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<document id="9DE612A55101E96BD5E00C883E1182B2" ID="10.11646/zootaxa.4169.3.7" ID-DOI="10.11646/zootaxa.4169.3.7" ID-GBIF-Dataset="faeafa7a-a105-4443-bbc3-2bea598d8cd5" ID-ISSN="1175-5326" ID-Zenodo-Dep="258027" ID-ZooBank="8A1FB454-AC73-4D54-A5CB-D12D6CD82AC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474440730591" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2016" docId="8706BE21FFB2FF911DC8F9BFFDBBA03E" docLanguage="en" docName="zootaxa.4169.3.7.pdf" docOrigin="Zootaxa 4169 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexachaetus brunki Tian &amp; Deuve, 2016, n. sp." docType="treatment" docVersion="7" lastPageNumber="549" masterDocId="7B3FC659FFBAFF981D5FFF88FFF0A546" masterDocTitle="Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini)" masterLastPageNumber="554" masterPageNumber="540" pageNumber="548" updateTime="1698386596725" updateUser="plazi">
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<mods:title id="D0F4BD503B50BAD1EDC42C57E8804258">Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini)</mods:title>
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<mods:namePart id="EA1C3447B23952D2DBF4573142ADC5F9">Tian, Mingyi</mods:namePart>
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<mods:namePart id="3621C900A9EB0E0CF5BD1BB66E83D755">Deuve, Thierry</mods:namePart>
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<mods:title id="718BC6F1F1F510EDE99E50A0A3FC1777">Zootaxa</mods:title>
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<mods:date id="C472F9ABBDF2482B1F520133B6C161BE">2016</mods:date>
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<treatment id="8706BE21FFB2FF911DC8F9BFFDBBA03E" ID-DOI="http://doi.org/10.5281/zenodo.6075827" ID-GBIF-Taxon="124552519" ID-Zenodo-Dep="6075827" LSID="urn:lsid:plazi:treatment:8706BE21FFB2FF911DC8F9BFFDBBA03E" httpUri="http://treatment.plazi.org/id/8706BE21FFB2FF911DC8F9BFFDBBA03E" lastPageId="9" lastPageNumber="549" pageId="8" pageNumber="548">
<subSubSection id="47B55CBCFFB2FF901DC8F9BFFE96A335" pageId="8" pageNumber="548" type="nomenclature">
<paragraph id="0F100F37FFB2FF901DC8F9BFFE2FA314" blockId="8.[151,479,1591,1651]" box="[151,479,1591,1618]" pageId="8" pageNumber="548">
<heading id="5458B85BFFB2FF901DC8F9BFFE2FA314" bold="true" box="[151,479,1591,1618]" fontSize="11" level="1" pageId="8" pageNumber="548" reason="1">
<emphasis id="3DDBD325FFB2FF901DC8F9BFFE2FA314" bold="true" box="[151,479,1591,1618]" pageId="8" pageNumber="548">
<taxonomicName id="C8AF74B4FFB2FF901DC8F9BFFE7DA317" ID-CoL="3LG2C" box="[151,397,1591,1617]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="548" phylum="Arthropoda" rank="species" species="brunki" status="sp. nov.">
<emphasis id="3DDBD325FFB2FF901DC8F9BFFE7DA317" bold="true" box="[151,397,1591,1617]" italics="true" pageId="8" pageNumber="548">Hexachaetus brunki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="26E86E5EFFB2FF901CC3F9B0FE2FA314" box="[412,479,1592,1618]" pageId="8" pageNumber="548" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0F100F37FFB2FF901DC8F9D2FE96A335" blockId="8.[151,479,1591,1651]" box="[151,358,1626,1651]" pageId="8" pageNumber="548">
(
<figureCitation id="979413B2FFB2FF901DC1F9D2FEFBA334" box="[158,267,1626,1651]" captionStart="FIGURES 5 8" captionStartId="5.[151,264,1945,1968]" captionTargetBox="[424,1148,585,1915]" captionTargetId="figure@5.[417,1162,575,1919]" captionTargetPageId="5" captionText="FIGURES 5 8. Habitus of Hexachaetus species (5. H. maculatus Tian &amp; Deuve, male, the holotype; 6. H. kirschenhoferi n. sp., male, holotype (photo: E. Kirschhofer); 7. H. brunki n. sp., male holotype; 8. H. laevissimus Chaudoir, 1878, female. Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258029/files/figure.png" pageId="8" pageNumber="548">Figures 7</figureCitation>
,
<figureCitation id="979413B2FFB2FF901C48F9D2FEAEA335" box="[279,350,1626,1651]" captionStart="FIGURES 12 19" captionStartId="7.[151,264,1737,1760]" captionTargetBox="[277,1307,185,1721]" captionTargetId="figure@7.[259,1328,173,1732]" captionTargetPageId="7" captionText="FIGURES 12 19. Male genitalia of Hexachaetus species, right lateral view and dorsal view for apical lamella (12 13. H. maculatus Tian &amp; Deuve; 14 15. H. kirschenhoferi n. sp.; 16 17. H. brunki n. sp.; 18 19. H. vietnamensis n. sp.)" httpUri="https://zenodo.org/record/258031/files/figure.png" pageId="8" pageNumber="548">1617</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB2FF901DC8F92AFA68A3FD" box="[151,1432,1698,1723]" pageId="8" pageNumber="548" type="materials_examined">
<paragraph id="0F100F37FFB2FF901DC8F92AFA68A3FD" blockId="8.[151,1436,1698,2011]" box="[151,1432,1698,1723]" pageId="8" pageNumber="548">
<materialsCitation id="BFC7056AFFB2FF901DC8F92AFA68A3FD" ID-GBIF-Occurrence="1313880541" box="[151,1432,1698,1723]" collectingDate="2011-05" collectionCode="CIB" collectorName="K. P. Yeb" country="Malaysia" elevation="1500" location="Boneo-North" pageId="8" pageNumber="548" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="3DDBD325FFB2FF901DC8F92AFEF6A3FD" bold="true" box="[151,262,1698,1723]" pageId="8" pageNumber="548">Material.</emphasis>
<typeStatus id="D014B195FFB2FF901C53F92AFE84A3FD" box="[268,372,1698,1723]" pageId="8" pageNumber="548" type="holotype">Holotype</typeStatus>
male, “
<location id="0A7059ECFFB2FF901C95F92AFD93A3FD" LSID="urn:lsid:plazi:treatment:8706BE21FFB2FF911DC8F9BFFDBBA03E:0A7059ECFFB2FF901C95F92AFD93A3FD" box="[458,611,1698,1723]" country="Malaysia" name="Boneo-North" pageId="8" pageNumber="548" stateProvince="Sabah">Boneo-North</location>
,
<collectingRegion id="CD6BC1D5FFB2FF901F30F92AFD43A3FD" box="[623,691,1698,1723]" country="Malaysia" name="Sabah" pageId="8" pageNumber="548">Sabah</collectingRegion>
Trus Madi,
<elevation id="8482E804FFB2FF901E63F92AFC63A3FC" box="[828,915,1698,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="548" unit="m" value="1500.0">
<quantity id="C857A2D2FFB2FF901E63F92AFC63A3FC" box="[828,915,1698,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="548" unit="m" value="1500.0">1500 m</quantity>
</elevation>
,
<collectingDate id="6B55D01FFFB2FF901EC0F92BFBE3A3FD" box="[927,1043,1699,1723]" pageId="8" pageNumber="548" value="2011-05">
<date id="7B1129F7FFB2FF901EC0F92BFBE3A3FD" box="[927,1043,1699,1723]" pageId="8" pageNumber="548" value="2011-05">May 2011</date>
</collectingDate>
, leg.
<collectorName id="A25A6AE1FFB2FF901910F92BFB47A3FD" box="[1103,1207,1698,1723]" pageId="8" pageNumber="548">K. P. Yeb</collectorName>
(
<collectingCountry id="77B84FA7FFB2FF901999F92AFAC2A3FD" box="[1222,1330,1698,1723]" name="Malaysia" pageId="8" pageNumber="548">Malaysia</collectingCountry>
), in
<collectionCode id="69BE97F2FFB2FF90183CF92BFA65A3FD" box="[1379,1429,1699,1723]" country="Mexico" httpUri="http://grbio.org/cool/vxkr-andy" name="Centro de Investigaciones Biologicas del Noroeste, S.C. (Mexico)" pageId="8" pageNumber="548">CIB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB2FF901D98F94EFCDBA2D4" pageId="8" pageNumber="548" type="diagnosis">
<paragraph id="0F100F37FFB2FF901D98F94EFB10A228" blockId="8.[151,1436,1698,2011]" pageId="8" pageNumber="548">
<emphasis id="3DDBD325FFB2FF901D98F94EFEB1A399" bold="true" box="[199,321,1734,1759]" pageId="8" pageNumber="548">Diagnosis.</emphasis>
Medium-sized for
<taxonomicName id="C8AF74B4FFB2FF901F6DF94EFD7FA398" box="[562,655,1734,1758]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="548" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="3DDBD325FFB2FF901F6DF94EFD7FA398" box="[562,655,1734,1758]" italics="true" pageId="8" pageNumber="548">lateralis</emphasis>
</taxonomicName>
species group, body moderately elongate-ovate, smooth, polish, impunctate and yellowish; head, longitudinal median portion of pronotum, and central marking of elytra black; elytral marking anchoreaform, wider; ligula 6-setose, clypeus bisetose, middle tibia slightly dilated in male; pronotum strongly convex, lateral expanded margin indistinctly reflexed, hind angle round off, elytra strongly and obliquely truncate at apex, inner apical angle acute and sharp, striae rather deep, intervals convex.
</paragraph>
<paragraph id="0F100F37FFB2FF901D98F8F2FCDBA2D4" blockId="8.[151,1436,1698,2011]" box="[199,811,1914,1939]" pageId="8" pageNumber="548">
Length:
<quantity id="C857A2D2FFB2FF901C7AF8F2FE7CA2D4" box="[293,396,1914,1939]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="8" pageNumber="548" unit="mm" value="15.5">15.5 mm</quantity>
; width: 6.0 mm. Habitus as in
<figureCitation id="979413B2FFB2FF901FBCF8F2FCD7A2D4" box="[739,807,1914,1939]" captionStart="FIGURES 5 8" captionStartId="5.[151,264,1945,1968]" captionTargetBox="[424,1148,585,1915]" captionTargetId="figure@5.[417,1162,575,1919]" captionTargetPageId="5" captionText="FIGURES 5 8. Habitus of Hexachaetus species (5. H. maculatus Tian &amp; Deuve, male, the holotype; 6. H. kirschenhoferi n. sp., male, holotype (photo: E. Kirschhofer); 7. H. brunki n. sp., male holotype; 8. H. laevissimus Chaudoir, 1878, female. Scale bar: 5.0 mm)" httpUri="https://zenodo.org/record/258029/files/figure.png" pageId="8" pageNumber="548">Fig. 7</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB2FF911D98F816FE6EA183" lastPageId="9" lastPageNumber="549" pageId="8" pageNumber="548" type="description">
<paragraph id="0F100F37FFB2FF911D98F816FB3AA5BE" blockId="8.[151,1436,1698,2011]" lastBlockId="9.[151,1437,151,1400]" lastPageId="9" lastPageNumber="549" pageId="8" pageNumber="548">
<emphasis id="3DDBD325FFB2FF901D98F816FEA9A2F1" bold="true" box="[199,345,1950,1975]" pageId="8" pageNumber="548">Description.</emphasis>
Yellow, but head (except labrum and median areas of clypeus and frons), median portion of pronotum and elytra median, and tibiae dark brown; palps, antennae and tarsi brown; elytra marking anchoreaform, occupied intervals
<quantity id="C857A2D2FFB3FF911C2BFF10FE2EA5F6" box="[372,478,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="9" pageNumber="549" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1 to 4 in</quantity>
the widest points, ending at about apical 2/11 portion of elytra, margins without marking throughout; head and pronotum smooth, glabrous, elytra with tiny and irregular punctures on intervals; microsculptural engraved meshes vanished on head, striate on pronotum, isodiametric on elytra.
</paragraph>
<paragraph id="0F100F37FFB3FF911D98FE8CFB36A77A" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="549">Head stout, as long as wide, clypeus somewhat uneven at base, bisetose; eyes large and very prominent, frontal impressions well-marked, foveate, short, finishing the level of fore supraorbital punctures; frons and vertex moderately convex; mandibles stout, apex moderately hooked; mentum and submentum each with a pair of setae, palpiger unsetose, ligula slightly enlarged, 6-setose at apex, central one pair of setae longest, middle pair longer than outer pair; palps subcylindrical, short and stout, maxillary palpomeres 3 and 4 subequal; labial palpomere 2 much longer (1.5 times) than 3, bisetose on inner margin; labrum nearly quadrate, front margin straight, 6-setose, surface smooth. Antennae moderately long, extending over base of elytra, pubescent from antennomere 4, concolorous; antennomere 13 glabrous, antennomere 1 with a long seta at subapex, and several shorter setae at apexes of 13; antennomere 1 about twice as long as antennomere 2, and slightly longer than 3;</paragraph>
<paragraph id="0F100F37FFB3FF911D98FDC0FE02A7B6" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="549">Pronotum transverse, PW/PL=1.70, widest at about middle; disc markedly convex, smooth and glabrous; both front and basal margin bordered, transversal impressions well-marked, median line distinct; lateral expanded margins narrow at front, then gradually widened towards base, quite flat throughout; front margin straight at middle, basal margin faintly bisinuate, median portion hardly produced backwards; hind and front angles round off; basal foveae shallow and faint.</paragraph>
<paragraph id="0F100F37FFB3FF911D98FD74FC0CA6C6" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="549">Elytra elongate-ovate, convex; EL/EW=1.70; near parallel-sided, widest at middle; base well bordered, apex deeply and obliquely truncate, outer angles pointed, inner angles very sharp; striae rather deep, intervals convex, odd and even intervals subequal at middle; interval 3 with basal and subapical setiferous pores, basal one wellmarked, closed to stria 3, subapical one small, at about middle of the interval.</paragraph>
<paragraph id="0F100F37FFB3FF911D98FC05FEF4A156" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="549">Fore tibia slightly dilated at apex, apex slightly arcuate, outer margin sub-serrate; middle tibia not evidently dilated at middle in male; hind legs slender, apex only slightly enlarged, apical spurs long and sharp, tarsomere 1.5 times as long as tarsomere 2; tarsomere 3 as long as 4 which is deeply bilobed at apex; all tarsal claws strongly pectinate.</paragraph>
<paragraph id="0F100F37FFB3FF911D98FB94FA67A173" blockId="9.[151,1437,151,1400]" box="[199,1431,1052,1077]" pageId="9" pageNumber="549">Prosternal process unbordered at apex; abdominal ventrite VII of male small but deeply emarginate at middle.</paragraph>
<paragraph id="0F100F37FFB3FF911D98FBB7FD1DA1E6" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="549">
Male genitalia (
<figureCitation id="979413B2FFB3FF911C29FBB7FDF0A11E" box="[374,512,1087,1112]" captionStart="FIGURES 12 19" captionStartId="7.[151,264,1737,1760]" captionTargetBox="[277,1307,185,1721]" captionTargetId="figure@7.[259,1328,173,1732]" captionTargetPageId="7" captionText="FIGURES 12 19. Male genitalia of Hexachaetus species, right lateral view and dorsal view for apical lamella (12 13. H. maculatus Tian &amp; Deuve; 14 15. H. kirschenhoferi n. sp.; 16 17. H. brunki n. sp.; 18 19. H. vietnamensis n. sp.)" httpUri="https://zenodo.org/record/258031/files/figure.png" pageId="9" pageNumber="549">Figs. 1617</figureCitation>
): The median lobe of aedeagus short and stout, slightly sinuate ventrally, expanded medially, apex sharp in lateral view, dorsal open quite large; in dorsal view, the apical lamella is rather long though distinctly shorter than wide, broadly rounded at apex.
</paragraph>
<paragraph id="0F100F37FFB3FF911D98FB24FE6EA183" blockId="9.[151,1437,151,1400]" box="[199,414,1196,1221]" pageId="9" pageNumber="549">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB3FF911D98FB47FC96A076" pageId="9" pageNumber="549" type="discussion">
<paragraph id="0F100F37FFB3FF911D98FB47FC96A076" blockId="9.[151,1437,151,1400]" pageId="9" pageNumber="549">
<emphasis id="3DDBD325FFB3FF911D98FB47FECBA1AE" bold="true" box="[199,315,1231,1256]" pageId="9" pageNumber="549">Remarks.</emphasis>
This species is allied to
<taxonomicName id="C8AF74B4FFB3FF911F09FB58FCECA1AE" box="[598,796,1232,1256]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="549" phylum="Arthropoda" rank="species" species="kirschenhoferi" status="sp. nov.">
<emphasis id="3DDBD325FFB3FF911F09FB58FCECA1AE" box="[598,796,1232,1256]" italics="true" pageId="9" pageNumber="549">O. kirschenhoferi</emphasis>
</taxonomicName>
<emphasis id="3DDBD325FFB3FF911E7BFB47FC94A1AE" bold="true" box="[804,868,1231,1256]" pageId="9" pageNumber="549">
<taxonomicNameLabel id="26E86E5EFFB3FF911E7BFB47FC94A1AE" box="[804,868,1231,1256]" pageId="9" pageNumber="549" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but differs from the latter species by: large-sized body, more transverse pronotum in front, elytra marking not extended to margin areas, and aedeagus sharp at apex (versus blunt in
<taxonomicName id="C8AF74B4FFB3FF911C13FA90FDE0A076" box="[332,528,1304,1328]" class="Insecta" family="Carabidae" genus="Hexachaetus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="549" phylum="Arthropoda" rank="species" species="kirschenhoferi">
<emphasis id="3DDBD325FFB3FF911C13FA90FDE0A076" box="[332,528,1304,1328]" italics="true" pageId="9" pageNumber="549">O. kirschenhoferi</emphasis>
</taxonomicName>
), with a longer apical lamella.
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB3FF911D98FAB4FC76A012" box="[199,902,1340,1365]" pageId="9" pageNumber="549" type="etymology">
<paragraph id="0F100F37FFB3FF911D98FAB4FC76A012" blockId="9.[151,1437,151,1400]" box="[199,902,1340,1365]" pageId="9" pageNumber="549">
<emphasis id="3DDBD325FFB3FF911D98FAB4FEBEA013" bold="true" box="[199,334,1340,1365]" pageId="9" pageNumber="549">Etymology.</emphasis>
In honour of Dr. Ingo Brunk (Dresden,
<collectingCountry id="77B84FA7FFB3FF911E52FAB4FC8BA012" box="[781,891,1340,1364]" name="Germany" pageId="9" pageNumber="549">Germany</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="47B55CBCFFB3FF911D98FAD7FDBBA03E" box="[199,587,1375,1400]" pageId="9" pageNumber="549" type="distribution">
<paragraph id="0F100F37FFB3FF911D98FAD7FDBBA03E" blockId="9.[151,1437,151,1400]" box="[199,587,1375,1400]" pageId="9" pageNumber="549">
<emphasis id="3DDBD325FFB3FF911D98FAD7FE90A03E" bold="true" box="[199,352,1375,1400]" pageId="9" pageNumber="549">Distribution.</emphasis>
<collectingCountry id="77B84FA7FFB3FF911C38FAD7FE3EA03E" box="[359,462,1375,1400]" name="Malaysia" pageId="9" pageNumber="549">Malaysia</collectingCountry>
(
<figureCitation id="979413B2FFB3FF911C82FAD7FDC1A03E" box="[477,561,1375,1400]" captionStart="FIGURE 20" captionStartId="12.[151,250,1789,1811]" captionTargetBox="[226,1360,198,1765]" captionTargetId="figure@12.[223,1363,195,1768]" captionTargetPageId="12" captionText="FIGURE 20. Distribution of Hexachaetus species (square: angulatus species group; round: lateralis species group; star: laevissimus species group; heart: vietnamensis species group; triangle: mulan species group; a. H. lateralis lateralis; b. H. lateralis maindroni; c. H. maculatus; d. H. kirschenhoferi n. sp.; e. H. brunki n. sp.; f. H. laevissimus; g. H. vietnamensis n. sp.; h. H. prodigus; i. H. mulan n. sp.)" httpUri="https://zenodo.org/record/258032/files/figure.png" pageId="9" pageNumber="549">Fig. 20</figureCitation>
e).
</paragraph>
</subSubSection>
</treatment>
</document>