pageNumber="457">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
in the much deeper cleft on the posterior margin of the telson and in the general shape of the chelipeds (Komai & Fujita 2018: figs. 2D, 4C–E). Therefore, these two genera seem to be more distantly related, in addition to be geographically widely separated. Interestingly, within the non-closely related alpheid genus
|
||
<taxonomicName id="4C214D78FFFAFFCDFB3F1B1E07ACFE23" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Parabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="457" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFAFFCDFB3F1B1E0250FECF" box="[1184,1326,333,359]" italics="true" pageId="7" pageNumber="457">Parabetaeus</emphasis>
|
||
<bibRefCitation id="EFB04B0AFFFAFFCDFAAA1B1E07ACFE23" author="Coutiere, H." pageId="7" pageNumber="457" pagination="380 - 386" refId="ref11301" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
, one recently described species (
|
||
<taxonomicName id="4C214D78FFFAFFCDFDD91B210422FE23" authority="Anker, 2015" authorityName="Anker" authorityYear="2015" box="[582,860,368,395]" class="Malacostraca" family="Alpheidae" genus="Parabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="457" phylum="Arthropoda" rank="species" species="acanthus">
|
||
<emphasis id="B955EAE9FFFAFFCDFDD91B2105B6FE23" box="[582,712,369,395]" italics="true" pageId="7" pageNumber="457">P. acanthus</emphasis>
|
||
<bibRefCitation id="EFB04B0AFFFAFFCDFD511B220422FE23" author="Anker, A." box="[718,860,368,395]" pageId="7" pageNumber="457" pagination="585 - 595" refId="ref10962" refString="Anker, A. (2015) Description of a new distinctive species of Parabetaeus Coutiere, 1897 (Decapoda: Caridea: Alpheidae) from the Indo-West Pacific. Zootaxa, 3957 (5), 585 - 595. https: // doi. org / 10.11646 / zootaxa. 3957.5.7" type="journal article" year="2015">Anker, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
) also has a sharp, anteriorly directed tooth in postrostral position, in contrast to the other two species of the genus, in which the post-rostral area is unarmed (
|
||
<bibRefCitation id="EFB04B0AFFFAFFCDFAA01BC60646FE7B" author="Nomura, K. & Anker, A." pageId="7" pageNumber="457" pagination="43 - 54" refId="ref11571" refString="Nomura, K. & Anker, A. (2001) Redescription of Parabetaeus culliereti Coutiere and status of the genus Neoalpheopsis Banner (Decapoda: Caridea: Alpheidae). Crustacean Research, 30, 43 - 54. https: // doi. org / 10.18353 / crustacea. 30.0 _ 43" type="journal article" year="2001">Nomura & Anker 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB04B0AFFFAFFCDFEDE1BEA06B4FE7B" author="Anker, A." box="[321,458,440,467]" pageId="7" pageNumber="457" pagination="585 - 595" refId="ref10962" refString="Anker, A. (2015) Description of a new distinctive species of Parabetaeus Coutiere, 1897 (Decapoda: Caridea: Alpheidae) from the Indo-West Pacific. Zootaxa, 3957 (5), 585 - 595. https: // doi. org / 10.11646 / zootaxa. 3957.5.7" type="journal article" year="2015">Anker 2015</bibRefCitation>
|
||
). This example shows that the armature in the form of a strong, sharp, anteriorly directed tooth on the mid-dorsal line of the carapace is not limited to
|
||
<taxonomicName id="4C214D78FFFAFFCDFCD41B8E0487FE5F" authorityName="Anker" authorityYear="2010" box="[843,1017,477,503]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="457" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFAFFCDFCD41B8E0487FE5F" box="[843,1017,477,503]" italics="true" pageId="7" pageNumber="457">Triacanthoneus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D78FFFAFFCDFBAD1B8E03CCFE5F" authorityName="Komai & Fujita" authorityYear="2018" box="[1074,1202,477,503]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="457" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B955EAE9FFFAFFCDFBAD1B8E03CCFE5F" box="[1074,1202,477,503]" italics="true" pageId="7" pageNumber="457">Caligoneus</emphasis>
|
||
</taxonomicName>
|
||
and that is also may be variable within a genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |