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<document id="92AF790AB75FC1468B1C72ADFB4477F1" ID-CLB-Dataset="27584" ID-DOI="10.11646/zootaxa.4551.4.9" ID-GBIF-Dataset="af287f7c-0f70-43de-9bb1-b6275833a1c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623143" ID-ZooBank="F70F17F4-B821-4F34-90A7-AA9502F06604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554270592672" checkinUser="plazi" docAuthor="Araújo, Maíra Xavier &amp; Bravo, Freddy" docDate="2019" docId="038687B3FFCAFF9BFF7116843D392507" docLanguage="en" docName="zootaxa.4551.4.9.pdf" docOrigin="Zootaxa 4551 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidiella maculosa Araújo &amp; Bravo 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="489" masterDocId="FFBFFFCBFFCAFF99FFE6107E3F22263D" masterDocTitle="Two new species of Lepidiella Enderlein, 1937 (Diptera: Psychodidae) from the Neotropical Region with taxonomic comments about the species of the genus" masterLastPageNumber="493" masterPageNumber="487" pageNumber="487" updateTime="1698716187655" updateUser="ExternalLinkService">
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<mods:title id="113B0AB002F4448DBEA023D8E961215E">Two new species of Lepidiella Enderlein, 1937 (Diptera: Psychodidae) from the Neotropical Region with taxonomic comments about the species of the genus</mods:title>
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<mods:namePart id="AC4FA0C385D919CA7852D6D2C44932D0">Araújo, Maíra Xavier</mods:namePart>
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<mods:namePart id="9020723C4738E23E8C521239D168CEDC">Bravo, Freddy</mods:namePart>
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<mods:title id="D4AA189E645AF73A5C8D209ED90FADD3">Zootaxa</mods:title>
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<mods:date id="F58B2DD3F2CDF8899EFC6D740DBF66F2">2019</mods:date>
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<mods:number id="1ACBA0757558B1CE4343257A2B7BFE6A">2019-02-04</mods:number>
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<treatment id="038687B3FFCAFF9BFF7116843D392507" ID-DOI="http://doi.org/10.5281/zenodo.5931395" ID-GBIF-Taxon="154534314" ID-Zenodo-Dep="5931395" LSID="urn:lsid:plazi:treatment:038687B3FFCAFF9BFF7116843D392507" httpUri="http://treatment.plazi.org/id/038687B3FFCAFF9BFF7116843D392507" lastPageId="2" lastPageNumber="489" pageId="0" pageNumber="487">
<subSubSection id="C335652EFFCAFF99FF7116843ED52128" box="[151,503,1786,1813]" pageId="0" pageNumber="487" type="nomenclature">
<paragraph id="8B9036A5FFCAFF99FF7116843ED52128" blockId="0.[151,503,1786,1846]" box="[151,503,1786,1813]" pageId="0" pageNumber="487">
<heading id="D0D881C9FFCAFF99FF7116843ED52128" bold="true" box="[151,503,1786,1813]" fontSize="11" level="1" pageId="0" pageNumber="487" reason="1">
<emphasis id="B95BEAB7FFCAFF99FF7116843ED52128" bold="true" box="[151,503,1786,1813]" pageId="0" pageNumber="487">
<emphasis id="B95BEAB7FFCAFF99FF7116843EB32129" bold="true" box="[151,401,1786,1812]" italics="true" pageId="0" pageNumber="487">
<taxonomicName id="4C2F4D26FFCAFF99FF7116843EA92129" authority="Araújo &amp; Bravo, 2019" authorityName="Araújo &amp; Bravo" authorityYear="2019" box="[151,395,1786,1812]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="487" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">Lepidiella maculosa</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26857CCFFCAFF99FE7F16853ED52128" box="[409,503,1787,1813]" pageId="0" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C335652EFFCAFF99FF7117633E05210B" box="[151,295,1821,1846]" pageId="0" pageNumber="487" type="description">
<paragraph id="8B9036A5FFCAFF99FF7117633E05210B" blockId="0.[151,503,1786,1846]" box="[151,295,1821,1846]" pageId="0" pageNumber="487">
(
<figureCitation id="13142A20FFCAFF99FF7917633FC5210B" box="[159,231,1821,1846]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="0" pageNumber="487">Figs. 1</figureCitation>
AF)
</paragraph>
</subSubSection>
<subSubSection id="C335652EFFCAFF99FF71171B3C1621FB" pageId="0" pageNumber="487" type="materials_examined">
<paragraph id="8B9036A5FFCAFF99FF71171B3C1621FB" blockId="0.[151,1436,1893,2026]" pageId="0" pageNumber="487">
<emphasis id="B95BEAB7FFCAFF99FF71171B3E712143" bold="true" box="[151,339,1893,1918]" pageId="0" pageNumber="487">Type material.</emphasis>
<materialsCitation id="3B473CF8FFCAFF99FE8E171B3C1621FB" ID-GBIF-Occurrence="2234226767" collectingDate="2017-05-10" collectionCode="GAMBA, MZFS" collectorName="Silva-Neto &amp; Mendes &amp; Moura" country="Brazil" latitude="-12.85" location="Sede" longLatPrecision="1292" longitude="-39.466667" municipality="Serra da Jiboia" pageId="0" pageNumber="487" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="54948807FFCAFF99FE8E171B3EF12143" box="[360,467,1893,1918]" pageId="0" pageNumber="487" type="holotype">Holotype</typeStatus>
, male:
<collectingCountry id="F3387635FFCAFF99FDA617183D8A2143" box="[576,680,1894,1918]" name="Brazil" pageId="0" pageNumber="487">BRAZIL</collectingCountry>
,
<collectingRegion id="49EBF847FFCAFF99FD27171B3C272143" box="[705,773,1893,1918]" country="Brazil" name="Bahia" pageId="0" pageNumber="487">Bahia</collectingRegion>
, [Elísio Medrado],
<collectingMunicipality id="6BF4ACDFFFCAFF99FBEF17183BED2143" box="[1033,1231,1893,1918]" pageId="0" pageNumber="487">Serra da Jibóia</collectingMunicipality>
,
<location id="8EF0607EFFCAFF99FB0E171B3A3C2143" LSID="urn:lsid:plazi:treatment:038687B3FFCAFF9BFF7116843D392507:8EF0607EFFCAFF99FB0E171B3A3C2143" box="[1256,1310,1893,1918]" country="Brazil" latitude="-12.85" longLatPrecision="1292" longitude="-39.466667" municipality="Serra da Jiboia" name="Sede" pageId="0" pageNumber="487" stateProvince="Bahia">Sede</location>
<collectionCode id="ED3EAE60FFCAFF99FAD217183ABE2143" box="[1332,1436,1894,1918]" pageId="0" pageNumber="487">GAMBA</collectionCode>
[Environmentalist Group of
<collectingRegion id="49EBF847FFCAFF99FE3B17F73D03219F" box="[477,545,1929,1954]" country="Brazil" name="Bahia" pageId="0" pageNumber="487">Bahia</collectingRegion>
] [
<geoCoordinate id="EE1B5062FFCAFF99FDDF17F43D84219F" box="[569,678,1929,1954]" direction="south" orientation="latitude" pageId="0" pageNumber="487" precision="925" value="-12.85">12° 51' S</geoCoordinate>
;
<geoCoordinate id="EE1B5062FFCAFF99FD5317F43C07219C" box="[693,805,1930,1954]" direction="west" orientation="longitude" pageId="0" pageNumber="487" precision="925" value="-39.466667">39 28' W</geoCoordinate>
], (
<collectingMethod id="526E4EB2FFCAFF99FCA417F73C92219F" box="[834,944,1929,1954]" pageId="0" pageNumber="487">ligth trap</collectingMethod>
),
<date id="FF911065FFCAFF99FC2017F43B15219F" box="[966,1079,1930,1954]" pageId="0" pageNumber="487" value="2017-05-10">
<collectingDate id="EFD5E98DFFCAFF99FC2017F43B15219F" box="[966,1079,1930,1954]" pageId="0" pageNumber="487" value="2017-05-10">10.v.2017</collectingDate>
</date>
,
<collectorName id="26DA5373FFCAFF99FBA317F73BE3219F" box="[1093,1217,1929,1954]" pageId="0" pageNumber="487">Silva-Neto</collectorName>
,
<collectorName id="26DA5373FFCAFF99FB3617F73A0B219F" box="[1232,1321,1929,1954]" pageId="0" pageNumber="487">Mendes</collectorName>
&amp;
<collectorName id="26DA5373FFCAFF99FAB717F43ABE219C" box="[1361,1436,1930,1953]" pageId="0" pageNumber="487">Moura</collectorName>
cols. (
<collectionCode id="ED3EAE60FFCAFF99FF3C17D03E0B21FB" box="[218,297,1966,1990]" pageId="0" pageNumber="487">MZFS</collectionCode>
), specimen dissected, mounted on micro-slide.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C335652EFFCAFF99FF2117AF3A0E21D7" box="[199,1324,2001,2026]" pageId="0" pageNumber="487" type="etymology">
<paragraph id="8B9036A5FFCAFF99FF2117AF3A0E21D7" blockId="0.[151,1436,1893,2026]" box="[199,1324,2001,2026]" pageId="0" pageNumber="487">
<emphasis id="B95BEAB7FFCAFF99FF2117AF3E6C21D7" bold="true" box="[199,334,2001,2026]" pageId="0" pageNumber="487">Etymology.</emphasis>
The specific epithet, from Latin
<emphasis id="B95BEAB7FFCAFF99FD5917AF3C1721D4" box="[703,821,2001,2025]" italics="true" pageId="0" pageNumber="487">maculosus</emphasis>
, spotted, refers to the marks on hypandrium.
</paragraph>
</subSubSection>
<caption id="DF50662DFFCBFF98FF7116E93E272134" httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="1" pageNumber="488" startId="1.[151,250,1687,1709]" targetBox="[151,1429,219,1666]" targetPageId="1">
<paragraph id="8B9036A5FFCBFF98FF7116E93E272134" blockId="1.[151,1436,1686,1801]" pageId="1" pageNumber="488">
<emphasis id="B95BEAB7FFCBFF98FF7116E93E7C2090" bold="true" box="[151,350,1686,1709]" pageId="1" pageNumber="488">FIGURE 1 (AH).</emphasis>
<taxonomicName id="4C2F4D26FFCBFF98FE8316E93D172090" authority="Araújo &amp; Bravo, 2019" authorityName="Araújo &amp; Bravo" authorityYear="2019" box="[357,565,1687,1709]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="488" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="B95BEAB7FFCBFF98FE8316E93D172090" box="[357,565,1687,1709]" italics="true" pageId="1" pageNumber="488">Lepidiella maculosa</emphasis>
</taxonomicName>
<emphasis id="B95BEAB7FFCBFF98FDDA16E93DAF2091" bold="true" box="[572,653,1687,1708]" pageId="1" pageNumber="488">
<taxonomicNameLabel id="A26857CCFFCBFF98FDDA16E93DAF2091" box="[572,653,1687,1708]" pageId="1" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere).
</paragraph>
</caption>
<subSubSection id="C335652EFFCBFF98FF21174E3CD12188" pageId="1" pageNumber="488" type="diagnosis">
<paragraph id="8B9036A5FFCBFF98FF21174E3CD12188" blockId="1.[151,1436,1840,2009]" pageId="1" pageNumber="488">
<emphasis id="B95BEAB7FFCBFF98FF21174E3E632174" bold="true" box="[199,321,1840,1865]" pageId="1" pageNumber="488">Diagnosis.</emphasis>
Head without cornicula; eyes separated by 2.0 facet diameters; scape without internal protuberance; interocular suture inverted V-shaped; hypandrium with microtrichia arranged in circular groups; aedeagus symmetrical, bifid, digitiform with a medial external lobe; two pairs of parameres, external pair fused medially forming a U-shaped plate; internal pair well sclerotized, lanceolate.
</paragraph>
</subSubSection>
<subSubSection id="C335652EFFCBFF9BFF2117BE3EB42496" lastPageId="2" lastPageNumber="489" pageId="1" pageNumber="488" type="description">
<paragraph id="8B9036A5FFCBFF9BFF2117BE3C0024BA" blockId="1.[151,1436,1840,2009]" lastBlockId="2.[151,1436,151,827]" lastPageId="2" lastPageNumber="489" pageId="1" pageNumber="488">
<emphasis id="B95BEAB7FFCBFF98FF2117BE3E7B21E4" bold="true" box="[199,345,1984,2009]" pageId="1" pageNumber="488">Description.</emphasis>
Male
<typeStatus id="54948807FFCBFF98FE4617BE3D2421E4" box="[416,518,1984,2009]" pageId="1" pageNumber="488" type="holotype">holotype</typeStatus>
. Head without corniculum. Vertex dorsally expanded. Eyes separated by 2.0 facet diameters; eye bridge with four facet rows; interocular suture inverted V-shaped; frons with hair patch not divided at center, extending dorsally and reaching the interocular suture (
<figureCitation id="13142A20FFC8FF9BFC9F10C23CF626E9" box="[889,980,188,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Fig. 1A</figureCitation>
). Scape subspherical, approximately the same length as pedicel, without internal protuberances. Antenna incomplete; first three flagellomeres fusiform, each one with pair of filiform ascoids (
<figureCitation id="13142A20FFC8FF9BFDBC117A3D912721" box="[602,691,260,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Fig. 1B</figureCitation>
). Relative palpomere proportion: 1.0:1.1:1.2:2.0 (
<figureCitation id="13142A20FFC8FF9BFB08117A3A6A2721" box="[1262,1352,260,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Fig. 1C</figureCitation>
). Wing (
<figureCitation id="13142A20FFC8FF9BFF7911593FD9277D" box="[159,251,295,320]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Fig. 1D</figureCitation>
). 2.5 times longer than wide; wing membrane brownish, with costal cell darker; lighter spots between veins R
<subScript id="17AB34E0FFC8FF9BFF0D11253FD12754" attach="left" box="[235,243,347,361]" fontSize="6" pageId="2" pageNumber="489">5</subScript>
/M
<subScript id="17AB34E0FFC8FF9BFEF511253E392754" attach="left" box="[275,283,347,361]" fontSize="6" pageId="2" pageNumber="489">1</subScript>
to CuA; Sc not reaching C; sc-r absent; radial fork distal to medial fork; R
<subScript id="17AB34E0FFC8FF9BFBB111253B7D2754" attach="left" box="[1111,1119,347,361]" fontSize="6" pageId="2" pageNumber="489">4</subScript>
ending at wing tip; CuA not reaching wing margin. Male terminalia: hypandrium plate-like, subtriangular with microtrichia arranged in circular groups (
<figureCitation id="13142A20FFC8FF9BFF1511E83EA02792" box="[243,386,406,431]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Figs. 1E, 1F</figureCitation>
); length of gonocoxite 0.3 times the length of gonostylus; gonostyli narrow, basally inflated, bare, except in the base with patch of alveoli; aedeagus symmetrical, bifid, distally digitiform; two pairs of parameres, external pair fused forming a U-shaped plate; internal pair ending at the same place of aedeagus, well sclerotized, lanciform, ending in pointed apex; ejaculatory apodeme dorsoventrally flattened (
<figureCitation id="13142A20FFC8FF9BFB5E127C3A2F2426" box="[1208,1293,514,539]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Fig. 1E</figureCitation>
); epandrium 2.5 times wider than long, with two foramina and two distal patches of alveoli (
<figureCitation id="13142A20FFC8FF9BFBF512583B4E2402" box="[1043,1132,550,575]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Fig. 1G</figureCitation>
); cercus digitiform, 1.2 the length of epandrium; left cercus with 12 tenacula (
<figureCitation id="13142A20FFC8FF9BFD1612343C64245E" box="[752,838,586,611]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Fig. 1F</figureCitation>
) and right cercus with 11 tenacula; hypoproct and epiproct subtriangular with apical micropilosity (
<figureCitation id="13142A20FFC8FF9BFD5A12103C3724BA" box="[700,789,622,647]" captionStart="FIGURE 1" captionStartId="1.[151,250,1687,1709]" captionTargetBox="[151,1429,219,1666]" captionTargetId="figure@1.[151,1435,193,1666]" captionTargetPageId="1" captionText="FIGURE 1 (AH). Lepidiella maculosa sp. nov. (A) head; (B) base of antenna: scape, pedicel and three basal flagellomeres; (C) palpus; (D) wing; (E) male terminalia, dorsal; (F) male terminália, dorsal: hypandrium plate-like; (G) male terminalia, ventral: epandrium, cerci and hypoproct; (H) cercus. (Abbreviations: ae = aedeagus, ep = external paramere, ip = internal paramere)." httpUri="https://zenodo.org/record/2623145/files/figure.png" pageId="2" pageNumber="489">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="8B9036A5FFC8FF9BFF2112EC3EB42496" blockId="2.[151,1436,151,827]" box="[199,406,658,683]" pageId="2" pageNumber="489">Female: Unknown</paragraph>
</subSubSection>
<subSubSection id="C335652EFFC8FF9BFF2112C83D392507" pageId="2" pageNumber="489" type="discussion">
<paragraph id="8B9036A5FFC8FF9BFF2112C83D392507" blockId="2.[151,1436,151,827]" pageId="2" pageNumber="489">
<emphasis id="B95BEAB7FFC8FF9BFF2112C83E1924F2" bold="true" box="[199,315,694,719]" pageId="2" pageNumber="489">Remarks.</emphasis>
Males of five species of
<taxonomicName id="4C2F4D26FFC8FF9BFDB712C93DE124F2" authorityName="Enderlein" authorityYear="1937" box="[593,707,695,719]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEAB7FFC8FF9BFDB712C93DE124F2" box="[593,707,695,719]" italics="true" pageId="2" pageNumber="489">Lepidiella</emphasis>
</taxonomicName>
have a head without cornicula: the Brazilian species
<taxonomicName id="4C2F4D26FFC8FF9BFAF512C63EB624CE" authority="Bravo &amp; Santos, 2011" authorityName="Bravo &amp; Santos" authorityYear="2011" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="flabellata">
<emphasis id="B95BEAB7FFC8FF9BFAF512C63A0024F2" box="[1299,1314,696,719]" italics="true" pageId="2" pageNumber="489">L</emphasis>
.
<emphasis id="B95BEAB7FFC8FF9BFAD712C93ABE24F2" box="[1329,1436,695,719]" italics="true" pageId="2" pageNumber="489">flabellata</emphasis>
<bibRefCitation id="EFBE4B54FFC8FF9BFF7112A53EB624CE" author="Bravo, F. &amp; Santos, C. B." box="[151,404,731,755]" pageId="2" pageNumber="489" pagination="264 - 268" refId="ref4301" refString="Bravo, F. &amp; Santos, C. B. (2011) Two new species of Lepidiella (Diptera: Psychodidae: Psychodinae) from the Atlantic Rainforest of southeastern Brazil, Zoologia, 28, 264 - 268. https: // doi. org / 10.1590 / s 1984 - 46702011000200017" type="journal article" year="2011">Bravo &amp; Santos, 2011</bibRefCitation>
</taxonomicName>
, L. maculata
<emphasis id="B95BEAB7FFC8FF9BFDCA12A43DA424CE" bold="true" box="[556,646,730,755]" pageId="2" pageNumber="489">sp. nov.</emphasis>
,
<taxonomicName id="4C2F4D26FFC8FF9BFD7212A53CD424CE" authority="Bravo &amp; Araujo, 2013" authorityName="Bravo &amp; Araujo" authorityYear="2013" box="[660,1014,730,755]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="olgae">
<emphasis id="B95BEAB7FFC8FF9BFD7212A53D8124CF" box="[660,675,731,754]" italics="true" pageId="2" pageNumber="489">L</emphasis>
.
<emphasis id="B95BEAB7FFC8FF9BFD5512A53DD324CE" box="[691,753,731,755]" italics="true" pageId="2" pageNumber="489">olgae</emphasis>
<bibRefCitation id="EFBE4B54FFC8FF9BFD1F12A53CD424CE" author="Bravo, F. &amp; Araujo, M. X." box="[761,1014,730,755]" pageId="2" pageNumber="489" pagination="85 - 90" refId="ref4356" refString="Bravo, F. &amp; Araujo, M. X. (2013) Two new species of moth flies (Diptera, Psychodidae) from the semi-arid region of Brazil. Zootaxa, 3693 (1), 85 - 90. https: // doi. org / 10.11646 / zootaxa. 3693.1.6" type="journal article" year="2013">Bravo &amp; Araújo, 2013</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2F4D26FFC8FF9BFBC812A53A1A24CE" authority="Bravo, 2005" authorityName="Bravo" authorityYear="2005" box="[1070,1336,730,755]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="B95BEAB7FFC8FF9BFBC812A53B1F24CF" box="[1070,1085,731,754]" italics="true" pageId="2" pageNumber="489">L</emphasis>
.
<emphasis id="B95BEAB7FFC8FF9BFBAA12A53B8324CF" box="[1100,1185,731,754]" italics="true" pageId="2" pageNumber="489">spinosa</emphasis>
<bibRefCitation id="EFBE4B54FFC8FF9BFB4C12A53A1A24CE" author="Bravo, F." box="[1194,1336,730,755]" pageId="2" pageNumber="489" pagination="490 - 493" refId="ref4246" refString="Bravo, F. (2005) Primeiro registro de Lepidiella Enderlein (Diptera, Psychodidae, Psychodinae) no Brasil e descricao de uma especie nova. Revista Brasileira de Zoologia, 22 (2), 490 - 493. https: // doi. org / 10.1590 / S 0101 - 81752005000200027" type="journal article" year="2005">Bravo, 2005</bibRefCitation>
</taxonomicName>
, and one Costa Rican species,
<taxonomicName id="4C2F4D26FFC8FF9BFE6F137E3D8B252A" authority="(Quate, 1996)" authorityName="Quate" authorityYear="1996" baseAuthorityName="Quate" baseAuthorityYear="1996" box="[393,681,766,791]" class="Insecta" family="Psychodidae" genus="Lepidiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="hansoni">
<emphasis id="B95BEAB7FFC8FF9BFE6F137E3EBA252A" box="[393,408,768,791]" italics="true" pageId="2" pageNumber="489">L</emphasis>
.
<emphasis id="B95BEAB7FFC8FF9BFE4E12813D23252A" box="[424,513,767,791]" italics="true" pageId="2" pageNumber="489">hansoni</emphasis>
(
<bibRefCitation id="EFBE4B54FFC8FF9BFDF612813D82252A" author="Quate, L. W." box="[528,672,766,791]" pageId="2" pageNumber="489" pagination="1 - 81" refId="ref4937" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae, Revista de Biologia Tropical, 44 (Supl. 1), 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
)
</taxonomicName>
. The males of these five species can be recognized using the male identification key proposed below.
</paragraph>
</subSubSection>
</treatment>
</document>