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<document id="0A6E021908FE4ABCA33AB434490A7D33" ID-CLB-Dataset="53342" ID-DOI="10.11646/zootaxa.5292.1.1" ID-GBIF-Dataset="be05b7e2-ccc8-49cf-9173-1238edee4d40" ID-ISSN="1175-5326" ID-Zenodo-Dep="7959518" ID-ZooBank="8DEB1E66-92FA-4200-91A9-4631057B0600" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1684795458951" checkinUser="plazi" docAuthor="Chatzimanolis, Stylianos" docDate="2023" docId="038687A2FFE0FF96728614D6D8E8FED5" docLanguage="en" docName="zootaxa.5292.1.1.pdf" docOrigin="Zootaxa 5292 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5292.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Photinopygus koptopeos Chatzimanolis 2023, new species" docType="treatment" docUuid="C5B6EEEE-FE42-4530-A6FD-4D38052DD3D1" docUuidSource="ZooBank" docVersion="8" lastPageNumber="66" masterDocId="FFBFFFDAFFDFFFD77211157FDC73FF94" masterDocTitle="A revision of the genus Photinopygus Chatzimanolis (Staphylinidae: Xanthopygina)" masterLastPageNumber="100" masterPageNumber="1" pageNumber="64" updateTime="1699507402151" updateUser="plazi">
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<mods:titleInfo id="337441FD7A38331F13E31FF839BA36C0">
<mods:title id="36E1D26E229A048B283C6CC47A633AF8">A revision of the genus Photinopygus Chatzimanolis (Staphylinidae: Xanthopygina)</mods:title>
</mods:titleInfo>
<mods:name id="F8C68CE1DBBCE777D84DAF1A56EF99B3" type="personal">
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<mods:namePart id="720576EC6389B6C09E32FB46C0FF564F">Chatzimanolis, Stylianos</mods:namePart>
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<mods:title id="8E0873ED20EE8F41E2FE3489DFFD1BE4">Zootaxa</mods:title>
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<mods:part id="25BD817E1D34CE6E5B66FB069582019F">
<mods:date id="197B98ACC690CCBB9B8CFF630D36E3B1">2023</mods:date>
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<mods:number id="DCD5F41FA8417294CDB7056FB26788AC">2023-05-18</mods:number>
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<mods:classification id="DE7CDB726C21C8BF8DDB882C46A49A22">journal article</mods:classification>
<mods:identifier id="542B00B0DB3ABCF3EBE024C0ABE8E432" type="CLB-Dataset">53342</mods:identifier>
<mods:identifier id="145FE5AC6F586B9518AAA56B41F6A792" type="DOI">10.11646/zootaxa.5292.1.1</mods:identifier>
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<treatment id="038687A2FFE0FF96728614D6D8E8FED5" ID-DOI="http://doi.org/10.5281/zenodo.7959712" ID-GBIF-Taxon="209629576" ID-Zenodo-Dep="7959712" LSID="urn:lsid:zoobank.org:act:C5B6EEEE-FE42-4530-A6FD-4D38052DD3D1" httpUri="http://treatment.plazi.org/id/038687A2FFE0FF96728614D6D8E8FED5" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64">
<subSubSection id="C335653FFFE0FFE8728614D6DF6BFE57" box="[151,792,425,452]" pageId="63" pageNumber="64" type="nomenclature">
<paragraph id="8B9036B4FFE0FFE8728614D6DF6BFE57" blockId="63.[151,982,425,524]" box="[151,792,425,452]" pageId="63" pageNumber="64">
<heading id="D0D881D8FFE0FFE8728614D6DF6BFE57" bold="true" box="[151,792,425,452]" fontSize="11" level="2" pageId="63" pageNumber="64" reason="4">
<emphasis id="B95BEAA6FFE0FFE8728614D6DF6BFE57" bold="true" box="[151,792,425,452]" pageId="63" pageNumber="64">
<taxonomicName id="4C2F4D37FFE0FFE8728614D6DE09FE57" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2023" box="[151,634,425,452]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="koptopeos" status="sp. nov.">
<emphasis id="B95BEAA6FFE0FFE8728614D6DDCDFE50" bold="true" box="[151,446,425,452]" italics="true" pageId="63" pageNumber="64">Photinopygus koptopeos</emphasis>
Chatzimanolis
</taxonomicName>
,
<taxonomicNameLabel id="A26857DDFFE0FFE8709914D6DF6BFE57" box="[648,792,425,451]" pageId="63" pageNumber="64" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C335653FFFE0FFE8728614B2DFA5FD98" pageId="63" pageNumber="64" type="description">
<paragraph id="8B9036B4FFE0FFE8728614B2DD07FE7C" blockId="63.[151,982,425,524]" box="[151,372,461,488]" pageId="63" pageNumber="64">
(
<figureCitation id="13142A31FFE0FFE8728E14B2DC8EFE73" box="[159,253,461,487]" captionStart="FIGURE 39" captionStartId="20.[151,250,1841,1865]" captionTargetBox="[303,1285,1035,1818]" captionTargetId="figure-64@20.[303,1285,1035,1818]" captionTargetPageId="20" captionText="FIGURE 39. Distribution map of Photinopygus assingi Chatzimanolis (stars), P. chapareanus (Scheerpeltz) (triangles), P. faustus (Erichson) (circles), and P. koptopeos Chatzimanolis (squares)." figureDoi="http://doi.org/10.5281/zenodo.7959546" httpUri="https://zenodo.org/record/7959546/files/figure.png" pageId="63" pageNumber="64">Figs. 39</figureCitation>
,
<figureCitation id="13142A31FFE0FFE8731814B2DD1EFE7C" box="[265,365,461,488]" captionStart-0="FIGURES 153–157" captionStart-1="FIGURES 158–160" captionStartId-0="63.[151,264,1740,1764]" captionStartId-1="64.[151,264,962,986]" captionTargetBox-0="[287,1301,548,1713]" captionTargetBox-1="[151,1431,182,933]" captionTargetId-0="figure-102@63.[287,1301,547,1716]" captionTargetId-1="figure-30@64.[151,1436,181,938]" captionTargetPageId-0="63" captionTargetPageId-1="64" captionText-0="FIGURES 153–157. Photinopygus koptopeos Chatzimanolis. 153. Habitus. 154. Sternite 7. 155. Sternite 8. 156. Pronotum. 157. Antenna. Not to scale." captionText-1="FIGURES 158–160. Aedeagus of Photinopygus koptopeos Chatzimanolis. 158. Lateral (slightly oblique) view. 159. Dorsal view. 160. Detail of paramere, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.7959632" figureDoi-1="http://doi.org/10.5281/zenodo.7959634" httpUri-0="https://zenodo.org/record/7959632/files/figure.png" httpUri-1="https://zenodo.org/record/7959634/files/figure.png" pageId="63" pageNumber="64">153–160</figureCitation>
)
</paragraph>
<paragraph id="8B9036B4FFE0FFE87286148EDFA5FD98" blockId="63.[151,982,425,524]" box="[151,982,497,524]" pageId="63" pageNumber="64">
<uri id="FFBE3AB6FFE0FFE87286148EDFA5FD98" box="[151,982,497,524]" pageId="63" pageNumber="64">
urn:lsid:zoobank.org:act:
<uuid id="FF890C61FFE0FFE873B9148EDFA5FD98" box="[424,982,497,524]" pageId="63" pageNumber="64">C5B6EEEE-FE42-4530-A6FD-4D38052DD3D1</uuid>
</uri>
</paragraph>
</subSubSection>
<caption id="DF50663CFFE0FFE8728613B3DDD8F89D" ID-DOI="http://doi.org/10.5281/zenodo.7959632" ID-Zenodo-Dep="7959632" httpUri="https://zenodo.org/record/7959632/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,264,1740,1764]" targetBox="[287,1301,548,1713]" targetPageId="63" targetType="figure">
<paragraph id="8B9036B4FFE0FFE8728613B3DDD8F89D" blockId="63.[151,1437,1740,1801]" pageId="63" pageNumber="64">
<emphasis id="B95BEAA6FFE0FFE8728613B3DD03F971" bold="true" box="[151,368,1740,1765]" pageId="63" pageNumber="64">FIGURES 153–157.</emphasis>
<taxonomicName id="4C2F4D37FFE0FFE8736913B2DF67F971" authority="Chatzimanolis." authorityName="Chatzimanolis." box="[376,788,1741,1765]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="koptopeos">
<emphasis id="B95BEAA6FFE0FFE8736913B2DE01F971" box="[376,626,1741,1765]" italics="true" pageId="63" pageNumber="64">Photinopygus koptopeos</emphasis>
Chatzimanolis.
</taxonomicName>
153. Habitus. 154. Sternite 7. 155. Sternite 8. 156. Pronotum. 157. Antenna. Not to scale.
</paragraph>
</caption>
<subSubSection id="C335653FFFE0FFE872861231D8B5F86C" pageId="63" pageNumber="64" type="materials_examined">
<paragraph id="8B9036B4FFE0FFE872861231D8B5F86C" blockId="63.[151,1437,1869,2040]" pageId="63" pageNumber="64">
<emphasis id="B95BEAA6FFE0FFE872861231DD35F8F3" bold="true" box="[151,326,1869,1896]" pageId="63" pageNumber="64">Type material.</emphasis>
<materialsCitation id="3B473CE9FFE0FFE8735F1231D851F824" ID-GBIF-Occurrence="4116786631" collectingDate="1962-10" collectorName="In" country="Brazil" determinerName="A. Davies" location="Jatai" pageId="63" pageNumber="64" specimenCount="2" stateProvince="Goias" typeStatus="holotype">
<emphasis id="B95BEAA6FFE0FFE8735F1231DDC8F8FC" bold="true" box="[334,443,1870,1896]" pageId="63" pageNumber="64">
<typeStatus id="54948816FFE0FFE8735F1231DDC8F8FC" box="[334,443,1870,1896]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
</emphasis>
, here designated, male, with labels: “Faz. Cachoeirinha,
<location id="8EF0606FFFE0FFE876501231D807F8FC" LSID="urn:lsid:plazi:treatment:038687A2FFE0FF96728614D6D8E8FED5:8EF0606FFFE0FFE876501231D807F8FC" box="[1089,1140,1870,1896]" country="Brazil" name="Jatai" pageId="63" pageNumber="64" stateProvince="Goias">Jatai</location>
[-17.88°, -51.83°],
<collectingRegion id="49EBF856FFE0FFE877451231D9EBF8FC" box="[1364,1432,1870,1896]" country="Brazil" name="Goias" pageId="63" pageNumber="64">Goiás</collectingRegion>
, Brasil [
<collectingCountry id="F3387624FFE0FFE872F6120DDD40F818" box="[231,307,1906,1932]" name="Brazil" pageId="63" pageNumber="64">Brazil</collectingCountry>
],
<date id="FF911074FFE0FFE87355120EDDE0F81F" box="[324,403,1905,1931]" pageId="63" pageNumber="64" value="1962-10">
<collectingDate id="EFD5E99CFFE0FFE87355120EDDE0F81F" box="[324,403,1905,1931]" pageId="63" pageNumber="64" value="1962-10">x.1962</collectingDate>
</date>
, Exp. Dep. Zool.” / “
<taxonomicName id="4C2F4D37FFE0FFE8706A120DDFB4F818" authority="Ndm." authorityName="Ndm." box="[635,967,1906,1932]" class="Insecta" family="Staphylinidae" genus="Xanthopygus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="chrysurus">
<emphasis id="B95BEAA6FFE0FFE8706A120DDFF2F818" box="[635,897,1906,1932]" italics="true" pageId="63" pageNumber="64">Xanthopygus chrysurus</emphasis>
Ndm.
</taxonomicName>
det.
<determinerName id="7E4CDA70FFE0FFE871E8120DD819F818" box="[1017,1130,1906,1932]" pageId="63" pageNumber="64">A. Davies</determinerName>
” / “Holotype
<taxonomicName id="4C2F4D37FFE0FFE87713120DDDCBF824" authority="Chatzimanolis" authorityName="Chatzimanolis" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="koptopeos">
<emphasis id="B95BEAA6FFE0FFE87713120DDD7AF824" italics="true" pageId="63" pageNumber="64">Photinopygus koptopeos</emphasis>
Chatzimanolis
</taxonomicName>
, des.
<bibRefCitation id="EFBE4B45FFE0FFE873ED12E9DE96F824" author="Chatzimanolis, S." box="[508,741,1942,1968]" pageId="63" pageNumber="64" pagination="151 - 178" refId="ref72514" refString="Chatzimanolis, S. (2022) A revision of the genus Xanthopygus Kraatz. Zootaxa, 5099 (2), 151 - 178. https: // doi. org / 10.11646 / zootaxa. 5099.2.1" type="journal article" year="2022">Chatzimanolis 2022</bibRefCitation>
”.
<collectorName id="26DA5362FFE0FFE870EC12E9DF67F824" box="[765,788,1942,1968]" pageId="63" pageNumber="64">In</collectorName>
the collection of CNC
</materialsCitation>
.
<materialsCitation id="3B473CE9FFE0FFE8762112E9D8B1F86C" ID-GBIF-Occurrence="4116787009" collectingDate="1984-04" collectorName="F. M. Oliveira" country="Brazil" location="Alagoas" pageId="63" pageNumber="64" specimenCount="2" stateProvince="Alagoas" typeStatus="paratype">
<typeStatus id="54948816FFE0FFE8762112E9D8E6F824" box="[1072,1173,1942,1968]" pageId="63" pageNumber="64" type="paratype">Paratype</typeStatus>
, here designated, male with labels: “
<collectingCountry id="F3387624FFE0FFE8732512C5DD0DF840" box="[308,382,1978,2004]" name="Brazil" pageId="63" pageNumber="64">Brazil</collectingCountry>
:
<collectingRegion id="49EBF856FFE0FFE8739D12C5DD98F840" box="[396,491,1978,2004]" country="Brazil" name="Alagoas" pageId="63" pageNumber="64">Alagoas</collectingRegion>
, Sao Miguel dos Campos [-9.78°, -36.09°],
<date id="FF911074FFE0FFE871E712C5D83FF840" box="[1014,1100,1978,2004]" pageId="63" pageNumber="64" value="1984-04">
<collectingDate id="EFD5E99CFFE0FFE871E712C5D83FF840" box="[1014,1100,1978,2004]" pageId="63" pageNumber="64" value="1984-04">iv.1984</collectingDate>
</date>
,
<collectorName id="26DA5362FFE0FFE8764D12C5D976F840" box="[1116,1285,1978,2004]" pageId="63" pageNumber="64">F. M. Oliveira</collectorName>
” / “Paratype
<taxonomicName id="4C2F4D37FFE0FFE8728612A1DE25F86C" authority="Chatzimanolis" authorityName="Chatzimanolis" box="[151,598,2014,2040]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="koptopeos">
<emphasis id="B95BEAA6FFE0FFE8728612A1DDD9F86C" box="[151,426,2014,2040]" italics="true" pageId="63" pageNumber="64">Photinopygus koptopeos</emphasis>
Chatzimanolis
</taxonomicName>
, des.
<bibRefCitation id="EFBE4B45FFE0FFE8708512A1DF09F86C" author="Chatzimanolis, S." box="[660,890,2014,2040]" pageId="63" pageNumber="64" pagination="151 - 178" refId="ref72514" refString="Chatzimanolis, S. (2022) A revision of the genus Xanthopygus Kraatz. Zootaxa, 5099 (2), 151 - 178. https: // doi. org / 10.11646 / zootaxa. 5099.2.1" type="journal article" year="2022">Chatzimanolis 2022</bibRefCitation>
”. In the collection of CMNH
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF50663CFF9FFF97728616BDDE27FC6A" ID-DOI="http://doi.org/10.5281/zenodo.7959634" ID-Zenodo-Dep="7959634" httpUri="https://zenodo.org/record/7959634/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,264,962,986]" targetBox="[151,1431,182,933]" targetPageId="64" targetType="figure">
<paragraph id="8B9036B4FF9FFF97728616BDDE27FC6A" blockId="64.[151,1437,962,1022]" pageId="64" pageNumber="65">
<emphasis id="B95BEAA6FF9FFF97728616BDDD03FC4E" bold="true" box="[151,368,962,986]" pageId="64" pageNumber="65">FIGURES 158–160.</emphasis>
Aedeagus of
<taxonomicName id="4C2F4D37FF9FFF97701316BDDFECFC4E" authority="Chatzimanolis." authorityName="Chatzimanolis." box="[514,927,962,986]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="koptopeos">
<emphasis id="B95BEAA6FF9FFF97701316BDDE8EFC4E" box="[514,765,962,986]" italics="true" pageId="64" pageNumber="65">Photinopygus koptopeos</emphasis>
Chatzimanolis.
</taxonomicName>
158. Lateral (slightly oblique) view. 159. Dorsal view. 160. Detail of paramere, ventral view.
</paragraph>
</caption>
<subSubSection id="C335653FFF9FFF9772D6113DD861FB7F" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="8B9036B4FF9FFF9772D6113DD861FB7F" blockId="64.[151,1437,1090,2040]" pageId="64" pageNumber="65">
<emphasis id="B95BEAA6FF9FFF9772D6113DDD31FBC8" bold="true" box="[199,322,1090,1116]" pageId="64" pageNumber="65">Diagnosis.</emphasis>
<taxonomicName id="4C2F4D37FF9FFF97735D113DDE11FBC8" box="[332,610,1090,1116]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="koptopeos">
<emphasis id="B95BEAA6FF9FFF97735D113DDE11FBC8" box="[332,610,1090,1116]" italics="true" pageId="64" pageNumber="65">Photinopygus koptopeos</emphasis>
</taxonomicName>
belongs in the narrow pronotum species group. Among species of that group that have pronotum with sparse punctation (i.e., with 3–5 loosely organized rows of punctures on each half beside median impunctate line), and antennomeres 4–5 longer than wide,
<taxonomicName id="4C2F4D37FF9FFF9771F711F5D806FB37" box="[998,1141,1161,1187]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="koptopeos">
<emphasis id="B95BEAA6FF9FFF9771F711F5D806FB37" box="[998,1141,1161,1187]" italics="true" pageId="64" pageNumber="65">P. koptopeos</emphasis>
</taxonomicName>
can be recognized by the following combination of characters: tip of paramere in dorsal view rounded (
<figureCitation id="13142A31FF9FFF9771EF11D2D82EFB5C" box="[1022,1117,1197,1224]" captionStart="FIGURES 158–160" captionStartId="64.[151,264,962,986]" captionTargetBox="[151,1431,182,933]" captionTargetId="figure-30@64.[151,1436,181,938]" captionTargetPageId="64" captionText="FIGURES 158–160. Aedeagus of Photinopygus koptopeos Chatzimanolis. 158. Lateral (slightly oblique) view. 159. Dorsal view. 160. Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7959634" httpUri="https://zenodo.org/record/7959634/files/figure.png" pageId="64" pageNumber="65">Fig. 159</figureCitation>
) median lobe in lateral view with lateral teeth and converging to a tip without hook-like structure (
<figureCitation id="13142A31FF9FFF9771B211AFD875FB7F" box="[931,1030,1232,1259]" captionStart="FIGURES 158–160" captionStartId="64.[151,264,962,986]" captionTargetBox="[151,1431,182,933]" captionTargetId="figure-30@64.[151,1436,181,938]" captionTargetPageId="64" captionText="FIGURES 158–160. Aedeagus of Photinopygus koptopeos Chatzimanolis. 158. Lateral (slightly oblique) view. 159. Dorsal view. 160. Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7959634" httpUri="https://zenodo.org/record/7959634/files/figure.png" pageId="64" pageNumber="65">Fig. 158</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335653FFF9FFF9772D6118BD8EDF8FD" pageId="64" pageNumber="65" type="description">
<paragraph id="8B9036B4FF9FFF9772D6118BD8EDF8FD" blockId="64.[151,1437,1090,2040]" pageId="64" pageNumber="65">
<emphasis id="B95BEAA6FF9FFF9772D6118BDD2AFA9A" bold="true" box="[199,345,1268,1294]" pageId="64" pageNumber="65">Description.</emphasis>
Forebody (
<figureCitation id="13142A31FF9FFF9773F4118BDE39FA9B" box="[485,586,1268,1295]" captionStart="FIGURES 153–157" captionStartId="63.[151,264,1740,1764]" captionTargetBox="[287,1301,548,1713]" captionTargetId="figure-102@63.[287,1301,547,1716]" captionTargetPageId="63" captionText="FIGURES 153–157. Photinopygus koptopeos Chatzimanolis. 153. Habitus. 154. Sternite 7. 155. Sternite 8. 156. Pronotum. 157. Antenna. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7959632" httpUri="https://zenodo.org/record/7959632/files/figure.png" pageId="64" pageNumber="65">Fig. 153</figureCitation>
) length
<quantity id="4CD79B51FF9FFF9770A0118BDF33FA9A" box="[689,832,1268,1295]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.6" metricValueMin="4.9" pageId="64" pageNumber="65" unit="mm" value="5.25" valueMax="5.6" valueMin="4.9">4.9–5.6 mm</quantity>
. Color of head, pronotum and mesoscutellum dark brown to black; antennae dark brown to black; legs dark brown to black; elytra metallic blue with green or purple overtones; abdomen dark brown to black except segments 7–8 (orange). Antenna (
<figureCitation id="13142A31FF9FFF9776481044D8CFFAC2" box="[1113,1212,1339,1366]" captionStart="FIGURES 153–157" captionStartId="63.[151,264,1740,1764]" captionTargetBox="[287,1301,548,1713]" captionTargetId="figure-102@63.[287,1301,547,1716]" captionTargetPageId="63" captionText="FIGURES 153–157. Photinopygus koptopeos Chatzimanolis. 153. Habitus. 154. Sternite 7. 155. Sternite 8. 156. Pronotum. 157. Antenna. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7959632" httpUri="https://zenodo.org/record/7959632/files/figure.png" pageId="64" pageNumber="65">Fig. 157</figureCitation>
) with antennomere 3 without tomentose pubescence; antennomere 4 with tomentose pubescence; antennomeres 4–5 longer than wide; antennomeres 6–7 subquadrate; antennomeres 8–10 transverse. Head transverse; HW/HL ratio = 1.42–1.43. Posterior margin of head slightly extended posteriad on each side of neck. Head with medium-sized punctures, distance between punctures as wide as 1–2 punctures but punctures denser posteriorly. Left mandible with bicuspid tooth. Pronotum (
<figureCitation id="13142A31FF9FFF9773731093DDB6F993" box="[354,453,1516,1543]" captionStart="FIGURES 153–157" captionStartId="63.[151,264,1740,1764]" captionTargetBox="[287,1301,548,1713]" captionTargetId="figure-102@63.[287,1301,547,1716]" captionTargetPageId="63" captionText="FIGURES 153–157. Photinopygus koptopeos Chatzimanolis. 153. Habitus. 154. Sternite 7. 155. Sternite 8. 156. Pronotum. 157. Antenna. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7959632" httpUri="https://zenodo.org/record/7959632/files/figure.png" pageId="64" pageNumber="65">Fig. 156</figureCitation>
) subquadrate; PW/PL ratio = 1.06–1.12. Lateral margins of pronotum in dorsal view posteriad of midpoint strongly converging; pronotum with 3–4 sparse rows of punctures on each half beside median impunctate line; distance between punctures as wide as 1–2 punctures but areas of pronotum without punctures. EL/PL ratio = 1.47–1.75. Elytra with dense punctation; distance between punctures as wide as 0.5–1 punctures. Metepisternum covered with punctures (impunctate area less than 1/3). Abdomen with tergites 3–4 setose; tergites 3–5 with curved carina (arch-like). In males, sternite 7 without porose structure, sternite 7 with shallow and broad emargination posteriorly; sternite 8 with small V-shaped emargination posteriorly (
<figureCitation id="13142A31FF9FFF97762C13BED8D3F948" box="[1085,1184,1729,1756]" captionStart="FIGURES 153–157" captionStartId="63.[151,264,1740,1764]" captionTargetBox="[287,1301,548,1713]" captionTargetId="figure-102@63.[287,1301,547,1716]" captionTargetPageId="63" captionText="FIGURES 153–157. Photinopygus koptopeos Chatzimanolis. 153. Habitus. 154. Sternite 7. 155. Sternite 8. 156. Pronotum. 157. Antenna. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7959632" httpUri="https://zenodo.org/record/7959632/files/figure.png" pageId="64" pageNumber="65">Fig. 154</figureCitation>
). Aedeagus as in
<figureCitation id="13142A31FF9FFF97777413BEDC8FF96B" captionStart="FIGURES 158–160" captionStartId="64.[151,264,962,986]" captionTargetBox="[151,1431,182,933]" captionTargetId="figure-30@64.[151,1436,181,938]" captionTargetPageId="64" captionText="FIGURES 158–160. Aedeagus of Photinopygus koptopeos Chatzimanolis. 158. Lateral (slightly oblique) view. 159. Dorsal view. 160. Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7959634" httpUri="https://zenodo.org/record/7959634/files/figure.png" pageId="64" pageNumber="65">Figs. 158–160</figureCitation>
; in dorsal view paramere becoming narrower before expanding to broad rounded apex; paramere shorter and as wide than median lobe; in lateral (slightly oblique) view paramere expanding apically; paramere with peg setae in lateral rows as in
<figureCitation id="13142A31FF9FFF9773A91254DE6BF8D1" box="[440,536,1835,1861]" captionStart="FIGURES 158–160" captionStartId="64.[151,264,962,986]" captionTargetBox="[151,1431,182,933]" captionTargetId="figure-30@64.[151,1436,181,938]" captionTargetPageId="64" captionText="FIGURES 158–160. Aedeagus of Photinopygus koptopeos Chatzimanolis. 158. Lateral (slightly oblique) view. 159. Dorsal view. 160. Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7959634" httpUri="https://zenodo.org/record/7959634/files/figure.png" pageId="64" pageNumber="65">Fig. 160</figureCitation>
. Median lobe in dorsal view gradually converging to rounded tip; in lateral view (slightly oblique) median lobe becoming narrower; median lobe with two large lateral teeth.
</paragraph>
</subSubSection>
<subSubSection id="C335653FFF9FFF9772D6120DDEE5F824" pageId="64" pageNumber="65" type="distribution">
<paragraph id="8B9036B4FF9FFF9772D6120DDEE5F824" blockId="64.[151,1437,1090,2040]" pageId="64" pageNumber="65">
<emphasis id="B95BEAA6FF9FFF9772D6120DDD12F818" bold="true" box="[199,353,1906,1932]" pageId="64" pageNumber="65">Distribution.</emphasis>
Known from Chacoan and
<collectingRegion id="49EBF856FF9FFF97708B120DDE95F818" box="[666,742,1906,1932]" country="Brazil" name="Parana" pageId="64" pageNumber="65">Paraná</collectingRegion>
biogeographic dominion. Distributed in the states of
<collectingRegion id="49EBF856FF9FFF977751120DD9EFF818" box="[1344,1436,1906,1932]" country="Brazil" name="Alagoas" pageId="64" pageNumber="65">Alagoas</collectingRegion>
and
<collectingRegion id="49EBF856FF9FFF9772D612E9DD7BF824" box="[199,264,1942,1968]" country="Brazil" name="Goias" pageId="64" pageNumber="65">Goiás</collectingRegion>
in
<collectingCountry id="F3387624FF9FFF97733D12E9DD07F824" box="[300,372,1942,1968]" name="Brazil" pageId="64" pageNumber="65">Brazil</collectingCountry>
. Map is shown in
<figureCitation id="13142A31FF9FFF97702E12E9DEE2F824" box="[575,657,1942,1968]" captionStart="FIGURE 39" captionStartId="20.[151,250,1841,1865]" captionTargetBox="[303,1285,1035,1818]" captionTargetId="figure-64@20.[303,1285,1035,1818]" captionTargetPageId="20" captionText="FIGURE 39. Distribution map of Photinopygus assingi Chatzimanolis (stars), P. chapareanus (Scheerpeltz) (triangles), P. faustus (Erichson) (circles), and P. koptopeos Chatzimanolis (squares)." figureDoi="http://doi.org/10.5281/zenodo.7959546" httpUri="https://zenodo.org/record/7959546/files/figure.png" pageId="64" pageNumber="65">Fig. 39</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C335653FFF9FFF9672D612C6D921FF25" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="etymology">
<paragraph id="8B9036B4FF9FFF9772D612C6DF5BF863" blockId="64.[151,1437,1090,2040]" pageId="64" pageNumber="65">
<emphasis id="B95BEAA6FF9FFF9772D612C6DD3DF847" bold="true" box="[199,334,1977,2003]" pageId="64" pageNumber="65">Etymology.</emphasis>
The specific epithet is derived from the Greek words κόπτω (to cut) and πέος (penis) and refers to the shape of the median lobe that appears to be broken off.
</paragraph>
<paragraph id="8B9036B4FF9EFF9672D615E9D921FF25" blockId="65.[151,1437,150,321]" box="[199,1362,150,177]" pageId="65" pageNumber="66">
<emphasis id="B95BEAA6FF9EFF9672D615E9DD59FF24" bold="true" box="[199,298,150,176]" pageId="65" pageNumber="66">Habitat.</emphasis>
Unknown but specimens have been collected from imprecise localities ranging from
<quantity id="4CD79B51FF9EFF9676F915E8D93DFF25" box="[1256,1358,151,177]" metricMagnitude="2" metricUnit="m" metricValue="4.01" metricValueMax="8.0" metricValueMin="0.02" pageId="65" pageNumber="66" unit="m" value="401.0" valueMax="800.0" valueMin="2.0">2–800 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C335653FFF9EFF9672D615C4D8E8FED5" pageId="65" pageNumber="66" type="discussion">
<paragraph id="8B9036B4FF9EFF9672D615C4D8E8FED5" blockId="65.[151,1437,150,321]" pageId="65" pageNumber="66">
<emphasis id="B95BEAA6FF9EFF9672D615C4DD48FF41" bold="true" box="[199,315,187,213]" pageId="65" pageNumber="66">Remarks.</emphasis>
The aedeagus of this species is remarkable, and unmistakable among
<taxonomicName id="4C2F4D37FF9EFF96765215C4D8AEFF41" authorityName="Chatzimanolis" authorityYear="2021" box="[1091,1245,187,213]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEAA6FF9EFF96765215C4D8AEFF41" box="[1091,1245,187,213]" italics="true" pageId="65" pageNumber="66">Photinopygus</emphasis>
</taxonomicName>
; it is reminiscent of the aedeagi found in
<emphasis id="B95BEAA6FF9EFF9673B215A0DEBBFF6D" box="[419,712,223,249]" italics="true" pageId="65" pageNumber="66">
<taxonomicName id="4C2F4D37FF9EFF9673B215A0DE62FF6C" box="[419,529,223,248]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="assingi">P. assingi</taxonomicName>
,
<taxonomicName id="4C2F4D37FF9EFF96700D15A0DEBBFF6D" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1969" box="[540,712,223,249]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="chapareanus">P. chapareanus</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C2F4D37FF9EFF96711715A0DF02FF6D" baseAuthorityName="Erichson" baseAuthorityYear="1839" box="[774,881,223,249]" class="Insecta" family="Staphylinidae" genus="Photinopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="faustus">
<emphasis id="B95BEAA6FF9EFF96711715A0DF02FF6D" box="[774,881,223,249]" italics="true" pageId="65" pageNumber="66">P. faustus</emphasis>
</taxonomicName>
but without the hook-like tip of the median lobe. When I dissected the first specimen, I thought that the tip of the median lobe was broken and only after I saw the second specimen with the same aedeagus I was convinced that this was the entire aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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