211 lines
15 KiB
XML
211 lines
15 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.4.4.13" ID-GBIF-Dataset="6cd3d259-465d-4ab3-8a32-d57a812bb6e0" ID-ISSN="2624-2834" ID-Zenodo-Dep="5507925" ID-ZooBank="D1F5A853-32D1-470B-A609-D54F7B8C60CA" checkinTime="1631645857228" checkinUser="carolina" docAuthor="JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S. & CAI, CHEN-YANG" docDate="2021" docId="03B28787AB4E0671FF0D87B2FE97F302" docLanguage="en" docName="palaeoentomology.4.4.13.pdf" docOrigin="Palaeoentomology 4 (4)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.4.13" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Tanytermitalis philetaerus JIANG & ZHANG & ELDREDGE & SONG & LI & TIHELKA & HUANG & WANG & ENGEL & CAI 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="386" masterDocId="FF8BFFFFAB42067DFF858453FF89FB0A" masterDocTitle="Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera)" masterLastPageNumber="389" masterPageNumber="374" pageNumber="386" updateTime="1661815476951" updateUser="admin">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>JIANG, RI-XIN</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>ZHANG, HONG-RUI</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>ELDREDGE, K. TARO</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>SONG, XIAO-BIN</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>LI, YAN-DA</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>TIHELKA, ERIK</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>HUANG, DI-YING</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>WANG, SHUO</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>ENGEL, MICHAEL S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>CAI, CHEN-YANG</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-08-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>374</mods:start>
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<mods:end>389</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/palaeoentomology.4.4.13</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.4.4.13</mods:identifier>
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<mods:identifier type="GBIF-Dataset">6cd3d259-465d-4ab3-8a32-d57a812bb6e0</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5507925</mods:identifier>
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<mods:identifier type="ZooBank">D1F5A853-32D1-470B-A609-D54F7B8C60CA</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5515618" ID-GBIF-Taxon="188191030" ID-Zenodo-Dep="5515618" LSID="urn:lsid:plazi:treatment:03B28787AB4E0671FF0D87B2FE97F302" httpUri="http://treatment.plazi.org/id/03B28787AB4E0671FF0D87B2FE97F302" lastPageNumber="386" pageId="12" pageNumber="386">
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<subSubSection box="[136,696,993,1020]" pageId="12" pageNumber="386" type="nomenclature">
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<paragraph blockId="12.[136,696,993,1055]" box="[136,696,993,1020]" pageId="12" pageNumber="386">
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<heading bold="true" box="[136,696,993,1020]" fontSize="10" level="2" pageId="12" pageNumber="386" reason="3">
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<emphasis bold="true" box="[136,696,993,1020]" pageId="12" pageNumber="386">
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<taxonomicName authority="Engel & Cai" authorityName="JIANG & ZHANG & ELDREDGE & SONG & LI & TIHELKA & HUANG & WANG & ENGEL & CAI" authorityYear="2021" box="[136,593,993,1020]" class="Insecta" family="Kalotermitidae" genus="Tanytermitalis" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="386" phylum="Arthropoda" rank="species" species="philetaerus" status="sp. nov.">
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<emphasis bold="true" box="[136,438,993,1020]" italics="true" pageId="12" pageNumber="386">Tanytermitalis philetaerus</emphasis>
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Engel & Cai
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</taxonomicName>
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,
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<taxonomicNameLabel box="[606,696,993,1019]" pageId="12" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[136,305,1029,1055]" pageId="12" pageNumber="386" type="description">
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<paragraph blockId="12.[136,696,993,1055]" box="[136,305,1029,1055]" pageId="12" pageNumber="386">
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(
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<figureCitation box="[144,270,1029,1055]" captionStart="FIGURE 1" captionStartId="2.[136,235,1689,1713]" captionTargetBox="[183,1404,179,1665]" captionTargetId="figure-90@2.[183,1404,179,1665]" captionTargetPageId="2" captionText="FIGURE 1. Mid-Cretaceous termites with their putative rove beetle termitophiles. A, Arceotermes hospitis gen. et sp. nov. and Cretotrichopsenius burmiticus Cai et al., 2017a (QUST-INSECT-0015). B, Tanytermitalis philetaerus gen. et sp. nov. and C. burmiticus (NIGP174707). C, The entire amber piece of QUST-INSECT-0015. D, The entire amber piece of NIGP174707. Scale bars = 1 mm in A & C; 2 mm in B; 10 mm in D." figureDoi="http://doi.org/10.5281/zenodo.5507927" httpUri="https://zenodo.org/record/5507927/files/figure.png" pageId="12" pageNumber="386">Figs 1B, D</figureCitation>
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,
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<figureCitation box="[282,297,1029,1055]" captionStart="FIGURE 7" captionStartId="11.[136,235,1309,1333]" captionTargetBox="[151,1436,189,1286]" captionTargetId="figure-295@11.[151,1436,189,1286]" captionTargetPageId="11" captionText="FIGURE 7. Holotype alate of Tanytermitalis philetaerus Engel & Cai, gen. et sp. nov., alate (NIGP174707). A, Dorsal habitus. B, Detail of portion of right antenna as preserved. C, Dorsal view of head. D, Ventral view of head, green fluorescence light source. E, Right foreleg. F, Left midleg. G, Left hind leg. Scale bars = 1 mm in A–D; 400 μm in E–G." figureDoi="http://doi.org/10.5281/zenodo.5507943" httpUri="https://zenodo.org/record/5507943/files/figure.png" pageId="12" pageNumber="386">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="386" type="materials_examined">
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<paragraph blockId="12.[136,778,1100,2056]" pageId="12" pageNumber="386">
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<emphasis bold="true" box="[136,252,1100,1126]" pageId="12" pageNumber="386">
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<typeStatus box="[136,247,1100,1126]" pageId="12" pageNumber="386">Holotype</typeStatus>
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.
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3353728302" collectionCode="NIGP" country="Myanmar" county="Nanjing" location="In" municipality="Chinese Academy of Sciences" pageId="12" pageNumber="386" specimenCode="NIGP174707" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<typeStatus box="[278,382,1100,1126]" pageId="12" pageNumber="386">Holotype</typeStatus>
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alate,
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<specimenCode box="[494,647,1100,1126]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="12" pageNumber="386">NIGP174707</specimenCode>
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.
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<location LSID="urn:lsid:plazi:treatment:03B28787AB4E0671FF0D87B2FE97F302:8EC4604AAB4E0671FD21801EFD32FF6D" box="[676,699,1101,1127]" country="Myanmar" county="Nanjing" municipality="Chinese Academy of Sciences" name="In" pageId="12" pageNumber="386" stateProvince="Kachin">In</location>
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mid-
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<location LSID="urn:lsid:plazi:treatment:03B28787AB4E0671FF0D87B2FE97F302:8EC4604AAB4E0671FF0D8023FE8DFF80" box="[136,260,1136,1162]" country="Myanmar" county="Nanjing" municipality="Chinese Academy of Sciences" name="Cretaceous" pageId="12" pageNumber="386" stateProvince="Kachin">Cretaceous</location>
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amber from
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<collectingRegion box="[451,656,1136,1162]" country="Myanmar" name="Kachin" pageId="12" pageNumber="386">Kachin Province</collectingRegion>
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, northern
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<collectingCountry box="[136,247,1172,1198]" name="Myanmar" pageId="12" pageNumber="386">Myanmar</collectingCountry>
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.
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<location LSID="urn:lsid:plazi:treatment:03B28787AB4E0671FF0D87B2FE97F302:8EC4604AAB4E0671FE8780C7FEFDFFA4" box="[258,372,1172,1198]" country="Myanmar" county="Nanjing" municipality="Chinese Academy of Sciences" name="Deposited" pageId="12" pageNumber="386" stateProvince="Kachin">Deposited</location>
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in the
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<location LSID="urn:lsid:plazi:treatment:03B28787AB4E0671FF0D87B2FE97F302:8EC4604AAB4E0671FE4180C7FC80FFA4" box="[452,777,1172,1198]" country="Myanmar" county="Nanjing" municipality="Chinese Academy of Sciences" name="Nanjing Institute of Geology" pageId="12" pageNumber="386" stateProvince="Kachin">Nanjing Institute of Geology</location>
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and
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<location LSID="urn:lsid:plazi:treatment:03B28787AB4E0671FF0D87B2FE97F302:8EC4604AAB4E0671FF4280E4FEE1FFDB" box="[199,360,1207,1233]" country="Myanmar" county="Nanjing" municipality="Chinese Academy of Sciences" name="Palaeontology" pageId="12" pageNumber="386" stateProvince="Kachin">Palaeontology</location>
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,
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<collectingMunicipality box="[386,774,1207,1233]" pageId="12" pageNumber="386">Chinese Academy of Sciences</collectingMunicipality>
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,
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<collectingCounty box="[136,229,1243,1269]" pageId="12" pageNumber="386">Nanjing</collectingCounty>
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,
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<collectingCountry box="[240,311,1243,1269]" name="China" pageId="12" pageNumber="386">China</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="386" type="etymology">
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<paragraph blockId="12.[136,778,1100,2056]" pageId="12" pageNumber="386">
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<emphasis bold="true" box="[189,324,1279,1305]" pageId="12" pageNumber="386">Etymology.</emphasis>
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The specific epithet is taken from the Greek term
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<emphasis box="[273,391,1314,1341]" italics="true" pageId="12" pageNumber="386">ΦΙΛέταΙΡΟΣ</emphasis>
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(
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<emphasis box="[408,528,1315,1341]" italics="true" pageId="12" pageNumber="386">philetairos</emphasis>
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) [composed of
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<emphasis box="[709,778,1316,1343]" italics="true" pageId="12" pageNumber="386">ΦῘÌΛΟΣ</emphasis>
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(
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<emphasis box="[145,213,1350,1376]" italics="true" pageId="12" pageNumber="386">phílos</emphasis>
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), meaning, “beloved”, and
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<emphasis box="[562,644,1350,1377]" italics="true" pageId="12" pageNumber="386">ἑταῖΡΟΣ</emphasis>
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(
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<emphasis box="[671,762,1350,1376]" italics="true" pageId="12" pageNumber="386">hetaîros</emphasis>
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), meaning, “friend”), and refers to the relationship between these termites and their inquilines.
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</paragraph>
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</subSubSection>
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<subSubSection box="[189,651,1458,1484]" pageId="12" pageNumber="386" type="diagnosis">
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<paragraph blockId="12.[136,778,1100,2056]" box="[189,651,1458,1484]" pageId="12" pageNumber="386">
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<emphasis bold="true" box="[189,312,1458,1484]" pageId="12" pageNumber="386">Diagnosis.</emphasis>
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As for the genus (
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<emphasis box="[518,635,1458,1484]" italics="true" pageId="12" pageNumber="386">vide supra</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="386" type="description">
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<paragraph blockId="12.[136,778,1100,2056]" pageId="12" pageNumber="386">
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<emphasis bold="true" box="[189,335,1493,1519]" pageId="12" pageNumber="386">Description.</emphasis>
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Imago: Total body length (as preserved)
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<quantity box="[136,244,1529,1556]" metricMagnitude="-3" metricUnit="m" metricValue="5.28" pageId="12" pageNumber="386" unit="mm" value="5.28">5.28 mm</quantity>
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; length of head to apex of labrum
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<quantity box="[666,773,1529,1556]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="12" pageNumber="386" unit="mm" value="1.12">1.12 mm</quantity>
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; head width
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<quantity box="[270,375,1565,1592]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="12" pageNumber="386" unit="mm" value="0.68">0.68 mm</quantity>
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; compound eye diameter
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||
<quantity box="[669,773,1565,1592]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="386" unit="mm" value="0.22">0.22 mm</quantity>
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; distance from compound eye to posterior margin
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<quantity metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="12" pageNumber="386" unit="mm" value="0.36">0.36 mm</quantity>
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; pronotal medial length
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||
<quantity box="[449,551,1636,1663]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="386" unit="mm" value="0.5">0.50 mm</quantity>
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; pronotal width
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<quantity metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="12" pageNumber="386" unit="mm" value="0.87">0.87 mm</quantity>
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||
; metatibial length
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||
<quantity box="[393,497,1672,1699]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="12" pageNumber="386" unit="mm" value="0.58">0.58 mm</quantity>
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||
; length of forewing with scale
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||
<quantity box="[197,299,1708,1735]" metricMagnitude="-3" metricUnit="m" metricValue="6.72" pageId="12" pageNumber="386" unit="mm" value="6.72">6.72 mm</quantity>
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; forewing width
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||
<quantity box="[484,586,1708,1735]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" pageId="12" pageNumber="386" unit="mm" value="2.24">2.24 mm</quantity>
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. Integument dark brown, microsculpturing of head finely imbricate. Setation of head and body sparse; setae not evident on pronotum. Rs with four simple branches, Rs terminating before wing apex; M with four main branches, M terminating at wing apex, basalmost branch terminating at apical quarter of posterior wing margin; CuA pectinately branched, with at least four simple branches (remainder of CuA obscured in
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<typeStatus box="[164,266,1994,2021]" pageId="12" pageNumber="386">holotype</typeStatus>
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), cubital field occupying majority of posterior wing margin.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |