treatments-xml/data/87/99/07/879907CFBA195DBC9AADE049FB0FB18E.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.999.52831" ID-GBIF-Dataset="11e8fa4e-ab97-47a5-a8d1-a9748ce5382c" ID-PMC="PMC7723878" ID-Pensoft-Pub="1313-2970-999-147" ID-Pensoft-UUID="69960D48D282536EAC45F3D50DD62BF3" ID-PubMed="33343215" ID-ZooBank="A1116A782C1141FBA8B315AB7369A942" ModsDocID="1313-2970-999-147" checkinTime="1607107216301" checkinUser="pensoft" docAuthor="Kato, Daichi, Watanabe, Kozo &amp; Kolcsar, Levente-Peter" docDate="2020" docId="879907CFBA195DBC9AADE049FB0FB18E" docLanguage="en" docName="ZooKeys 999: 147-163" docOrigin="ZooKeys 999" docSource="http://dx.doi.org/10.3897/zookeys.999.52831" docTitle="Ulomorpha longipenis Kato &amp; Kolcsar 2020, sp. nov." docType="treatment" docUuid="E2EA56ED-B89A-43B8-B668-03E0E4E9FDAC" docUuidSource="ZooBank" docVersion="4" id="69960D48D282536EAC45F3D50DD62BF3" lastPageNumber="147" masterDocId="69960D48D282536EAC45F3D50DD62BF3" masterDocTitle="Synopsis of the genus Ulomorpha Osten Sacken, 1869 (Diptera, Limoniidae) in Japan" masterLastPageNumber="163" masterPageNumber="147" pageNumber="147" updateTime="1668169993919" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Synopsis of the genus Ulomorpha Osten Sacken, 1869 (Diptera, Limoniidae) in Japan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kato, Daichi</mods:namePart>
<mods:affiliation>Echigo-Matsunoyama Museum of Natural Sciences, ' Kyororo', 1712 - 2 Matsunoyama, Tokamachi, 942 - 1411, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Watanabe, Kozo</mods:namePart>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7062-595X</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kolcsar, Levente-Peter</mods:namePart>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-7784-2386</mods:nameIdentifier>
<mods:nameIdentifier type="email">kolcsar.peter@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>999</mods:number>
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<mods:start>147</mods:start>
<mods:end>163</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.999.52831</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.999.52831</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-999-147</mods:identifier>
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<treatment ID-GBIF-Taxon="170678109" LSID="urn:lsid:zoobank.org:act:E2EA56ED-B89A-43B8-B668-03E0E4E9FDAC" httpUri="http://treatment.plazi.org/id/879907CFBA195DBC9AADE049FB0FB18E" lastPageNumber="147" pageId="0" pageNumber="147">
<subSubSection pageId="0" pageNumber="147" type="nomenclature">
<paragraph pageId="0" pageNumber="147">
<taxonomicName LSID="879907CF-BA19-5DBC-9AAD-E049FB0FB18E" authority="Kato &amp; Kolcsár" authorityName="Kato &amp; Kolcsar" authorityYear="2020" class="Insecta" family="Limoniidae" genus="Ulomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulomorpha longipenis" order="Diptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="longipenis" status="sp. nov.">
Ulomorpha longipenis Kato &amp; 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="147">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">Figs 2</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Japanese species of Ulomorpha. A U. amamiana Kato &amp; Kolcsar, sp. nov. (orange), U. nigricolor Alexander, 1924 (blue), U. polytricha Alexander, 1930 (green) B U. longipenis Kato &amp; Kolcsar, sp. nov. (red)." figureDoi="10.3897/zookeys.999.52831.figure5" httpUri="https://binary.pensoft.net/fig/481464" pageId="0" pageNumber="147">, 5B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="materials_examined">
<paragraph pageId="0" pageNumber="147">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" italics="true" pageId="0" pageNumber="147">Holotype</emphasis>
♂, pinned. Original label: &quot;JAPAN, Hokkaido, Sapporo-shi, Minami-ku, Moiwashita, Mt Moiwa-yama; alt. 200 m; 
<geoCoordinate degrees="43" direction="north" minutes="0.84" orientation="latitude" precision="9" value="43.014">43°0.84'N</geoCoordinate>
, 
<geoCoordinate degrees="141" direction="east" minutes="20.01" orientation="longitude" precision="9" value="141.3335">141°20.01'E</geoCoordinate>
; 23 Jun. 2014; D. Kato leg.&quot; &quot;HOLOTYPE 
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2020" class="Insecta" family="Limoniidae" genus="Ulomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ulomorpha longipenis" order="Diptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="longipenis">
<emphasis italics="true" pageId="0" pageNumber="147">Ulomorpha longipenis</emphasis>
</taxonomicName>
Kato &amp; 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. [red label]&quot; (BLKU).
</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" pageId="0" pageNumber="147">
<emphasis italics="true" pageId="0" pageNumber="147">Paratypes</emphasis>
.
</emphasis>
Japan, [Hokkaido] • 3♂; same data as holotype • 1♀; 
<normalizedToken originalValue="Hidaka-chô">Hidaka-cho</normalizedToken>
, Chisaka, tributary of Saru-gawa River; alt. 739 m; 
<geoCoordinate degrees="42" direction="north" minutes="58.39" orientation="latitude" precision="9" value="42.973167">42°58.39'N</geoCoordinate>
, 
<geoCoordinate degrees="142" direction="east" minutes="40.79" orientation="longitude" precision="9" value="142.67984">142°40.79'E</geoCoordinate>
; 28 Jul. 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP). [Honshu] • 1♀; Aomori, Nishimeya-mura, Kawaratai, 
<normalizedToken originalValue="Ôkawa">Okawa</normalizedToken>
Path; alt. 300 m; 
<geoCoordinate degrees="40" direction="north" minutes="30.04" orientation="latitude" precision="9" value="40.500668">40°30.04'N</geoCoordinate>
, 
<geoCoordinate degrees="140" direction="east" minutes="12.24" orientation="longitude" precision="9" value="140.204">140°12.24'E</geoCoordinate>
; 30 Jun. 2013; D. Kato leg. (BLKU) • 3♂; Aomori, Towada-shi, Okuse, Tsutanuma Path; alt. 460 m; 
<geoCoordinate degrees="40" direction="north" minutes="35.45" orientation="latitude" precision="9" value="40.59083">40°35.45'N</geoCoordinate>
, 
<geoCoordinate degrees="140" direction="east" minutes="57.42" orientation="longitude" precision="9" value="140.957">140°57.42'E</geoCoordinate>
; 1 Jun. 2014; D. Kato leg. (BLKU) • 1♂; same data as previous except 21 Jun. 2014 • 1♂; Niigata, 
<normalizedToken originalValue="Tôkamachi-shi">Tokamachi-shi</normalizedToken>
, Matsunoyama-Amamizukoshi, Mt Amamizu-yama; alt. 920 m; 
<geoCoordinate degrees="37" direction="north" minutes="1.46" orientation="latitude" precision="9" value="37.024334">37°1.46'N</geoCoordinate>
, 
<geoCoordinate degrees="138" direction="east" minutes="33.77" orientation="longitude" precision="9" value="138.56284">138°33.77'E</geoCoordinate>
; 2 Aug. 2019; D. Kato leg. (BLKU) • 2♂; Aichi, Seto-shi, 
<normalizedToken originalValue="Iwaya-chô">Iwaya-cho</normalizedToken>
, near 
<normalizedToken originalValue="Iwayadô">Iwayado</normalizedToken>
Park; alt. 300 m; 
<geoCoordinate degrees="35" direction="north" minutes="14.37" orientation="latitude" precision="9" value="35.2395">35°14.37'N</geoCoordinate>
, 
<geoCoordinate degrees="137" direction="east" minutes="9.05" orientation="longitude" precision="9" value="137.15083">137°9.05'E</geoCoordinate>
; 4 May 2016; D. Kato leg. (BLKU) • 1♀; Kyoto, Kibune; 
<geoCoordinate degrees="35" direction="north" minutes="7.29" orientation="latitude" precision="9" value="35.1215">35°7.29'N</geoCoordinate>
, 
<geoCoordinate degrees="135" direction="east" minutes="45.45" orientation="longitude" precision="9" value="135.7575">135°45.45'E</geoCoordinate>
(rough coordinate, altitude unknown);?.IV.1935; M. Tokunaga leg. (USNM). [Shikoku] • 1♂, 1♀; Ehime, 
<normalizedToken originalValue="Kumakôgen-chô">Kumakogen-cho</normalizedToken>
, Chichinokawa; alt. 580 m; 
<geoCoordinate degrees="33" direction="north" minutes="36.29" orientation="latitude" precision="9" value="33.60483">33°36.29'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="51.35" orientation="longitude" precision="9" value="132.85583">132°51.35'E</geoCoordinate>
; 19 May 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 1♂; Ehime, 
<normalizedToken originalValue="Kumakôgen-chô">Kumakogen-cho</normalizedToken>
, Hinoura; alt. 722 m; 
<geoCoordinate degrees="33" direction="north" minutes="35.13" orientation="latitude" precision="9" value="33.5855">33°35.13'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="57.71" orientation="longitude" precision="9" value="132.96184">132°57.71'E</geoCoordinate>
; 19 May 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 2♂; Ehime, 
<normalizedToken originalValue="Kumakôgen-chô">Kumakogen-cho</normalizedToken>
, Nishidani, near Prefectural road 328; alt. 1430 m; 
<geoCoordinate degrees="33" direction="north" minutes="34" orientation="latitude" precision="925" value="33.566666">33°34'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="56.2" orientation="longitude" precision="92" value="132.93666">132°56.2'E</geoCoordinate>
; 17 Jun. 2019; D. Kato leg. (BLKU) • 1♂; same data as previous except alt. 1387 m; 
<geoCoordinate degrees="33" direction="north" minutes="33.89" orientation="latitude" precision="9" value="33.564835">33°33.89'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="56.1" orientation="longitude" precision="92" value="132.935">132°56.1'E</geoCoordinate>
; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 1♀; same data as previous except alt. 890 m; 
<geoCoordinate degrees="33" direction="north" minutes="32.92" orientation="latitude" precision="9" value="33.548668">33°32.92'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="56.88" orientation="longitude" precision="9" value="132.948">132°56.88'E</geoCoordinate>
; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 2♂; Ehime, 
<normalizedToken originalValue="Kumakôgen-chô">Kumakogen-cho</normalizedToken>
, Wakayama; alt. 930 m; 
<geoCoordinate degrees="33" direction="north" minutes="42.95" orientation="latitude" precision="9" value="33.71583">33°42.95'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="6.5" orientation="longitude" precision="92" value="132.10834">132°6.5'E</geoCoordinate>
; 18 May 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 1♂; Ehime, Matsuyama-shi, Jikibamachi; alt. 180 m; 
<geoCoordinate degrees="33" direction="north" minutes="51.69" orientation="latitude" precision="9" value="33.8615">33°51.69'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="49.55" orientation="longitude" precision="9" value="132.82584">132°49.55'E</geoCoordinate>
; 16 May 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 1♀; Ehime, Matsuyama-shi, Shukunomachi; alt. 240 m; 
<geoCoordinate degrees="33" direction="north" minutes="52.08" orientation="latitude" precision="9" value="33.868">33°52.08'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="50.09" orientation="longitude" precision="9" value="132.83484">132°50.09'E</geoCoordinate>
; 3 May 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 1♀; Ehime, Matsuyama-shi, Yuyamayanagi; alt. 250 m; 
<geoCoordinate degrees="33" direction="north" minutes="53.41" orientation="latitude" precision="9" value="33.890167">33°53.41'N</geoCoordinate>
, 
<geoCoordinate degrees="132" direction="east" minutes="50.52" orientation="longitude" precision="9" value="132.842">132°50.52'E</geoCoordinate>
; 3 May 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 1♂, 1♀; Ehime, 
<normalizedToken originalValue="Saijô-shi">Saijo-shi</normalizedToken>
, Nishinokawa-Tei, Mt Ishizuchi-san; alt. 1530 m; 
<geoCoordinate degrees="33" direction="north" minutes="45.29" orientation="latitude" precision="9" value="33.754833">33°45.29'N</geoCoordinate>
, 
<geoCoordinate degrees="133" direction="east" minutes="9.19" orientation="longitude" precision="9" value="133.15317">133°9.19'E</geoCoordinate>
; 16 Jun. 2019; D. Kato leg. (BLKU) • 1♂, 2♀; same data as previous except alt. 1480 m; 
<geoCoordinate degrees="33" direction="north" minutes="45.3" orientation="latitude" precision="92" value="33.755">33°45.3'N</geoCoordinate>
, 
<geoCoordinate degrees="133" direction="east" minutes="9.23" orientation="longitude" precision="9" value="133.15384">133°9.23'E</geoCoordinate>
; 16 May 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP). [Kyushu] • 2♂, 2♀; Fukuoka, Fukuoka-shi, 
<normalizedToken originalValue="Jônan-ku">Jonan-ku</normalizedToken>
, Katae, Mt Abura-yama; alt. 230 m; 
<geoCoordinate degrees="33" direction="north" minutes="31.83" orientation="latitude" precision="9" value="33.5305">33°31.83'N</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="21.96" orientation="longitude" precision="9" value="130.366">130°21.96'E</geoCoordinate>
; 19 Apr. 2014; D. Kato leg. (BLKU) • 1♂, 2♀; Fukuoka, Fukuoka-shi, Sawara-ku, Itaya, Mt Sefuri-san; alt. 970 m; 
<geoCoordinate degrees="33" direction="north" minutes="26.29" orientation="latitude" precision="9" value="33.438168">33°26.29'N</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="22" orientation="longitude" precision="925" value="130.36667">130°22'E</geoCoordinate>
; 4 Jun. 2015; D. Kato leg. (BLKU) • 2♀; same data as previous except 10 Jun. 2015 • 2♀; same data as previous except 17 Jun. 2015 • 1♂; Fukuoka, Miyako-machi, Saigawa-Hobashira, 
<normalizedToken originalValue="Notȏge">Notȏge</normalizedToken>
Pass; alt. 740 m; 
<geoCoordinate degrees="33" direction="north" minutes="29.74" orientation="latitude" precision="9" value="33.495667">33°29.74'N</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="57.69" orientation="longitude" precision="9" value="130.9615">130°57.69'E</geoCoordinate>
; 12 May 2019; D. Kato leg. (BLKU) • 2♀; Fukuoka, Miyawaka-shi, Inunaki, Mt Inunaki-san; alt. 300 m; 
<geoCoordinate degrees="33" direction="north" minutes="40.87" orientation="latitude" precision="9" value="33.681168">33°40.87'N</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="33.19" orientation="longitude" precision="9" value="130.55316">130°33.19'E</geoCoordinate>
; 5 May 2015; D. Kato leg. (BLKU) • 1♀; Saga, Kanzaki-shi, Sefuri-machi-Fukumaki; alt. 980 m; 
<geoCoordinate degrees="33" direction="north" minutes="26.04" orientation="latitude" precision="9" value="33.434">33°26.04'N</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="22.12" orientation="longitude" precision="9" value="130.36867">130°22.12'E</geoCoordinate>
; 23 May 2019; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg. (CKLP) • 1♂; Saga, Saga-shi, Fuji-machi-Sekiya, Kase-gawa River near Hokuzan Dam; alt. 320 m; 
<geoCoordinate degrees="33" direction="north" minutes="25.99" orientation="latitude" precision="9" value="33.433167">33°25.99'N</geoCoordinate>
, 
<geoCoordinate degrees="130" direction="east" minutes="13.93" orientation="longitude" precision="9" value="130.23216">130°13.93'E</geoCoordinate>
; 23 May 2015; D. Kato leg. (BLKU) • 2♂; Miyazaki, 
<normalizedToken originalValue="Takachiho-chô">Takachiho-cho</normalizedToken>
, Gokasho, Mt Sobo-san, near Kitadani trailhead; alt. 1150 m; 
<geoCoordinate degrees="32" direction="north" minutes="49.36" orientation="latitude" precision="9" value="32.822666">32°49.36'N</geoCoordinate>
, 
<geoCoordinate degrees="131" direction="east" minutes="19.64" orientation="longitude" precision="9" value="131.32733">131°19.64'E</geoCoordinate>
; 22 May 2019; D. Kato leg. (BLKU).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="diagnosis">
<paragraph pageId="0" pageNumber="147">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="147">Body blackish. Vertex sparsely pruinose, sometimes partly polished. Flagellomeres with basal 2-3 segments oval ventral sides with pubescences. Scutum sparsely pruinose or polished. Wing brownish tinged, with oval, dark-brown stigma and weakly dark seam on anterior part of cord. Halter yellowish. Interbase with outer lobe shorter than medial lobe in dorsal view; medial lobe medial to base of outer lobe about 2.3 times as long as wide. Aedeagus with rod-shaped distal part, 3.3-4.4 times as long as wide and almost straight.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="description">
<paragraph pageId="0" pageNumber="147">Description.</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" pageId="0" pageNumber="147">Male</emphasis>
. Body length 7.2-11.0 mm, wing length 7.1-11.9 mm.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" pageId="0" pageNumber="147">
<emphasis italics="true" pageId="0" pageNumber="147">Head</emphasis>
:
</emphasis>
subnitidous black, sparsely dusted with gray pruinosity; vertex with brighter gray pruinosity at anterior end; anterior part of vertex sometimes largely shiny except anterior end or with shiny and small, longitudinally long bacilliform area at middle posterior to anterior brighter area. Rostrum and mouthparts dark brown. Antenna with scape and pedicel dark brown; flagellum dusky yellow to brown; basal 2-3 flagellomeres oval, covered with pubescences ventrally.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" pageId="0" pageNumber="147">
<emphasis italics="true" pageId="0" pageNumber="147">Thorax</emphasis>
:
</emphasis>
subnitidous black, sparsely dusted with brownish pruinosity, sometimes polished and pruinosity absent on prescutum and most of scutal lobe; postpronotum yellowish or brownish. Wing (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2B</figureCitation>
) tinged with brown; basal and costal regions proximal to cord yellowish; stigma oval, dark brown, with faint outline; veins dark brown, yellowish on Sc and basal part of wing; weakly dark seam on fork of Rs to crossvein r-m. Halter yellow to dusky yellow. Legs mainly yellow to dusky yellow; fore coxa dark on basal half, sometimes entirely so, mid and hind coxae sometimes narrowly dark at bases; femora narrowly dark at tips; dark area on fore femur often extending to near middle; tibiae narrowly dark at tips; tarsi weakly dark from tip of tarsomere 1 to apical segment.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" pageId="0" pageNumber="147">
<emphasis italics="true" pageId="0" pageNumber="147">Abdomen</emphasis>
:
</emphasis>
subnitidous dark brown to black, sparsely covered with brownish pruinosity.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" italics="true" pageId="0" pageNumber="147">Male terminalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2C-G</figureCitation>
): caudal margin of tergite 9 roundly produced at middle, with shallow U-shaped notch at center. Outer gonostylus in dorsal view (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2D</figureCitation>
) with tip narrowed and curved anteriorly, concaved margin with distinct teeth. Interbase with outer lobe shorter than medial lobe in dorsal view (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2E</figureCitation>
); outer lobe wide at base, inner basal end situated near middle of medial lobe (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2G</figureCitation>
); medial lobe strongly narrowed distally, weakly sinuous, distal part medial to base of outer lobe about 2.3 times as long as wide (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2G</figureCitation>
). Aedeagus with rod-shaped distal part, 3.3-4.4 times as long as wide and almost straight (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2E, F</figureCitation>
), usually shorter in specimens from southern part of Japan.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" pageId="0" pageNumber="147">Female</emphasis>
. Body length 7.0-11.7 mm, wing length 6.2-10.2 mm. Generally resembling male.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" pageId="0" pageNumber="147">
<emphasis italics="true" pageId="0" pageNumber="147">Ovipositor</emphasis>
:
</emphasis>
dark brown; yellow on cercus, hypogynial valve, and distal 1/4-1/3 of tergite 10; cercus 1.5 times as long as tergite 10.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="etymology">
<paragraph pageId="0" pageNumber="147">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="147">
The specific epithet is from the Latin 
<emphasis italics="true" pageId="0" pageNumber="147">longus/longi</emphasis>
(long) + 
<emphasis italics="true" pageId="0" pageNumber="147">penis</emphasis>
(penis) and refers to the long aedeagus of this species compared to the other Japanese species of the genus. The name is an adjective in nominative singular.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="distribution">
<paragraph pageId="0" pageNumber="147">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="147">
Japan (Hokkaido, Honshu, Shikoku, and Kyushu islands) (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Japanese species of Ulomorpha. A U. amamiana Kato &amp; Kolcsar, sp. nov. (orange), U. nigricolor Alexander, 1924 (blue), U. polytricha Alexander, 1930 (green) B U. longipenis Kato &amp; Kolcsar, sp. nov. (red)." figureDoi="10.3897/zookeys.999.52831.figure5" httpUri="https://binary.pensoft.net/fig/481464" pageId="0" pageNumber="147">5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="remarks">
<paragraph pageId="0" pageNumber="147">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="147">
This species is similar to 
<taxonomicName lsidName="U. polytricha" pageId="0" pageNumber="147" rank="species" species="polytricha">
<emphasis italics="true" pageId="0" pageNumber="147">U. polytricha</emphasis>
</taxonomicName>
Alexander, 1930 but is differentiated from it by the following characters: wing with stigma distinctly dark, fork of Rs to crossvein r-m weakly darkened (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2B</figureCitation>
) (stigma indistinct in 
<taxonomicName lsidName="U. polytricha" pageId="0" pageNumber="147" rank="species" species="polytricha">
<emphasis italics="true" pageId="0" pageNumber="147">U. polytricha</emphasis>
</taxonomicName>
; Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ulomorpha polytricha Alexander, 1930. A habitus, male B basal part of antenna, lateral view C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view. Scale bars: 3 mm (A, C); 0.1 mm (B, D-H)." figureDoi="10.3897/zookeys.999.52831.figure4" httpUri="https://binary.pensoft.net/fig/481463" pageId="0" pageNumber="147">4C</figureCitation>
); aedeagus with rod-shaped part more than three times as long as wide (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ulomorpha longipenis Kato &amp; Kolcsar, sp. nov. A habitus, male B wing C male terminalia, dorsal view D tip of outer gonostylus, dorsal view E aedeagal complex, dorsal view F aedeagal complex, lateral view G interbase, posterodorsal view. Scale bars: 3 mm (A, B); 0.1 mm (C-G)." figureDoi="10.3897/zookeys.999.52831.figure2" httpUri="https://binary.pensoft.net/fig/481461" pageId="0" pageNumber="147">2F</figureCitation>
) (twice as long as wide in in 
<taxonomicName lsidName="U. polytricha" pageId="0" pageNumber="147" rank="species" species="polytricha">
<emphasis italics="true" pageId="0" pageNumber="147">U. polytricha</emphasis>
</taxonomicName>
; Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ulomorpha polytricha Alexander, 1930. A habitus, male B basal part of antenna, lateral view C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view. Scale bars: 3 mm (A, C); 0.1 mm (B, D-H)." figureDoi="10.3897/zookeys.999.52831.figure4" httpUri="https://binary.pensoft.net/fig/481463" pageId="0" pageNumber="147">4G</figureCitation>
). This species also resembles a Nearctic species, 
<taxonomicName lsidName="U. nigronitida" pageId="0" pageNumber="147" rank="species" species="nigronitida">
<emphasis italics="true" pageId="0" pageNumber="147">U. nigronitida</emphasis>
</taxonomicName>
Alexander, 1920, according to the original description (
<bibRefCitation author="Alexander, CP" journalOrPublisher="Memoirs, Cornell University Agricultural Experiment Station" pageId="0" pageNumber="147" refId="B3" refString="Alexander, CP, 1920b. Undescribed Tipulidae (Diptera) from western North America. Proceedings of the California Academy of Sciences (Series 4) 10(5): 35-46." title="Undescribed Tipulidae (Diptera) from western North America. Proceedings of the California Academy of Sciences (Series 4) 10 (5): 35 - 46." year="1920 b">Alexander 1920b</bibRefCitation>
), but is distinguished from it by the following characters: antenna pale on flagellum, yellow to brown (black throughout in 
<taxonomicName lsidName="U. nigronitida" pageId="0" pageNumber="147" rank="species" species="nigronitida">
<emphasis italics="true" pageId="0" pageNumber="147">U. nigronitida</emphasis>
</taxonomicName>
); coxae with distal parts yellowish (coxae brownish black, hind pair paler in 
<taxonomicName lsidName="U. nigronitida" pageId="0" pageNumber="147" rank="species" species="nigronitida">
<emphasis italics="true" pageId="0" pageNumber="147">U. nigronitida</emphasis>
</taxonomicName>
); halter entirely yellowish (knob dark brownish black in 
<taxonomicName lsidName="U. nigronitida" pageId="0" pageNumber="147" rank="species" species="nigronitida">
<emphasis italics="true" pageId="0" pageNumber="147">U. nigronitida</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>