163 lines
23 KiB
XML
163 lines
23 KiB
XML
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<mods:title id="403E647A54D4D725534B0D5897085854">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part VII. Parabuthus Pocock, 1890 (Buthidae) with description of P. hamar sp. n. and P. kajibu sp. n. from Ethiopia</mods:title>
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<mods:namePart id="8746D9182612B933A2A50D83B96A01A2">Kovařík, František</mods:namePart>
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<mods:namePart id="CFBB7FD6D595459104426F000631AD65">Lowe, Graeme</mods:namePart>
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<mods:namePart id="ECECBEBAA70B92F8203708E9D263054E">Plíšková, Jana</mods:namePart>
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<mods:namePart id="9CA4F7C1523A1D39CBD4A083DF4E847A">Šťáhlavský, František</mods:namePart>
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<taxonomicName id="4C2F4D1FFFC4DC5AFEAF6CA5F69D30FB" ID-CoL="4CN4W" authority="Kovarik, 2004" authorityName="Kovarik" authorityYear="2004" box="[295,699,939,962]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="cimrmani">
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<emphasis id="B95BEA8EFFC4DC5AFEAF6CA5F63C30FB" bold="true" box="[295,538,939,962]" italics="true" pageId="20" pageNumber="19">Parabuthus cimrmani</emphasis>
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Kovařík, 2004
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<paragraph id="8B90369CFFC4DC5AFE846CC9F6F330E7" blockId="20.[268,725,939,990]" box="[268,725,967,990]" pageId="20" pageNumber="19">
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(
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<figureCitation id="13142A19FFC4DC5AFE9C6CC9F5B030E7" box="[276,406,967,990]" captionStart="Figures 36–39" captionStartId="14.[192,278,1824,1847]" captionTargetBox="[204,1425,199,1775]" captionTargetId="figure-14@14.[192,1439,192,1795]" captionTargetPageId="14" captionText="Figures 36–39: Parabuthus cimrmani. Figures 36–37. Male holotype in dorsal (36) and ventral (37) views. Figures 38–39: Female paratype in dorsal (38) and ventral (39) aspects. Scale bar: 10 mm." pageId="20" pageNumber="19">Figs. 36–39</figureCitation>
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,
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<figureCitation id="13142A19FFC4DC5AFE286CC9F59B30E7" box="[416,445,967,990]" captionStart="Figures 71–89" captionStartId="26.[192,278,1661,1684]" captionTargetBox="[204,1433,246,1645]" captionTargetId="figure-14@26.[192,1439,233,1656]" captionTargetPageId="26" captionText="Figures 71–89: Figures 71–83. Parabuthus hamar sp. n., pedipalp segments. Figures 71–72, 83. Female paratype from locality 16EH, chela dorsal (71) and external (72). Patella dorsal (83). Figures 73–82. Male paratype from locality 16EC, chela dorsal (73), external (74), and ventral (75). Patella dorsal (76), external (77) and ventral (78). Femur internal (79) and femur and trochanter dorsal (80). Movable finger (81) and fixed finger (82) dentition. Trichobothrial pattern is indicated in Figures 74–77 and 79–80. Figures 84–85. P. kajibu sp. n., male holotype, movable finger (84) and fixed finger (85) dentition. Figure 86. P. maximus, male from Kenya, Voi, movable finger dentition. Figure 87. P. pallidus, male from locality 16EA, movable finger dentition. Figure 88. P. cimrmani, female paratype, movable finger dentition. Figure 89. P. eritreaensis, male from Somalia, Gardo, Migiurtina, movable finger dentition." pageId="20" pageNumber="19">88</figureCitation>
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,
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<figureCitation id="13142A19FFC4DC5AFE406CC9F60E30E7" box="[456,552,967,990]" captionStart="Figures 164–182" captionStartId="50.[192,278,1614,1637]" captionTargetBox="[200,1421,197,1572]" captionTargetId="figure-37@50.[192,1439,192,1585]" captionTargetPageId="50" captionText="Figures 164–182: Figures 164–179. Comparison of male pedipalp chelae of Parabuthus spp., dorsal and external views. Figures 164–165. Parabuthus liosoma from Yemen Arab Republic, Al Hudaydah gov., 10 km W Al Manşuriah, 14°43'N 43°12'E, 110 m a.s.l. Figures 166–167. P. abyssinicus from Eritrea, locality 15EG. Figures 168–169. P. hamar sp. n. paratype from Ethiopia, locality 16EC. Figures 170–171. P. granimanus from Somaliland, locality 11SE. Figures 172–173. P. cimrmani holotype. Figures 174–175. P. eritreaensis from Somalia, Gardo, Migiurtina. Figures 176–177. P. heterurus from Somaliland, locality 11SQ. Figures 178–179. P. kajibu sp. n. holotype. Figures 180–182. Comparisons of male pedipalp patellae of Parabuthus spp., dorsal views. Figure 180. P. liosoma from Yemen Arab Republic, Al Hudaydah gov., 10 km W Al Manşuriah, 14°43'N 43°12'E, 110 m a.s.l. Figure 181. P. abyssinicus from Eritrea, locality 15EG. Figure 182. P. maximus from Tanzania, Arusha." pageId="20" pageNumber="19">172–173</figureCitation>
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,
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<figureCitation id="13142A19FFC4DC5AFDBB6CC9F67B30E7" box="[563,605,967,990]" captionStart="Figures 183–192" captionStartId="52.[192,278,1762,1785]" captionTargetBox="[201,1434,181,1732]" captionTargetId="figure-47@52.[192,1439,168,1732]" captionTargetPageId="52" captionText="Figures 183–192: Comparison of male metasomal segments I–III of Parabuthus spp., dorsal views. Figure 183. Parabuthus liosoma from Yemen Arab Republic, Al Hudaydah gov., 10 km W Al Manşuriah, 14°43'N 43°12'E, 110 m a.s.l. Figure 184. P. maximus from Kenya, Voi. Figure 185. P. abyssinicus from Ethiopia, locality 11EA. Figure 186. P. granimanus from Somaliland, locality 11SE. Figure 187. P. hamar sp. n. holotype. Figure 188. P. cimrmani holotype. Figure 189. P. eritreaensis from Somalia, Gardo, Migiurtina. Figure 190. P. heterurus from Somaliland, locality 11SQ. Figure 191. P. kajibu sp. n. holotype. Figure 192. P. pallidus from Ethiopia, locality 16EA." pageId="20" pageNumber="19">188</figureCitation>
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,
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<figureCitation id="13142A19FFC4DC5AFDE06CC9F6B430E7" box="[616,658,967,990]" captionStart="Figures 193–197" captionStartId="53.[192,278,1736,1759]" captionTargetBox="[196,1429,245,1696]" captionTargetId="figure-25@53.[192,1439,237,1704]" captionTargetPageId="53" captionText="Figures 193–197: Comparison of male metasomal segments III–V and telson of Parabuthus spp., lateral views. Figure 193. Parabuthus liosoma from Yemen Arab Republic, Al Hudaydah gov., 10 km W Al Manşuriah, 14°43'N 43°12'E, 110 m a.s.l. Figure 194. P. abyssinicus from Eritrea, locality 15EG. Figure 195. P. hamar sp. n. paratype from Ethiopia, locality 16EC. Figure 196. P. granimanus from Somaliland, locality 11SE. Figure 197. P. cimrmani holotype." pageId="20" pageNumber="19">197</figureCitation>
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,
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<figureCitation id="13142A19FFC4DC5AFD156CC9F6E330E7" box="[669,709,967,990]" captionStart="Figure 204" captionStartId="56.[192,268,1220,1243]" captionTargetBox="[196,1436,197,1186]" captionTargetId="figure-99@56.[192,1439,192,1190]" captionTargetPageId="56" captionText="Figure 204: Map showing confirmed distribution of Parabuthus spp. in Djibouti, Eritrea, Ethiopia, Somaliland and P. eritreaensis in Somalia (Puntland). Most points indicate sites sampled during 2011–2016 expeditions. We have not included in the map locality data for old specimens mainly from Somalia deposited in Italian museums which include large collections from this region (Kovařík & Whitman, 2005: 110–112), because these specimens are often mislabeled. Without corroboration from recently collected specimens, it is problematic to verify which of the locality labels are correct." pageId="20" pageNumber="19">204</figureCitation>
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)
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</paragraph>
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<taxonomicName id="4C2F4D1FFFC4DC5AFF486CF0F68A372C" ID-CoL="4CN4W" authority="Kovarik, 2004: 15 - 19" authorityName="Kovarik" authorityPageNumber="15 - 19" authorityYear="2004" box="[192,684,1022,1045]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="cimrmani">
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<emphasis id="B95BEA8EFFC4DC5AFF486CF0F58B372C" box="[192,429,1022,1045]" italics="true" pageId="20" pageNumber="19">Parabuthus cimrmani</emphasis>
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Kovařík, 2004: 15–19
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</taxonomicName>
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, figs. 1–5, table 1.
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</paragraph>
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<subSubSection id="C3356517FFC4DC5AFF486B5FF65437D6" pageId="20" pageNumber="19" type="materials_examined">
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<paragraph id="8B90369CFFC4DC5AFF486B5FF5B837BD" blockId="20.[192,802,1105,1156]" pageId="20" pageNumber="19">
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<materialsCitation id="3B473CC1FFC4DC5AFF486B5FF5983751" box="[192,446,1105,1128]" collectionCode="LOCALITY, AND" pageId="20" pageNumber="19" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5494883EFFC4DC5AFF486B5FF4DE3751" box="[192,248,1105,1128]" pageId="20" pageNumber="19">TYPE</typeStatus>
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<collectionCode id="ED3EAE59FFC4DC5AFE876B5BF5983751" box="[271,446,1109,1128]" pageId="20" pageNumber="19">LOCALITY AND</collectionCode>
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<materialsCitation id="3B473CC1FFC4DC5AFE5D6B5BF57737BD" collectionCode="REPOSITORY" country="Somalia" location="Maxaans" pageId="20" pageNumber="19" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5494883EFFC4DC5AFE5D6B5BF62C3751" box="[469,522,1109,1128]" pageId="20" pageNumber="19">TYPE</typeStatus>
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<collectionCode id="ED3EAE59FFC4DC5AFDA96B5BF6833751" box="[545,677,1109,1128]" pageId="20" pageNumber="19">REPOSITORY</collectionCode>
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.
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<collectingCountry id="F338760CFFC4DC5AFD4B6B5FF73B3751" box="[707,797,1105,1128]" name="Somalia" pageId="20" pageNumber="19">Somalia</collectingCountry>
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,
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<location id="8EF06047FFC4DC5AFF486B63F50637BD" LSID="urn:lsid:plazi:treatment:0386878AFFC4DC5AFEAF6CA5F5E335F8:8EF06047FFC4DC5AFF486B63F50637BD" box="[192,288,1133,1156]" country="Somalia" name="Maxaans" pageId="20" pageNumber="19">Maxaans</location>
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env
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</materialsCitation>
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;
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<collectionCode id="ED3EAE59FFC4DC5AFED36B63F5B837BD" box="[347,414,1133,1156]" pageId="20" pageNumber="19">FKCP</collectionCode>
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<materialsCitation id="3B473CC1FFC4DC5AFF486BAFF63A37D6" collectionCode="TYPE, MATERIAL, EXAMINED" country="Somalia" location="Maxaans" pageId="20" pageNumber="19" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
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<collectionCode id="ED3EAE59FFC4DC5AFF486BAFF5D3378E" box="[192,501,1185,1208]" pageId="20" pageNumber="19">TYPE MATERIAL EXAMINED</collectionCode>
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.
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<emphasis id="B95BEA8EFFC4DC5AFD846BAFF64E3781" bold="true" box="[524,616,1185,1208]" pageId="20" pageNumber="19">
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<collectingCountry id="F338760CFFC4DC5AFD846BAFF64E3781" box="[524,616,1185,1208]" name="Somalia" pageId="20" pageNumber="19">Somalia</collectingCountry>
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</emphasis>
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,
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<location id="8EF06047FFC4DC5AFDF66BAFF6F83781" LSID="urn:lsid:plazi:treatment:0386878AFFC4DC5AFEAF6CA5F5E335F8:8EF06047FFC4DC5AFDF66BAFF6F83781" box="[638,734,1185,1208]" country="Somalia" name="Maxaans" pageId="20" pageNumber="19">Maxaans</location>
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env., 1972,
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<specimenCount id="9D29FD15FFC4DC5AFE856BB2F50937EA" box="[269,303,1212,1235]" pageId="20" pageNumber="19" type="male">1♂</specimenCount>
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<specimenCount id="9D29FD15FFC4DC5AFEA76BB2F57637EA" box="[303,336,1212,1235]" pageId="20" pageNumber="19" type="female">1♀</specimenCount>
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(
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<figureCitation id="13142A19FFC4DC5AFEE16BB2F5D337EA" box="[361,501,1212,1235]" captionStart="Figures 36–39" captionStartId="14.[192,278,1824,1847]" captionTargetBox="[204,1425,199,1775]" captionTargetId="figure-14@14.[192,1439,192,1795]" captionTargetPageId="14" captionText="Figures 36–39: Parabuthus cimrmani. Figures 36–37. Male holotype in dorsal (36) and ventral (37) views. Figures 38–39: Female paratype in dorsal (38) and ventral (39) aspects. Scale bar: 10 mm." pageId="20" pageNumber="19">Figs. 36–39</figureCitation>
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,
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<figureCitation id="13142A19FFC4DC5AFD836BB2F60E37EA" box="[523,552,1212,1235]" captionStart="Figures 71–89" captionStartId="26.[192,278,1661,1684]" captionTargetBox="[204,1433,246,1645]" captionTargetId="figure-14@26.[192,1439,233,1656]" captionTargetPageId="26" captionText="Figures 71–89: Figures 71–83. Parabuthus hamar sp. n., pedipalp segments. Figures 71–72, 83. Female paratype from locality 16EH, chela dorsal (71) and external (72). Patella dorsal (83). Figures 73–82. Male paratype from locality 16EC, chela dorsal (73), external (74), and ventral (75). Patella dorsal (76), external (77) and ventral (78). Femur internal (79) and femur and trochanter dorsal (80). Movable finger (81) and fixed finger (82) dentition. Trichobothrial pattern is indicated in Figures 74–77 and 79–80. Figures 84–85. P. kajibu sp. n., male holotype, movable finger (84) and fixed finger (85) dentition. Figure 86. P. maximus, male from Kenya, Voi, movable finger dentition. Figure 87. P. pallidus, male from locality 16EA, movable finger dentition. Figure 88. P. cimrmani, female paratype, movable finger dentition. Figure 89. P. eritreaensis, male from Somalia, Gardo, Migiurtina, movable finger dentition." pageId="20" pageNumber="19">88</figureCitation>
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,
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<figureCitation id="13142A19FFC4DC5AFDB56BB2F6B837EA" box="[573,670,1212,1235]" captionStart="Figures 164–182" captionStartId="50.[192,278,1614,1637]" captionTargetBox="[200,1421,197,1572]" captionTargetId="figure-37@50.[192,1439,192,1585]" captionTargetPageId="50" captionText="Figures 164–182: Figures 164–179. Comparison of male pedipalp chelae of Parabuthus spp., dorsal and external views. Figures 164–165. Parabuthus liosoma from Yemen Arab Republic, Al Hudaydah gov., 10 km W Al Manşuriah, 14°43'N 43°12'E, 110 m a.s.l. Figures 166–167. P. abyssinicus from Eritrea, locality 15EG. Figures 168–169. P. hamar sp. n. paratype from Ethiopia, locality 16EC. Figures 170–171. P. granimanus from Somaliland, locality 11SE. Figures 172–173. P. cimrmani holotype. Figures 174–175. P. eritreaensis from Somalia, Gardo, Migiurtina. Figures 176–177. P. heterurus from Somaliland, locality 11SQ. Figures 178–179. P. kajibu sp. n. holotype. Figures 180–182. Comparisons of male pedipalp patellae of Parabuthus spp., dorsal views. Figure 180. P. liosoma from Yemen Arab Republic, Al Hudaydah gov., 10 km W Al Manşuriah, 14°43'N 43°12'E, 110 m a.s.l. Figure 181. P. abyssinicus from Eritrea, locality 15EG. Figure 182. P. maximus from Tanzania, Arusha." pageId="20" pageNumber="19">172–173</figureCitation>
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,
|
||
<figureCitation id="13142A19FFC4DC5AFD3B6BB2F6F837EA" box="[691,734,1212,1235]" captionStart="Figures 183–192" captionStartId="52.[192,278,1762,1785]" captionTargetBox="[201,1434,181,1732]" captionTargetId="figure-47@52.[192,1439,168,1732]" captionTargetPageId="52" captionText="Figures 183–192: Comparison of male metasomal segments I–III of Parabuthus spp., dorsal views. Figure 183. Parabuthus liosoma from Yemen Arab Republic, Al Hudaydah gov., 10 km W Al Manşuriah, 14°43'N 43°12'E, 110 m a.s.l. Figure 184. P. maximus from Kenya, Voi. Figure 185. P. abyssinicus from Ethiopia, locality 11EA. Figure 186. P. granimanus from Somaliland, locality 11SE. Figure 187. P. hamar sp. n. holotype. Figure 188. P. cimrmani holotype. Figure 189. P. eritreaensis from Somalia, Gardo, Migiurtina. Figure 190. P. heterurus from Somaliland, locality 11SQ. Figure 191. P. kajibu sp. n. holotype. Figure 192. P. pallidus from Ethiopia, locality 16EA." pageId="20" pageNumber="19">188</figureCitation>
|
||
,
|
||
<figureCitation id="13142A19FFC4DC5AFD7B6BB2F73837EA" box="[755,798,1212,1235]" captionStart="Figures 193–197" captionStartId="53.[192,278,1736,1759]" captionTargetBox="[196,1429,245,1696]" captionTargetId="figure-25@53.[192,1439,237,1704]" captionTargetPageId="53" captionText="Figures 193–197: Comparison of male metasomal segments III–V and telson of Parabuthus spp., lateral views. Figure 193. Parabuthus liosoma from Yemen Arab Republic, Al Hudaydah gov., 10 km W Al Manşuriah, 14°43'N 43°12'E, 110 m a.s.l. Figure 194. P. abyssinicus from Eritrea, locality 15EG. Figure 195. P. hamar sp. n. paratype from Ethiopia, locality 16EC. Figure 196. P. granimanus from Somaliland, locality 11SE. Figure 197. P. cimrmani holotype." pageId="20" pageNumber="19">197</figureCitation>
|
||
,
|
||
<typeStatus id="5494883EFFC4DC5AFF486BD6F53B37D6" box="[192,285,1240,1263]" pageId="20" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
and allotypic
|
||
<typeStatus id="5494883EFFC4DC5AFE3D6BD6F63337D6" box="[437,533,1240,1263]" pageId="20" pageNumber="19" type="paratype">paratype</typeStatus>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="ED3EAE59FFC4DC5AFDAF6BD6F64B37D6" box="[551,621,1240,1263]" pageId="20" pageNumber="19">FKCP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3356517FFC4DC5AFF486A00F6F235B3" pageId="20" pageNumber="19" type="diagnosis">
|
||
<paragraph id="8B90369CFFC4DC5AFF486A00F6F235B3" blockId="20.[192,802,1294,1674]" pageId="20" pageNumber="19">
|
||
DIAGNOSIS. Adults from
|
||
<quantity id="4CD79B79FFC4DC5AFE6B6A00F610361C" box="[483,566,1294,1317]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="20" pageNumber="19" unit="mm" value="83.0">83 mm</quantity>
|
||
(male) to
|
||
<quantity id="4CD79B79FFC4DC5AFD336A00F707361C" box="[699,801,1294,1317]" metricMagnitude="-2" metricUnit="m" metricValue="8.53" pageId="20" pageNumber="19" unit="mm" value="85.3">85.3 mm</quantity>
|
||
(female) long. Base color uniformly yellow to yellowish brown, only fourth and fifth metasomal segments and telson dark. Pectine teeth number
|
||
<quantity id="4CD79B79FFC4DC5AFDA56A69F6AD3647" box="[557,651,1383,1406]" metricMagnitude="0" metricUnit="m" metricValue="1.5621" metricValueMax="1.5748" metricValueMin="1.5493999999999999" pageId="20" pageNumber="19" unit="in" value="61.5" valueMax="62.0" valueMin="61.0">61–62 in</quantity>
|
||
male and
|
||
<quantity id="4CD79B79FFC4DC5AFD726A69F4DE36A5" metricMagnitude="-1" metricUnit="m" metricValue="8.254999999999999" metricValueMax="8.382" metricValueMin="8.128" pageId="20" pageNumber="19" unit="in" value="32.5" valueMax="33.0" valueMin="32.0">32– 33 in</quantity>
|
||
female. Stridulatory area present on dorsal surface of first and second segments, absent on third segment in female and small, in a disc only, in male. Metasoma densely hirsute. Movable finger of pedipalp more than twice as long as manus, bearing 14 rows of granules which include external and internal granules. Manus of pedipalp smooth and very narrow in both sexes, pedipalp chela length/ width ratio
|
||
<quantity id="4CD79B79FFC4DC5AFE78695BF6623555" box="[496,580,1621,1644]" metricMagnitude="-1" metricUnit="m" metricValue="1.4986" pageId="20" pageNumber="19" unit="in" value="5.9">5.90 in</quantity>
|
||
male and
|
||
<quantity id="4CD79B79FFC4DC5AFD46695BF7043555" box="[718,802,1621,1644]" metricMagnitude="-1" metricUnit="m" metricValue="1.88468" pageId="20" pageNumber="19" unit="in" value="7.42">7.42 in</quantity>
|
||
female. Tarsomere I of all legs with bristlecombs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3356517FFC4DC5AFF4869A4F5E335F8" box="[192,453,1706,1729]" pageId="20" pageNumber="19" type="distribution">
|
||
<paragraph id="8B90369CFFC4DC5AFF4869A4F5E335F8" blockId="20.[192,453,1706,1729]" box="[192,453,1706,1729]" pageId="20" pageNumber="19">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F338760CFFC4DC5AFEEF69A4F5E435F8" box="[359,450,1706,1729]" name="Somalia" pageId="20" pageNumber="19">Somalia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |