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<document id="CB81FC9D1C479640739460694330C951" ID-CLB-Dataset="112021" ID-DOI="10.11646/zootaxa.5143.1.1" ID-GBIF-Dataset="9d7cad94-f472-47eb-a988-5569e479204e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6601404" ID-ZooBank="F71553B2-7D58-4E61-A883-546B2A0124D5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654063338489" checkinUser="plazi" docAuthor="Burington, Zelia L." docDate="2022" docId="038687B669788F7EFF1BFE3AFD4686E7" docLanguage="en" docName="zootaxa.5143.1.1.pdf" docOrigin="Zootaxa 5143 (1)" docSource="http://zoobank.org/f71553b2-7d58-4e61-a883-546b2a0124d5" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Eucelatoria hafelei Burington 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="52" masterDocId="FFBFFFCE694A8F4DFF8CFFE0FFA28416" masterDocTitle="A taxonomic revision of the Eucelatoria ferox species group (Diptera: Tachinidae)" masterLastPageNumber="104" masterPageNumber="1" pageNumber="51" updateTime="1699338509909" updateUser="ExternalLinkService">
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<mods:title id="9390F01E4118411D7EE0DF2A12782AA1">A taxonomic revision of the Eucelatoria ferox species group (Diptera: Tachinidae)</mods:title>
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<treatment id="038687B669788F7EFF1BFE3AFD4686E7" ID-DOI="http://doi.org/10.5281/zenodo.6958427" ID-GBIF-Taxon="198574810" ID-Zenodo-Dep="6958427" LSID="urn:lsid:plazi:treatment:038687B669788F7EFF1BFE3AFD4686E7" httpUri="http://treatment.plazi.org/id/038687B669788F7EFF1BFE3AFD4686E7" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
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<heading id="D0D881CC69788F7FFF1BFE3AFE4685E2" bold="true" box="[151,484,474,501]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="4">
<emphasis id="B95BEAB269788F7FFF1BFE3AFE4685E2" bold="true" box="[151,484,474,501]" pageId="50" pageNumber="51">
<taxonomicName id="4C2F4D2369788F7FFF1BFE3AFEDD85E3" ID-CoL="9J366" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[151,383,474,501]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="hafelei" status="sp. nov.">
<emphasis id="B95BEAB269788F7FFF1BFE3AFEDD85E3" bold="true" box="[151,383,474,501]" italics="true" pageId="50" pageNumber="51">Eucelatoria hafelei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857C969788F7FFE0BFE3AFE4685E2" box="[391,484,474,500]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B9036A069788F7FFF1BFE1EFDA5860F" blockId="50.[151,519,474,537]" box="[151,519,510,537]" pageId="50" pageNumber="51">
(
<figureCitation id="13142A2569788F7FFF13FE1EFF55860E" box="[159,247,510,536]" captionStart="FIGURES 4245" captionStartId="72.[151,264,1943,1967]" captionTargetBox="[172,1415,181,1920]" captionTargetId="figure-31@72.[172,1415,181,1920]" captionTargetPageId="72" captionText="FIGURES 4245. Male habitus photographs of Eucelatoria ferox group spp., dorsal view. 42. E. hafelei sp. nov., Chapare Province (ZLB_E.Ferox 00386) (DMW). 43. E. kopis sp. nov., allotype (ZLB_E.Ferox 00003) (CNC). 44. E. makhaira sp. nov., allotype (ZLB_E.Ferox 00343) (USNM). 45. E. ritavargasae sp. nov., holotype (ZLB_E.Ferox 00004) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601428" httpUri="https://zenodo.org/record/6601428/files/figure.png" pageId="50" pageNumber="51">Figs 42</figureCitation>
,
<figureCitation id="13142A2569788F7FFE8EFE1EFE82860E" box="[258,288,510,536]" captionStart="FIGURES 8687" captionStartId="93.[151,264,1943,1967]" captionTargetBox="[198,1390,181,1919]" captionTargetId="figure-15@93.[198,1390,181,1920]" captionTargetPageId="93" captionText="FIGURES 8687. Male habitus photographs of Eucelatoria ferox group spp., lateral view. 86. E. gustavogutierrezi sp. nov., Guanacaste Province (ZLB_E.Ferox 00008) (CNC). 87. E. hafelei sp. nov., Chapare Province (ZLB_E.Ferox 00386) (DMW)." figureDoi="http://doi.org/10.5281/zenodo.6601484" httpUri="https://zenodo.org/record/6601484/files/figure.png" pageId="50" pageNumber="51">87</figureCitation>
,
<figureCitation id="13142A2569788F7FFEA0FE1EFE32860F" box="[300,400,510,537]" captionStart="FIGURES 136155" captionStartId="100.[151,264,1677,1701]" captionTargetBox="[277,1319,184,1650]" captionTargetId="figure-31@100.[264,1324,181,1653]" captionTargetPageId="100" captionText="FIGURES 136155. Male terminalia of Eucelatoria ferox group spp., left lateral view, and caudal view of cerci and surstyli. Numbers are “ZLB_E.Ferox” specimen ID numbers. Specimen information for caudal views is identical to that for Figs 124 139. 136. E. gustavogutierrezi sp. nov., lateral view, Guanacaste Province (00008) (CNC). 137. E. hafelei sp. nov., lateral view, Chapare Province (00386) (DMW). 138. E. ritavargasae sp. nov., lateral view, Narupa (00215) (USNM). 139. E. sica sp. nov., lateral view, Yanayacu (00213) (CNC). 140. E. aurata (Townsend), caudal view. 141. E. auriceps (Aldrich), caudal view. 142. E. borealis sp. nov., caudal view. 143. E. charapensis (Townsend), caudal view. 144. E. falcata sp. nov., caudal view. 145. E. fordlandia sp. nov., caudal view. 146. E. gladiatrix (Townsend), caudal view. 147. E. sabroskyi sp. nov., caudal view. 148. E. strigata (Wulp), caudal view. 149. E. texana (Reinhard), caudal view. 150. E. woodorum sp. nov., caudal view. 151. E. yanayacu sp. nov., caudal view. 152. E. gustavogutierrezi sp. nov., caudal view. 153. E. hafelei sp. nov., caudal view. 154. E. ritavargasae sp. nov., caudal view. 155. E. sica sp. nov., caudal view." figureDoi="http://doi.org/10.5281/zenodo.6601508" httpUri="https://zenodo.org/record/6601508/files/figure.png" pageId="50" pageNumber="51">137, 153</figureCitation>
,
<figureCitation id="13142A2569788F7FFE10FE1EFE6A860E" box="[412,456,510,536]" captionStart="FIGURES 156173" captionStartId="101.[151,264,1755,1779]" captionTargetBox="[261,1320,195,1718]" captionTargetId="figure-15@101.[253,1335,181,1731]" captionTargetPageId="101" captionText="FIGURES 156173. Male terminalia and fifth abdominal sternite of Eucelatoria ferox group spp. Specimen information for fifth sternites is identical to that for Figs 124139. 156. Generic male terminalia, left lateral view (left) and caudal view (right), setae and membrane omitted.Abbreviations: aed, aedeagal apodeme; bas, basal lobe of surstylus; C, depth of postgonite anterior emargination; cer, cercus; epa, epandrium; hyp, hypandrium; pha, phallus; pos, postgonite; pre, pregonite; sur, surstylus. 157. E. aurata (Townsend). 158. E. auriceps (Aldrich). 159. E. borealis sp. nov. 160. E. charapensis (Townsend). 161. E. falcata sp. nov. 162. E. fordlandia sp. nov. 163. E. gladiatrix (Townsend). 164. E. sabroskyi sp. nov. 165. E. strigata (Wulp). 166. E. texana (Reinhard). 167. E. woodorum sp. nov. 168. E. yanayacu sp. nov. 169. E. gustavogutierrezi sp. nov. 170. E. hafelei sp. nov. 171. E. ritavargasae sp. nov. 172. E. sica sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601512" httpUri="https://zenodo.org/record/6601512/files/figure.png" pageId="50" pageNumber="51">170</figureCitation>
,
<figureCitation id="13142A2569788F7FFE58FE1EFE5D860F" box="[468,511,510,537]" captionStart="FIGURES 174179" captionStartId="103.[151,264,1856,1880]" captionTargetBox="[203,1374,189,1818]" captionTargetId="figure-14@103.[193,1393,181,1832]" captionTargetPageId="103" captionText="FIGURES 174179. Distribution maps of Eucelatoria spp. 174. E. yanayacu sp. nov., E. gladiatrix (Townsend), E. falcata sp. nov., E. charapensis (Townsend), and E. fordlandia sp. nov. 175. E. strigata (Wulp) and E. aurata (Townsend). 176. E. hafelei sp. nov., E. gustavogutierrezi sp. nov., and E. ritavargasae sp. nov. 177. E. ferox (Townsend), E. huitepecensis sp. nov., and E. inclani sp. nov. 178. E. kopis sp. nov., E. sica sp. nov., E. makhaira sp. nov., and E. tenebrionis sp. nov. 179. E. rivalis (Reinhard), E. luctuosa (Wulp), E. woodorum sp. nov., E. crambivora sp. nov., and E. jorgecortesi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601514" httpUri="https://zenodo.org/record/6601514/files/figure.png" pageId="50" pageNumber="51">176</figureCitation>
)
</paragraph>
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<paragraph id="8B9036A069788F7FFF1BFDA6FE1186BE" blockId="50.[151,1437,582,2013]" pageId="50" pageNumber="51">
<emphasis id="B95BEAB269788F7FFF1BFDA6FEE88676" bold="true" box="[151,330,582,608]" pageId="50" pageNumber="51">Type material.</emphasis>
<materialsCitation id="3B473CFD69788F7FFEDAFDA6FE1186BE" ID-GBIF-Occurrence="3878846619" collectionCode="DMW" collectorName="G. &amp; M. Wood" country="Bolivia" elevation="1083" location="Villa Tunari" municipality="Cbba Chapare" pageId="50" pageNumber="51" specimenCount="2" stateProvince="Cochabamba" typeStatus="holotype">
<typeStatus id="5494880269788F7FFEDAFDA6FE1C8676" box="[342,446,582,608]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
male, labeled “
<collectingCountry id="F338763069788F7FFDF5FDA6FD4C8676" box="[633,750,582,608]" name="Bolivia" pageId="50" pageNumber="51">BOLIVIA</collectingCountry>
<collectingMunicipality id="6BF4ACDA69788F7FFD75FDA6FC3F8676" box="[761,925,582,608]" pageId="50" pageNumber="51">Cbba Chapare</collectingMunicipality>
/
<location id="8EF0607B69788F7FFC3DFDA6FB9E8676" LSID="urn:lsid:plazi:treatment:038687B669788F7EFF1BFE3AFD4686E7:8EF0607B69788F7FFC3DFDA6FB9E8676" box="[945,1084,582,608]" country="Bolivia" municipality="Cbba Chapare" name="Villa Tunari" pageId="50" pageNumber="51" stateProvince="Cochabamba">Villa Tunarí</location>
<collectingRegion id="49EBF84269788F7FFBC6FDA6FB7E8676" box="[1098,1244,582,608]" country="Bolivia" name="Cochabamba" pageId="50" pageNumber="51">Cochabamba</collectingRegion>
/ road - km
<quantity id="4CD79B4569788F7FFAFEFDA6FF4F8692" metricMagnitude="3" metricUnit="m" metricValue="1.0825" metricValueMax="1.8" metricValueMin="0.365" pageId="50" pageNumber="51" unit="m" value="1082.5" valueMax="1800.0" valueMin="365.0">
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</quantity>
/
<collectorName id="26DA537669788F7FFE8DFD8AFE168692" box="[257,436,618,645]" pageId="50" pageNumber="51">G. &amp; M. Wood</collectorName>
310.xii.96”, “
<typeStatus id="5494880269788F7FFDEAFD8AFD5E8692" box="[614,764,618,644]" pageId="50" pageNumber="51" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C2F4D2369788F7FFC83FD8AFC2C8692" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" box="[783,910,618,644]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ hafelei/
<collectorName id="26DA537669788F7FFB8CFD8BFB0F8692" box="[1024,1197,618,645]" pageId="50" pageNumber="51">Z.L. Burington</collectorName>
[red label]”, “ZLB_ E.Ferox 00386” (
<collectionCode id="ED3EAE6569788F7FFED5FD6EFE0586BE" box="[345,423,654,680]" pageId="50" pageNumber="51">DMW</collectionCode>
).
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<materialsCitation id="3B473CFD69788F7FFF4BFD53FDDD86DA" ID-GBIF-Occurrence="3878846318" box="[199,639,690,717]" location="Cochabamba" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="Cochabamba" typeStatus="paratype">
<typeStatus id="5494880269788F7FFF4BFD53FE8E86DB" box="[199,300,691,717]" pageId="50" pageNumber="51" type="paratype">Paratype</typeStatus>
,
<specimenCount id="9D29FD2969788F7FFEB7FD52FECF86DB" box="[315,365,690,717]" pageId="50" pageNumber="51" type="male">1 ♂</specimenCount>
. Bolivia
<emphasis id="B95BEAB269788F7FFE43FD53FE4A86DA" box="[463,488,691,716]" italics="true" pageId="50" pageNumber="51"></emphasis>
<collectingRegion id="49EBF84269788F7FFE64FD52FDDD86DA" box="[488,639,690,716]" country="Bolivia" name="Cochabamba" pageId="50" pageNumber="51">Cochabamba</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B473CFD69788F7FFD01FD52FCDC8702" ID-GBIF-Occurrence="3878846546" collectionCode="DMW" collectorName="Z. L. Burington" country="Bolivia" elevation="829" location="Villa Tunari" municipality="Cbba Chapare" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="Cochabamba" typeStatus="paratype">
<specimenCount id="9D29FD2969788F7FFD01FD52FD1F86DB" box="[653,701,690,717]" pageId="50" pageNumber="51" type="male">1 ♂</specimenCount>
, labeled “
<collectingCountry id="F338763069788F7FFCBFFD52FC0A86DA" box="[819,936,690,716]" name="Bolivia" pageId="50" pageNumber="51">BOLIVIA</collectingCountry>
<collectingMunicipality id="6BF4ACDA69788F7FFC3FFD52FBF586DA" box="[947,1111,690,716]" pageId="50" pageNumber="51">Cbba Chapare</collectingMunicipality>
/
<location id="8EF0607B69788F7FFBE6FD52FB5786DA" LSID="urn:lsid:plazi:treatment:038687B669788F7EFF1BFE3AFD4686E7:8EF0607B69788F7FFBE6FD52FB5786DA" box="[1130,1269,690,716]" country="Bolivia" municipality="Cbba Chapare" name="Villa Tunari" pageId="50" pageNumber="51" stateProvince="Cochabamba">Villa Tunarí</location>
<collectingRegion id="49EBF84269788F7FFA8EFD52FA3686DA" box="[1282,1428,690,716]" country="Bolivia" name="Cochabamba" pageId="50" pageNumber="51">Cochabamba</collectingRegion>
/ road - km
<quantity id="4CD79B4569788F7FFE81FD36FE3686E6" box="[269,404,726,753]" metricMagnitude="2" metricUnit="m" metricValue="8.29" metricValueMax="13.0" metricValueMin="3.5799999999999996" pageId="50" pageNumber="51" unit="m" value="829.0" valueMax="1300.0" valueMin="358.0">
<elevation id="0002D19369788F7FFE81FD36FE3686E6" box="[269,404,726,753]" metricMagnitude="2" metricUnit="m" metricValue="8.29" metricValueMax="13.0" metricValueMin="3.5799999999999996" pageId="50" pageNumber="51" unit="m" value="829.0" valueMax="1300.0" valueMin="358.0">358 -1300m</elevation>
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/ Pruett &amp; Wood 23.III.95”, “
<typeStatus id="5494880269788F7FFD57FD36FCCB86E6" box="[731,873,726,752]" pageId="50" pageNumber="51" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C2F4D2369788F7FFCF5FD36FC5A86E6" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" box="[889,1016,726,752]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ hafelei/
<collectorName id="26DA537669788F7FFBE8FD37FAAF86E6" box="[1124,1293,726,753]" pageId="50" pageNumber="51">Z.L. Burington</collectorName>
[red label]”, “ZLB_E.Ferox 00387” [genitalia in vial under specimen] (
<collectionCode id="ED3EAE6569788F7FFCA4FD1AFCD48702" box="[808,886,762,788]" pageId="50" pageNumber="51">DMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9036A069788F7FFF4BFCFEFE5787FA" blockId="50.[151,1437,582,2013]" pageId="50" pageNumber="51">
<emphasis id="B95BEAB269788F7FFF4BFCFEFEFF872E" bold="true" box="[199,349,798,824]" pageId="50" pageNumber="51">Recognition.</emphasis>
This species is closest to
<taxonomicName id="4C2F4D2369788F7FFDFBFCFFFC81872E" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[631,803,799,824]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="ritavargasae" status="sp. nov.">
<emphasis id="B95BEAB269788F7FFDFBFCFFFC81872E" box="[631,803,799,824]" italics="true" pageId="50" pageNumber="51">E. ritavargasae</emphasis>
</taxonomicName>
<emphasis id="B95BEAB269788F7FFCA5FCFEFC23872E" bold="true" box="[809,897,798,824]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="A26857C969788F7FFCA5FCFEFC23872E" box="[809,897,798,824]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; in both species the palpi are dark, the presutural thoracic vittae are fused into two, the postsutural thoracic vittae are fused into one, and the scutellum bears visible tomentum only at the extreme apex. Males of
<taxonomicName id="4C2F4D2369788F7FFD15FC87FCA18796" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[665,771,870,896]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="hafelei" status="sp. nov.">
<emphasis id="B95BEAB269788F7FFD15FC87FCA18796" box="[665,771,870,896]" italics="true" pageId="50" pageNumber="51">E. hafelei</emphasis>
</taxonomicName>
<emphasis id="B95BEAB269788F7FFC86FC86FCC68796" bold="true" box="[778,868,870,896]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="A26857C969788F7FFC86FC86FCC68796" box="[778,868,870,896]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are distinguished from
<taxonomicName id="4C2F4D2369788F7FFBE1FC87FABB8796" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[1133,1305,871,896]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="ritavargasae" status="sp. nov.">
<emphasis id="B95BEAB269788F7FFBE1FC87FABB8796" box="[1133,1305,871,896]" italics="true" pageId="50" pageNumber="51">E. ritavargasae</emphasis>
</taxonomicName>
<emphasis id="B95BEAB269788F7FFAACFC86FAD88796" bold="true" box="[1312,1402,870,896]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="A26857C969788F7FFAACFC86FAD88796" box="[1312,1402,870,896]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the broader, denser dorsal abdominal bands, the less dilated cercus apex in lateral view, the more narrowly separated basal lobes of the cercus, the long setae at the apex of the surstylus, and the more rounded, shallower anterior emargination of the postgonite.
</paragraph>
</subSubSection>
<subSubSection id="C335652B69788F7FFF4BFC16FE7F804E" pageId="50" pageNumber="51" type="etymology">
<paragraph id="8B9036A069788F7FFF4BFC16FE7F804E" blockId="50.[151,1437,582,2013]" pageId="50" pageNumber="51">
<emphasis id="B95BEAB269788F7FFF4BFC16FEEC8006" bold="true" box="[199,334,1014,1040]" pageId="50" pageNumber="51">Etymology.</emphasis>
This species is named in honor and memory of Dr. Joseph C. Hafele (19442014), physicist, grandfather of the author, known for conducting the first experimental test of time dilation under special relativity with Dr. Richard Keating in 1971.
</paragraph>
</subSubSection>
<subSubSection id="C335652B69788F7EFF4BFB82FE3D85E3" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
<paragraph id="8B9036A069788F7FFF4BFB82FC68806B" blockId="50.[151,1437,582,2013]" box="[199,970,1122,1149]" pageId="50" pageNumber="51">
<emphasis id="B95BEAB269788F7FFF4BFB82FEFB806A" bold="true" box="[199,345,1122,1148]" pageId="50" pageNumber="51">Description.</emphasis>
<emphasis id="B95BEAB269788F7FFEECFB82FE38806A" box="[352,410,1122,1148]" italics="true" pageId="50" pageNumber="51">Male</emphasis>
[described from
<specimenCount id="9D29FD2969788F7FFDD5FB82FD4B806B" box="[601,745,1122,1149]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
]. Length
<quantity id="4CD79B4569788F7FFCDCFB82FC67806B" box="[848,965,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="8.95" metricValueMax="9.0" metricValueMin="8.9" pageId="50" pageNumber="51" unit="mm" value="8.95" valueMax="9.0" valueMin="8.9">8.99 mm</quantity>
.
</paragraph>
<paragraph id="8B9036A069788F7FFF4BFB66FAC0823A" blockId="50.[151,1437,582,2013]" pageId="50" pageNumber="51">Head. Parafacial, fronto-orbital plate, vertex, upper half of post-ocular plate, and vibrissal angle tomentum dense yellow, appearing gold. Genal dilation, lower half of post-ocular plate, postgena, and occiput tomentum ashgray, diffuse on gena such that it appears close to black. Pale occipital setae gray on lower half, merging to yellow on upper half. Ocellar setae three-fours to subequal to length of posterior reclinate orbital seta. Gena with 45 setae, subvibrissal ridge with 2 setae. Facial ridge with setulae on lower one-third to one-half. Postocellar setae one-half to three-fourths length of ocellar setae. Paravertical seta one-half length of postocellar setae. Outer vertical seta not differentiated from post-ocular row. Reclinate orbital setae 3. Frontal setae 910, last frontal seta level with base of arista or slightly beyond. Eye with short setulae, each setula no longer than four facets. Eye height to head height ratio 0.800.84. Postpedicel length 0.370.40 height of head. Facial ridge length 0.440.45 height of head. Parafacial width 0.120.13 lateral length of head. Pedicel 0.360.41 length of postpedicel. Postpedicel 1.7 times width of parafacial in lateral view. Vertex 0.220.23 width of head in dorsal view. Palpus black, slightly flattened and expanded at apex in male, with many short, stout dorsal and ventral setae, and several longer, thinner setae mediolaterally.</paragraph>
<paragraph id="8B9036A069788F7FFF4BF9D6FE9C8366" blockId="50.[151,1437,582,2013]" pageId="50" pageNumber="51">Thorax. Dorsomedial length 1.4 width of thorax. Lateral tomentum ash-gray, merging to yellow on anepisternum. Dorsal tomentum ash-gray to yellow, usually more gray medially. Presutural outer and inner vittae fused, at most with a narrow line of diffuse tomentum near notopleural suture; the two subquadrate bands separated medially by subtriangular region of tomentum, pointed anteriorly. Postsutural vittae merged into single vitta covering most of area, with only narrow region of tomentum visible between postsutural supra-alar and intra-alar setae. Scutellar dorsal tomentum diffuse, gray to yellow, visible only between subapical setae at apex in dorsal view. Postpronotum with 3 setae. Presutural area with 2 supra-alar setae. Postsutural area with 3 dorsocentral setae. Fore tibia with 3 posterodorsal setae. Wing infuscated in membrane adjacent to costa, radius, and in the dm cell. Calypters tan to cream colored.</paragraph>
<paragraph id="8B9036A069788F7FFF4BF89AFA3B83CA" blockId="50.[151,1437,582,2013]" pageId="50" pageNumber="51">Abdomen. Cuticle and setulae entirely black. Dorsal tomentum bands ash-gray to yellow, covering one-third to one-half length of T35; with wide median vitta. Ventral tomentum bands ash-gray, covering one-half to two-thirds of T35. T4 median marginal setae and lateral marginal setae forming continuous, widely spaced row of 5 or more pairs.</paragraph>
<paragraph id="8B9036A069798F7EFF4BFF77FDD385C7" blockId="51.[151,1437,150,753]" pageId="51" pageNumber="52">
Male terminalia (
<figureCitation id="13142A2569798F7EFE00FF77FD8F84A7" box="[396,557,150,177]" captionStart="FIGURES 136155" captionStartId="100.[151,264,1677,1701]" captionTargetBox="[277,1319,184,1650]" captionTargetId="figure-31@100.[264,1324,181,1653]" captionTargetPageId="100" captionText="FIGURES 136155. Male terminalia of Eucelatoria ferox group spp., left lateral view, and caudal view of cerci and surstyli. Numbers are “ZLB_E.Ferox” specimen ID numbers. Specimen information for caudal views is identical to that for Figs 124 139. 136. E. gustavogutierrezi sp. nov., lateral view, Guanacaste Province (00008) (CNC). 137. E. hafelei sp. nov., lateral view, Chapare Province (00386) (DMW). 138. E. ritavargasae sp. nov., lateral view, Narupa (00215) (USNM). 139. E. sica sp. nov., lateral view, Yanayacu (00213) (CNC). 140. E. aurata (Townsend), caudal view. 141. E. auriceps (Aldrich), caudal view. 142. E. borealis sp. nov., caudal view. 143. E. charapensis (Townsend), caudal view. 144. E. falcata sp. nov., caudal view. 145. E. fordlandia sp. nov., caudal view. 146. E. gladiatrix (Townsend), caudal view. 147. E. sabroskyi sp. nov., caudal view. 148. E. strigata (Wulp), caudal view. 149. E. texana (Reinhard), caudal view. 150. E. woodorum sp. nov., caudal view. 151. E. yanayacu sp. nov., caudal view. 152. E. gustavogutierrezi sp. nov., caudal view. 153. E. hafelei sp. nov., caudal view. 154. E. ritavargasae sp. nov., caudal view. 155. E. sica sp. nov., caudal view." figureDoi="http://doi.org/10.5281/zenodo.6601508" httpUri="https://zenodo.org/record/6601508/files/figure.png" pageId="51" pageNumber="52">Figs 137, 153</figureCitation>
,
<figureCitation id="13142A2569798F7EFDB7FF77FDCB84A7" box="[571,617,151,177]" captionStart="FIGURES 156173" captionStartId="101.[151,264,1755,1779]" captionTargetBox="[261,1320,195,1718]" captionTargetId="figure-15@101.[253,1335,181,1731]" captionTargetPageId="101" captionText="FIGURES 156173. Male terminalia and fifth abdominal sternite of Eucelatoria ferox group spp. Specimen information for fifth sternites is identical to that for Figs 124139. 156. Generic male terminalia, left lateral view (left) and caudal view (right), setae and membrane omitted.Abbreviations: aed, aedeagal apodeme; bas, basal lobe of surstylus; C, depth of postgonite anterior emargination; cer, cercus; epa, epandrium; hyp, hypandrium; pha, phallus; pos, postgonite; pre, pregonite; sur, surstylus. 157. E. aurata (Townsend). 158. E. auriceps (Aldrich). 159. E. borealis sp. nov. 160. E. charapensis (Townsend). 161. E. falcata sp. nov. 162. E. fordlandia sp. nov. 163. E. gladiatrix (Townsend). 164. E. sabroskyi sp. nov. 165. E. strigata (Wulp). 166. E. texana (Reinhard). 167. E. woodorum sp. nov. 168. E. yanayacu sp. nov. 169. E. gustavogutierrezi sp. nov. 170. E. hafelei sp. nov. 171. E. ritavargasae sp. nov. 172. E. sica sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601512" httpUri="https://zenodo.org/record/6601512/files/figure.png" pageId="51" pageNumber="52">170</figureCitation>
). Sternite 5 basal plate with median teeth separated 0.37 width of basal notch; apical lobes 0.8 times length of basal plate. Postgonite broadly rounded on posterior margin; broadened on basal third, then narrowed to parallel sided, digitiform apex; anterior emargination depth subequal width of postgonite at mid length. Surstylus paddle shaped; three times as wide as long; posterior and anterior margins gradually curved; apex blunt with long setae; basal lobe margin rounded. Cercus in lateral view gradually narrowed from base to mid length, then slightly dilated at apex; in caudal view narrowed to mid length then broadened to blunt apex. Upper lobe rectangular; inner margin between lobes u-shaped; 0.24 length of cercus. Median section 0.5 length of cercus. Apical cleft divergent at approximately a 20 degree angle, 0.28 length of cercus. Syncercus apex width in caudal view 0.51 width of syncercus base.
</paragraph>
<paragraph id="8B9036A069798F7EFF4BFE3BFE3D85E3" blockId="51.[151,1437,150,753]" box="[199,415,475,501]" pageId="51" pageNumber="52">
<emphasis id="B95BEAB269798F7EFF4BFE3BFEB985E3" box="[199,283,475,501]" italics="true" pageId="51" pageNumber="52">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C335652B69798F7EFF4BFE1EFCD18674" pageId="51" pageNumber="52" type="biology_ecology">
<paragraph id="8B9036A069798F7EFF4BFE1EFE03860F" blockId="51.[151,1437,150,753]" box="[199,417,510,537]" pageId="51" pageNumber="52">
<emphasis id="B95BEAB269798F7EFF4BFE1EFE86860E" bold="true" box="[199,292,510,536]" pageId="51" pageNumber="52">Host(s).</emphasis>
Unknown.
</paragraph>
<paragraph id="8B9036A069798F7EFF4BFDC2FCD18674" blockId="51.[151,1437,150,753]" pageId="51" pageNumber="52">
<emphasis id="B95BEAB269798F7EFF4BFDC2FD6C862B" bold="true" box="[199,718,546,573]" pageId="51" pageNumber="52">Geographic extent and seasonal occurrence.</emphasis>
Individuals of
<taxonomicName id="4C2F4D2369798F7EFCF9FDC3FC7C862B" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[885,990,547,573]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="hafelei" status="sp. nov.">
<emphasis id="B95BEAB269798F7EFCF9FDC3FC7C862B" box="[885,990,547,573]" italics="true" pageId="51" pageNumber="52">E. hafelei</emphasis>
</taxonomicName>
<emphasis id="B95BEAB269798F7EFC68FDC3FB9F862B" bold="true" box="[996,1085,547,573]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A26857C969798F7EFC68FDC3FB9F862B" box="[996,1085,547,573]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have been collected from tropical montane forests in
<collectingCountry id="F338763069798F7EFE19FDA7FE458677" box="[405,487,583,609]" name="Bolivia" pageId="51" pageNumber="52">Bolivia</collectingCountry>
at elevations of
<quantity id="4CD79B4569798F7EFD2CFDA7FD578677" box="[672,757,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="51" pageNumber="52" unit="m" value="2200.0">2200 m</quantity>
(
<figureCitation id="13142A2569798F7EFC88FDA7FCC58674" box="[772,871,583,610]" captionStart="FIGURES 174179" captionStartId="103.[151,264,1856,1880]" captionTargetBox="[203,1374,189,1818]" captionTargetId="figure-14@103.[193,1393,181,1832]" captionTargetPageId="103" captionText="FIGURES 174179. Distribution maps of Eucelatoria spp. 174. E. yanayacu sp. nov., E. gladiatrix (Townsend), E. falcata sp. nov., E. charapensis (Townsend), and E. fordlandia sp. nov. 175. E. strigata (Wulp) and E. aurata (Townsend). 176. E. hafelei sp. nov., E. gustavogutierrezi sp. nov., and E. ritavargasae sp. nov. 177. E. ferox (Townsend), E. huitepecensis sp. nov., and E. inclani sp. nov. 178. E. kopis sp. nov., E. sica sp. nov., E. makhaira sp. nov., and E. tenebrionis sp. nov. 179. E. rivalis (Reinhard), E. luctuosa (Wulp), E. woodorum sp. nov., E. crambivora sp. nov., and E. jorgecortesi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601514" httpUri="https://zenodo.org/record/6601514/files/figure.png" pageId="51" pageNumber="52">Fig. 176</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335652B69798F7EFF4BFD8AFD4686E7" pageId="51" pageNumber="52" type="discussion">
<paragraph id="8B9036A069798F7EFF4BFD8AFD4686E7" blockId="51.[151,1437,150,753]" pageId="51" pageNumber="52">
<emphasis id="B95BEAB269798F7EFF4BFD8AFEEE8692" bold="true" box="[199,332,618,644]" pageId="51" pageNumber="52">Discussion.</emphasis>
Only
<specimenCount id="9D29FD2969798F7EFE1AFD8BFDA88693" box="[406,522,619,645]" pageId="51" pageNumber="52" type="male">two males</specimenCount>
are known for this species, but the genitalia and external characters are distinct enough from
<taxonomicName id="4C2F4D2369798F7EFEBFFD6FFE4086BE" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[307,482,655,680]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="ritavargasae" status="sp. nov.">
<emphasis id="B95BEAB269798F7EFEBFFD6FFE4086BE" box="[307,482,655,680]" italics="true" pageId="51" pageNumber="52">E. ritavargasae</emphasis>
</taxonomicName>
<emphasis id="B95BEAB269798F7EFE60FD6FFDEA86BF" bold="true" box="[492,584,655,681]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A26857C969798F7EFE60FD6FFDEA86BF" box="[492,584,655,681]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and other species with two presutural vittae and limited tomentum on the scutellum to warrant description. Females are likely similar in piercer length and shape to
<taxonomicName id="4C2F4D2369798F7EFB02FD53FA9986DA" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[1166,1339,691,716]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="ritavargasae" status="sp. nov.">
<emphasis id="B95BEAB269798F7EFB02FD53FA9986DA" box="[1166,1339,691,716]" italics="true" pageId="51" pageNumber="52">E. ritavargasae</emphasis>
</taxonomicName>
<emphasis id="B95BEAB269798F7EFACEFD53FA3E86DB" bold="true" box="[1346,1436,691,717]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A26857C969798F7EFACEFD53FA3E86DB" box="[1346,1436,691,717]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and should have wider abdominal bands as in males.
</paragraph>
</subSubSection>
</treatment>
</document>