313 lines
50 KiB
XML
313 lines
50 KiB
XML
<document id="DA47B98F11CC101C9F5A92E68D0C5D5A" ID-CLB-Dataset="5071" ID-DOI="10.11646/zootaxa.3857.4.3" ID-GBIF-Dataset="b8178736-7493-4f74-bc85-0188fc50473d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4930056" ID-ZooBank="76021E0C-7542-455B-82F4-C670A3DC8806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623441838049" checkinUser="felipe" docAuthor="Mooi, Rich, Kroh, Andreas & Srivastava, Dinesh K." docDate="2014" docId="0398C0247212D50BFF686B34438CF872" docLanguage="en" docName="zt03857p526.pdf" docOrigin="Zootaxa 3857 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lenicyamidia compta Brunnschweiler 1962" docType="treatment" docVersion="8" lastPageNumber="522" masterDocId="FFA1B85C7201D51EFFFF6C3A425CFFA0" masterDocTitle="Phylogenetic re-evaluation of fossil and extant micro-echinoids with revision of Tridium, Cyamidia, and Lenicyamidia (Echinoidea: Clypeasteroida)" masterLastPageNumber="526" masterPageNumber="501" pageNumber="520" updateTime="1699012996579" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F49F5F217E09DAA5ABA67FADAA4D8798">
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<mods:title id="973B328A71376D3FDD5F9D47FAED26E1">Phylogenetic re-evaluation of fossil and extant micro-echinoids with revision of Tridium, Cyamidia, and Lenicyamidia (Echinoidea: Clypeasteroida)</mods:title>
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<mods:name id="34BCB3079D648DC10F86CEE3EC3A27D1" type="personal">
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<mods:roleTerm id="9C331B631539CD17A00302AC981F99C3">Author</mods:roleTerm>
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<mods:namePart id="0C1BCF45275C674EC656EAEE690E3507">Mooi, Rich</mods:namePart>
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<mods:affiliation id="FC64FB42634EB739AB0E550FC7814731">Department of Invertebrate Zoology and Geology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118, USA. E-mail: rmooi @ calacademy. org</mods:affiliation>
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<mods:name id="06C41865C262EB16CD3C06F7D4FB7287" type="personal">
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<mods:roleTerm id="04BEF709A995BB2D7BF71F8BB3CB4627">Author</mods:roleTerm>
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<mods:namePart id="7BF5B875CEC29DC3F85EBDF613334B08">Kroh, Andreas</mods:namePart>
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<mods:affiliation id="ECD0858D3E3D2EF4586E40AD8BBF0E11">Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria. E-mail: andreas. kroh @ nhm-wien. ac. at</mods:affiliation>
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<mods:name id="538C2606FE67A912AB085991D29117A4" type="personal">
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<mods:namePart id="5C92B35F5B54DD42F4B5CC6799C22E58">Srivastava, Dinesh K.</mods:namePart>
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<mods:affiliation id="CDD7B034C05FBB391E211CCFD0401E07">Centre of Advanced Study in Geology, University of Lucknow, Lucknow 226 007, India. E-mail: deardrdk @ gmail. com Corresponding author</mods:affiliation>
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<mods:typeOfResource id="5FA7164AD6FF2141DD9524186E117B76">text</mods:typeOfResource>
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<mods:title id="634587F0D6BA1A5E9C77A51687E54766">Zootaxa</mods:title>
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<mods:part id="DB6F264045315516026CC8AF111C22AD">
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<mods:date id="97E9BF9E2DBD6D9F19F0AE77D646E36A">2014</mods:date>
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<mods:number id="4A771DEEEE96A4DEEAD218D382CB5181">2014-09-01</mods:number>
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<mods:number id="10CDEAD2472B158880ED13034A9E00BA">3857</mods:number>
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<treatment id="0398C0247212D50BFF686B34438CF872" ID-DOI="http://doi.org/10.5281/zenodo.4929736" ID-GBIF-Taxon="183925366" ID-Zenodo-Dep="4929736" LSID="urn:lsid:plazi:treatment:0398C0247212D50BFF686B34438CF872" httpUri="http://treatment.plazi.org/id/0398C0247212D50BFF686B34438CF872" lastPageId="21" lastPageNumber="522" pageId="19" pageNumber="520">
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<subSubSection id="C32B22B97212D50DFF686B3440CBF887" box="[151,663,1806,1831]" pageId="19" pageNumber="520" type="nomenclature">
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<paragraph id="8B8E71327212D50DFF686B3440CBF887" blockId="19.[151,663,1806,1867]" box="[151,663,1806,1831]" pageId="19" pageNumber="520">
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<heading id="D0C6C65E7212D50DFF686B3440CBF887" bold="true" box="[151,663,1806,1831]" fontSize="10" level="2" pageId="19" pageNumber="520" reason="2">
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<taxonomicName id="4C310AB17212D50DFF686B3440CBF887" authority="Brunnschweiler, 1962" authorityName="Brunnschweiler" authorityYear="1962" box="[151,663,1806,1831]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="19" pageNumber="520" phylum="Echinodermata" rank="species" species="compta">
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<emphasis id="B945AD207212D50DFF686B3440CBF887" bold="true" box="[151,663,1806,1831]" pageId="19" pageNumber="520">
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<emphasis id="B945AD207212D50DFF686B3443D3F886" bold="true" box="[151,399,1806,1831]" italics="true" pageId="19" pageNumber="520">Lenicyamidia compta</emphasis>
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<bibRefCitation id="EFA00CC37212D50DFE6A6B3440CBF887" author="Brunnschweiler, R. O." box="[405,663,1806,1831]" pageId="19" pageNumber="520" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler, 1962</bibRefCitation>
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||
</emphasis>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C32B22B97212D50DFF686B08410FF80F" pageId="19" pageNumber="520" type="description">
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<paragraph id="8B8E71327212D50DFF686B084072F8EB" blockId="19.[151,663,1806,1867]" box="[151,558,1842,1867]" pageId="19" pageNumber="520">
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<figureCitation id="130A6DB77212D50DFF686B08435EF8EB" box="[151,258,1842,1867]" captionStart="FIGURE 4" captionStartId="9.[151,250,935,957]" captionTargetBox="[155,1430,415,893]" captionTargetId="figure-294@9.[151,1436,409,896]" captionTargetPageId="9" captionText="FIGURE 4. Morphology of the peristomial region in six species of fibulariids, including Cyamidia, Tridium (= Fibularia), Lenicyamidia, and Leniechinus. On the left for each species is the oral view depicting tuberculation, sculpturing, and positions of buccal pores and sphaeridial chambers, with accompanying scale bar. On the right for each species is a sagittal section through the test (hatched) around the peristome, anterior to the left, with accompanying scale bar. In all cases, scale bar is 1.0 mm long." figureDoi="http://doi.org/10.5281/zenodo.4930064" httpUri="https://zenodo.org/record/4930064/files/figure.png" pageId="19" pageNumber="520">Figures 4</figureCitation>
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,
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<figureCitation id="130A6DB77212D50DFEF16B094312F8EA" box="[270,334,1843,1866]" captionStart="FIGURE 7" captionStartId="13.[151,250,1004,1026]" captionTargetBox="[228,1360,193,982]" captionTargetId="figure-107@13.[228,1360,193,982]" captionTargetPageId="13" captionText="FIGURE 7. Brunnschweiler's (1962) holotypes: A–C. Cyamidia paucipora (= juvenile Lenicyamidia compta), GA CPC 2825; D–F. Lenicyamidia compta, GA CPC 2827. Aboral (A, D), oral (B, E) and left lateral views (C, F)." figureDoi="http://doi.org/10.5281/zenodo.4930072" httpUri="https://zenodo.org/record/4930072/files/figure.png" pageId="19" pageNumber="520">7D–F</figureCitation>
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,
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<figureCitation id="130A6DB77212D50DFEA66B0843F1F8EB" box="[345,429,1842,1867]" captionStart="FIGURE 10" captionStartId="15.[151,250,1188,1210]" captionTargetBox="[160,1428,189,1167]" captionTargetId="figure-378@15.[151,1436,181,1173]" captionTargetPageId="15" captionText="FIGURE 10. Partial plate architecture of Lenicyamidia compta and Cyamidia paucipora (= juvenile Lenicyamidia compta). A–C. L. compta: A. aboral view of GA CPC 41774; B. oral view of the holotype GA CPC 2827; C. left lateral view of GA CPC 41774; D. C. paucipora, holotype CPC 2825, adapical part of interambulacrum 2. Labelling conventions as in Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.4930078" httpUri="https://zenodo.org/record/4930078/files/figure.png" pageId="19" pageNumber="520">10A–C</figureCitation>
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,
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<figureCitation id="130A6DB77212D50DFE476B084389F8EB" box="[440,469,1842,1867]" captionStart="FIGURE" captionStartId="16.[151,168,836,861]" captionTargetBox="[345,1243,152,809]" captionTargetId="figure-380@16.[341,1249,147,812]" captionTargetPageId="16" captionText="FIGURE 11. Scatter plot of test width (solid squares; n = 30), height (open triangles; n = 27) and respiratory pore number (solid circles; n = 17) vs. test length for the material attributed to Lenicyamidia compta and Cyamidia paucipora by Brunnschweiler (1962). Size range of C. paucipora highlighted by grey backfill. Thumbnail images show outline and profile for the smallest and largest complete specimens. Within profiles, scale bars 1.0 mm long, arrows indicate anterior end. APP = anterior paired petals (i.e. petal II or IV)." figureDoi="http://doi.org/10.5281/zenodo.4930082" httpUri="https://zenodo.org/record/4930082/files/figure.png" pageId="19" pageNumber="520">11</figureCitation>
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,
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<figureCitation id="130A6DB77212D50DFE1E6B084077F8EB" box="[481,555,1842,1867]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="21.[151,250,1052,1074]" captionStartId-1="22.[151,250,1343,1365]" captionTargetBox-0="[226,1363,203,1023]" captionTargetBox-1="[151,1436,181,1331]" captionTargetId-0="figure-393@21.[217,1370,193,1030]" captionTargetId-1="figure-305@22.[151,1436,181,1331]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURE 13. SEM images of the test of three specimens of Lenicyamidia compta. A–C. GA CPC 41774, aboral (A), oral (B) and right lateral view (C); D–F. GA CPC 41773, aboral (D), oral (E) and right lateral view (F)." captionText-1="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi-0="http://doi.org/10.5281/zenodo.4930086" figureDoi-1="http://doi.org/10.5281/zenodo.4930088" httpUri-0="https://zenodo.org/record/4930086/files/figure.png" httpUri-1="https://zenodo.org/record/4930088/files/figure.png" pageId="19" pageNumber="520">13–14</figureCitation>
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.
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</paragraph>
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<paragraph id="8B8E71327212D50DFF686B404141F830" blockId="19.[151,851,1914,1967]" box="[151,797,1914,1936]" pageId="19" pageNumber="520">
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1962
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<taxonomicName id="4C310AB17212D50DFF2F6B404009F830" authority="Brunnschweiler" authorityName="Brunnschweiler" authorityYear="1962" box="[208,597,1914,1936]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="19" pageNumber="520" phylum="Echinodermata" rank="species" species="compta">
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<emphasis id="B945AD207212D50DFF2F6B4043F6F830" box="[208,426,1914,1936]" italics="true" pageId="19" pageNumber="520">Lenicyamidia compta</emphasis>
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Brunnschweiler
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</taxonomicName>
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: 165–169, figs 2–3.
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</paragraph>
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<paragraph id="8B8E71327212D50DFF686BA3410FF80F" blockId="19.[151,851,1914,1967]" box="[151,851,1945,1967]" pageId="19" pageNumber="520">
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1966
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<taxonomicName id="4C310AB17212D50DFF2F6BA3400FF80F" authority="Brunnschweiler" authorityName="Brunnschweiler" authorityYear="1962" box="[208,595,1945,1967]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="19" pageNumber="520" phylum="Echinodermata" rank="species" species="compta">
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<emphasis id="B945AD207212D50DFF2F6BA343F7F80F" box="[208,427,1945,1967]" italics="true" pageId="19" pageNumber="520">Lenicyamidia compta</emphasis>
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Brunnschweiler
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</taxonomicName>
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—Philip: 116–117, fig. 1.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B22B97212D50AFF686BE646F5FEC5" lastPageId="20" lastPageNumber="521" pageId="19" pageNumber="520" type="materials_examined">
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<paragraph id="8B8E71327212D50DFF686BE646DAF855" blockId="19.[151,1158,2012,2037]" box="[151,1158,2012,2037]" pageId="19" pageNumber="520">
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<emphasis id="B945AD207212D50DFF686BE64319F855" bold="true" box="[151,325,2012,2037]" pageId="19" pageNumber="520">Type material.</emphasis>
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<materialsCitation id="3B597B6F7212D50DFEB36BE646DEF855" ID-GBIF-Occurrence="3331843302" box="[332,1154,2012,2037]" collectionCode="CPC" pageId="19" pageNumber="520" specimenCode="CPC 2827, CPC 2828" specimenCount="2" typeStatus="holotype">
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GA
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<specimenCode id="DB97D9497212D50DFE846BE643B3F855" box="[379,495,2012,2037]" collectionCode="CPC" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="19" pageNumber="520">CPC 2827</specimenCode>
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(
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<typeStatus id="548ACF907212D50DFE016BE64038F855" box="[510,612,2012,2037]" pageId="19" pageNumber="520" type="holotype">holotype</typeStatus>
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,
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<figureCitation id="130A6DB77212D50DFD906BE640BBF853" box="[623,743,2012,2037]" captionStart="FIGURE 7" captionStartId="13.[151,250,1004,1026]" captionTargetBox="[228,1360,193,982]" captionTargetId="figure-107@13.[228,1360,193,982]" captionTargetPageId="13" captionText="FIGURE 7. Brunnschweiler's (1962) holotypes: A–C. Cyamidia paucipora (= juvenile Lenicyamidia compta), GA CPC 2825; D–F. Lenicyamidia compta, GA CPC 2827. Aboral (A, D), oral (B, E) and left lateral views (C, F)." figureDoi="http://doi.org/10.5281/zenodo.4930072" httpUri="https://zenodo.org/record/4930072/files/figure.png" pageId="19" pageNumber="520">Figs 7D–F</figureCitation>
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,
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<figureCitation id="130A6DB77212D50DFD0D6BE6417EF855" box="[754,802,2012,2037]" captionStart="FIGURE 10" captionStartId="15.[151,250,1188,1210]" captionTargetBox="[160,1428,189,1167]" captionTargetId="figure-378@15.[151,1436,181,1173]" captionTargetPageId="15" captionText="FIGURE 10. Partial plate architecture of Lenicyamidia compta and Cyamidia paucipora (= juvenile Lenicyamidia compta). A–C. L. compta: A. aboral view of GA CPC 41774; B. oral view of the holotype GA CPC 2827; C. left lateral view of GA CPC 41774; D. C. paucipora, holotype CPC 2825, adapical part of interambulacrum 2. Labelling conventions as in Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.4930078" httpUri="https://zenodo.org/record/4930078/files/figure.png" pageId="19" pageNumber="520">10B</figureCitation>
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) and GA
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<specimenCode id="DB97D9497212D50DFC6F6BE64658F855" box="[912,1028,2012,2037]" collectionCode="CPC" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="19" pageNumber="520">CPC 2828</specimenCode>
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||
(
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||
<typeStatus id="548ACF907212D50DFBEC6BE64626F855" box="[1043,1146,2012,2037]" pageId="19" pageNumber="520" type="paratype">paratype</typeStatus>
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||
)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8E71327215D50AFF386CAD4168FF10" blockId="20.[151,1437,151,1904]" box="[199,820,151,176]" pageId="20" pageNumber="521">
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<emphasis id="B945AD207215D50AFF386CAD43C8FF10" bold="true" box="[199,404,151,176]" inLineHeading="true" pageId="20" pageNumber="521" reason="1">Material studied.</emphasis>
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GA
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<collectionCode id="ED20E9F77215D50AFE346CAD405CFF10" box="[459,512,151,176]" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="20" pageNumber="521">CPC</collectionCode>
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2827–2829, 41773–41799.
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</paragraph>
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<paragraph id="8B8E71327215D50AFF386C8640A6FEBD" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
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||
<materialsCitation id="3B597B6F7215D50AFF386C8640AAFEBD" ID-GBIF-Occurrence="3331843301" latitude="-23.603334" location="Northwest Division" longLatPrecision="21" longitude="114.73722" municipality="Pleiades Hills" pageId="20" pageNumber="521" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<emphasis id="B945AD207215D50AFF386C864339FF75" bold="true" box="[199,357,188,213]" inLineHeading="true" pageId="20" pageNumber="521" reason="1">
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<typeStatus id="548ACF907215D50AFF386C86435EFF75" box="[199,258,188,213]" pageId="20" pageNumber="521">Type</typeStatus>
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locality.
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</emphasis>
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<collectorName id="26C414E47215D50AFE936C8643BCFF75" box="[364,480,188,213]" pageId="20" pageNumber="521">Sample M</collectorName>
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24 from point 221 on airphoto
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<collectorName id="26C414E47215D50AFCC66C86413FFF73" box="[825,867,188,211]" pageId="20" pageNumber="521">No.</collectorName>
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5170 on
|
||
<collectorName id="26C414E47215D50AFC336C8641A7FF73" box="[972,1019,188,211]" pageId="20" pageNumber="521">Run</collectorName>
|
||
2,
|
||
<collectorName id="26C414E47215D50AFBE26C86468FFF75" box="[1053,1235,188,213]" pageId="20" pageNumber="521">Moogooloo Hill</collectorName>
|
||
(hill crest located at
|
||
<geoCoordinate id="EE0517F57215D50AFF496CE54304FF58" box="[182,344,223,248]" degrees="23" direction="south" minutes="36" orientation="latitude" pageId="20" pageNumber="521" precision="15" seconds="12" value="-23.603334">23° 36′ 12″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0517F57215D50AFE966CE54047FF57" box="[361,539,223,248]" degrees="114" direction="east" minutes="44" orientation="longitude" pageId="20" pageNumber="521" precision="15" seconds="14" value="114.73722">114° 44′ 14″ E</geoCoordinate>
|
||
; exact coordinates of sampling locality unknown); about
|
||
<quantity id="4CC9DCD77215D50AFB436CE5474EFF58" box="[1212,1298,223,248]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="20" pageNumber="521" unit="mi" value="8.0">8 miles</quantity>
|
||
SSE of the
|
||
<collectingMunicipality id="6BEAEB487215D50AFF686D3E4368FEBD" box="[151,308,260,285]" pageId="20" pageNumber="521">Pleiades Hills</collectingMunicipality>
|
||
,
|
||
<location id="8EEE27E97215D50AFEBF6D3E4042FEBD" LSID="urn:lsid:plazi:treatment:0398C0247212D50BFF686B34438CF872:8EEE27E97215D50AFEBF6D3E4042FEBD" box="[320,542,260,285]" latitude="-23.603334" longLatPrecision="21" longitude="114.73722" municipality="Pleiades Hills" name="Northwest Division" pageId="20" pageNumber="521" stateProvince="Western Australia">Northwest Division</location>
|
||
,
|
||
<collectingRegion id="49F5BFD07215D50AFDD56D3E40AAFEBD" box="[554,758,260,285]" country="Australia" name="Western Australia" pageId="20" pageNumber="521">Western Australia</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF386D1D46ABFEE0" blockId="20.[151,1437,151,1904]" box="[199,1271,295,320]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386D1D4332FEE0" bold="true" box="[199,366,295,320]" inLineHeading="true" pageId="20" pageNumber="521" reason="1">
|
||
<typeStatus id="548ACF907215D50AFF386D1D435EFEE0" box="[199,258,295,320]" pageId="20" pageNumber="521">Type</typeStatus>
|
||
stratum.
|
||
</emphasis>
|
||
Merlinleigh Sandstone, Late Eocene (see
|
||
<bibRefCitation id="EFA00CC37215D50AFCB76D1D462FFEE0" author="Darragh, T. A. & Kendrick, G. W." box="[840,1139,295,320]" pageId="20" pageNumber="521" pagination="23 - 41" refId="ref11991" refString="Darragh, T. A. & Kendrick, G. W. (2010) Eocene molluscs from the Merlinleigh Sandstone, Carnarvon Basin, Western Australia. Records of the Western Australian Museum, 26, 23 - 41." type="journal article" year="2010">Darragh & Kendrick 2010</bibRefCitation>
|
||
: pp. 24–25)
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF386D7646F5FEC5" blockId="20.[151,1437,151,1904]" box="[199,1193,332,357]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386D7643D9FEC5" bold="true" box="[199,389,332,357]" pageId="20" pageNumber="521">
|
||
ZooBank
|
||
<collectionCode id="ED20E9F77215D50AFEC36D764323FEC5" box="[316,383,332,357]" pageId="20" pageNumber="521">LSID</collectionCode>
|
||
.
|
||
</emphasis>
|
||
<uri id="FFA07D307215D50AFE736D7646F5FEC5" box="[396,1193,332,357]" pageId="20" pageNumber="521">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FF974BE77215D50AFD626D7646F5FEC5" box="[669,1193,332,357]" pageId="20" pageNumber="521">90041316-5191-4933-A119-6C0E598DD5D2</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B22B97215D50BFF386D5540FEFAEA" lastPageId="21" lastPageNumber="522" pageId="20" pageNumber="521" type="description">
|
||
<paragraph id="8B8E71327215D50AFF386D55436DFE70" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386D554304FE28" bold="true" box="[199,344,367,392]" inLineHeading="true" pageId="20" pageNumber="521" reason="1">Description.</emphasis>
|
||
<emphasis id="B945AD207215D50AFE9F6D4A405BFE28" box="[352,519,367,392]" italics="true" pageId="20" pageNumber="521">Size and shape</emphasis>
|
||
—Corona small, not exceeding
|
||
<quantity id="4CC9DCD77215D50AFC926D5541E0FE27" box="[877,956,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="521" unit="mm" value="10.0">10 mm</quantity>
|
||
in TL among available specimens; outline in aboral view forming slightly angular circle; distinctly flattened in profile, maximum height between 30 and 40% of test length.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF386DE646F5FD9D" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386DE643FBFE55" box="[199,423,476,501]" italics="true" pageId="20" pageNumber="521">Internal buttressing</emphasis>
|
||
—Absent, but ambulacral plates bear large pits internally and are about half as thick as interambulacral plates in their centre (
|
||
<figureCitation id="130A6DB77215D50AFDBE6DC540F4FDB8" box="[577,680,511,536]" captionStart="FIGURE 14" captionStartId="22.[151,250,1343,1365]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-305@22.[151,1436,181,1331]" captionTargetPageId="22" captionText="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930088" httpUri="https://zenodo.org/record/4930088/files/figure.png" pageId="20" pageNumber="521">Fig. 14C</figureCitation>
|
||
), explaining peculiarly shaped internal casts (
|
||
<figureCitation id="130A6DB77215D50AFB576DC54751FDB8" box="[1192,1293,511,536]" captionStart="FIGURE 14" captionStartId="22.[151,250,1343,1365]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-305@22.[151,1436,181,1331]" captionTargetPageId="22" captionText="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930088" httpUri="https://zenodo.org/record/4930088/files/figure.png" pageId="20" pageNumber="521">Fig. 14C</figureCitation>
|
||
) reported by
|
||
<bibRefCitation id="EFA00CC37215D50AFF686E1E43C8FD9D" author="Brunnschweiler, R. O." box="[151,404,548,573]" pageId="20" pageNumber="521" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler (1962</bibRefCitation>
|
||
: fig. 2E); pits associated with pores for accessory tube feet (
|
||
<figureCitation id="130A6DB77215D50AFBCA6E1E46C0FD9D" box="[1077,1180,548,573]" captionStart="FIGURE 14" captionStartId="22.[151,250,1343,1365]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-305@22.[151,1436,181,1331]" captionTargetPageId="22" captionText="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930088" httpUri="https://zenodo.org/record/4930088/files/figure.png" pageId="20" pageNumber="521">Fig. 14B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF386E7D434FFD08" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386E7D433BFDC0" box="[199,359,583,608]" italics="true" pageId="20" pageNumber="521">Apical system</emphasis>
|
||
—Situated centrally, at apex of corona; monobasal, with four gonopores and single central hydropore, not situated in pit or groove; ocular pores small and indistinct, lying well outside area enclosed by gonopores.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF386E8E41EFFB95" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386E8E431FFD6D" box="[199,323,692,717]" italics="true" pageId="20" pageNumber="521">Ambulacra</emphasis>
|
||
—Ambulacral plating simple; ambulacra expanding only slightly towards ambitus where they are barely wider than interambulacra; petals short, with up to 21 non-conjugate respiratory pore pairs in each ambulacrum; pore pairs strongly oblique, crossing ambulacral plates; interporal ridge smooth and formed by unperforated stereom; distal pore pairs becoming even more strongly oblique with distance between pores in each pair slightly decreasing; width of interporiferous zones widest halfway along petals and becoming slightly narrower again distally; petaloid region large, extending about 70% of TL; food grooves absent; buccal pores large and located close to peristomial edge, facing horizontally into peristome, externally visible only in broken specimens (
|
||
<figureCitation id="130A6DB77215D50AFEDF6F9543D4FC68" box="[288,392,943,968]" captionStart="FIGURE 14" captionStartId="22.[151,250,1343,1365]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-305@22.[151,1436,181,1331]" captionTargetPageId="22" captionText="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930088" httpUri="https://zenodo.org/record/4930088/files/figure.png" pageId="20" pageNumber="521">Fig. 14E</figureCitation>
|
||
) since they are hidden by transverse stereom bars in oral view; accessory pores evenly distributed all over test on aboral surface (including interporiferous zones and interambulacral plates), but forming patches obliquely crossing ambulacral plates and less common elsewhere on oral surface; ambitus initiating at approximately fourth to fifth pair of ambulacral postbasicoronal plates.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF38680543CAFA3D" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386805432BFBF8" box="[199,375,1087,1112]" italics="true" pageId="20" pageNumber="521">Interambulacra</emphasis>
|
||
—Adapically, two unpaired plates lie in tandem adjacent to apical system; three or four postbasicoronal interambulacrals in each column visible in oral view; posterior unpaired interambulacrum expanding distinctly in region accommodating periproct; basicoronal plates usually extending only to first adjacent ambulacral plate, except in interambulacrum 2, where usually extending to second adjacent ambulacral plate on one or both sides; in largest specimen with plate sutures visible (CPC 41796), basicoronal plates of interambulacra 1, 3 and 4 also extend on one side to second adjacent ambulacral plate due to presence of large spine-bearing tubercles on margins of these plates (usually missing in smaller specimens) and fact that sutures are curved to make room for tubercles slightly modifies plate outline in such a way that they come into contact with second ambulacral plates; accessory pores evenly spread over aboral interambulacra, but less common, though present on oral interambulacral plates.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF38699D471AF98D" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF38699D433FFA60" box="[199,355,1447,1472]" italics="true" pageId="20" pageNumber="521">Tuberculation</emphasis>
|
||
—Primary tubercles crenulate, perforate, homogeneously distributed on aboral surface; on oral surface, in contrast, there is a narrow area between the periproct and peristome (
|
||
<figureCitation id="130A6DB77215D50AFBD769F646C4FA45" box="[1064,1176,1484,1509]" captionStart="FIGURE 14" captionStartId="22.[151,250,1343,1365]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-305@22.[151,1436,181,1331]" captionTargetPageId="22" captionText="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930088" httpUri="https://zenodo.org/record/4930088/files/figure.png" pageId="20" pageNumber="521">Fig. 14 D</figureCitation>
|
||
), as well as anterior of the latter (
|
||
<figureCitation id="130A6DB77215D50AFEF769D5432AF9A8" box="[264,374,1519,1544]" captionStart="FIGURE 14" captionStartId="22.[151,250,1343,1365]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-305@22.[151,1436,181,1331]" captionTargetPageId="22" captionText="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930088" httpUri="https://zenodo.org/record/4930088/files/figure.png" pageId="20" pageNumber="521">Fig. 14 A</figureCitation>
|
||
), free of primary tubercles; instead densely covered with glassy tubercles; lateral to this medial area are located primary tubercles with slightly sunken areoles, these tubercles larger than aboral tubercles.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF386A024024F8A5" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386A024365F9EF" box="[199,313,1592,1615]" italics="true" pageId="20" pageNumber="521">Peristome</emphasis>
|
||
—Larger than periproct, about 18% TL; infundibulum extremely shallow but with near-vertical walls; peristomial opening facing directly downwards almost at midpoint of oral surface; framed by basicoronal circlet in which ambulacral plates only slightly longer than adjacent interambulacrals; plates of basicoronal circlet distinctly longer in posterior half of test; only scattered single tubercles present on interambulacral basicoronals adjacent to peristome, mainly on larger specimens; peristome framed by broken circle of circumferential bars overlying sphaeridial chambers (
|
||
<figureCitation id="130A6DB77215D50AFDF96AD64037F8A5" box="[518,619,1772,1797]" captionStart="FIGURE 14" captionStartId="22.[151,250,1343,1365]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-305@22.[151,1436,181,1331]" captionTargetPageId="22" captionText="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930088" httpUri="https://zenodo.org/record/4930088/files/figure.png" pageId="20" pageNumber="521">Fig. 14E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E71327215D50AFF386B2A41EDF8D0" blockId="20.[151,1437,151,1904]" pageId="20" pageNumber="521">
|
||
<emphasis id="B945AD207215D50AFF386B2A436DF887" box="[199,305,1808,1831]" italics="true" pageId="20" pageNumber="521">Periproct</emphasis>
|
||
—Small, approximately 12% TL; facing down- and slightly backwards; located close to posterior margin on oral side; distinctly elongated along anterior-posterior axis; bounded by first (5.a.2, 5.b.2), second (5.a.3, 5.b.3), and one plate (5.b.4) of the third pair of post-basicoronal plates.
|
||
</paragraph>
|
||
<caption id="DF4E21BA7214D50BFF686826460AFBF0" ID-DOI="http://doi.org/10.5281/zenodo.4930086" ID-Zenodo-Dep="4930086" httpUri="https://zenodo.org/record/4930086/files/figure.png" pageId="21" pageNumber="522" startId="21.[151,250,1052,1074]" targetBox="[226,1363,203,1023]" targetPageId="21">
|
||
<paragraph id="8B8E71327214D50BFF686826460AFBF0" blockId="21.[151,1436,1052,1104]" pageId="21" pageNumber="522">
|
||
<emphasis id="B945AD207214D50BFF686826437CFB91" bold="true" box="[151,288,1052,1074]" pageId="21" pageNumber="522">FIGURE 13.</emphasis>
|
||
SEM images of the test of three specimens of
|
||
<taxonomicName id="4C310AB17214D50BFCFF68264187FB91" authorityName="Brunnschweiler" authorityYear="1962" box="[768,987,1052,1074]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD207214D50BFCFF68264187FB91" box="[768,987,1052,1074]" italics="true" pageId="21" pageNumber="522">Lenicyamidia compta</emphasis>
|
||
</taxonomicName>
|
||
. A–C. GA CPC 41774, aboral (A), oral (B) and right lateral view (C); D–F. GA CPC 41773, aboral (D), oral (E) and right lateral view (F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E71327214D50BFF38684640B7FB7E" blockId="21.[151,1436,1148,2002]" pageId="21" pageNumber="522">
|
||
<emphasis id="B945AD207214D50BFF38684643C5FB35" box="[199,409,1148,1173]" italics="true" pageId="21" pageNumber="522">Perignathic girdle</emphasis>
|
||
—Consisting of five small processes (auricles) attached to the internal surface of each interambulacral basicoronal; on their adoral side a pair of minute depressions (likely muscle scars for insertion of lantern protractors) located in each interambulacrum.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327214D50BFF3868D343D0FA84" blockId="21.[151,1436,1148,2002]" pageId="21" pageNumber="522">
|
||
<taxonomicName id="4C310AB17214D50BFF3868D3431EFAA2" box="[199,322,1257,1282]" class="Entognatha" family="Sminthurididae" genus="Sphaeridia" kingdom="Animalia" order="Collembola" pageId="21" pageNumber="522" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945AD207214D50BFF3868D3431EFAA2" box="[199,322,1257,1282]" italics="true" pageId="21" pageNumber="522">Sphaeridia</emphasis>
|
||
</taxonomicName>
|
||
—One per ambulacrum; fully enclosed (
|
||
<figureCitation id="130A6DB77214D50BFCEF68D34107FAA2" box="[784,859,1257,1282]" captionStart="FIGURE 4" captionStartId="9.[151,250,935,957]" captionTargetBox="[155,1430,415,893]" captionTargetId="figure-294@9.[151,1436,409,896]" captionTargetPageId="9" captionText="FIGURE 4. Morphology of the peristomial region in six species of fibulariids, including Cyamidia, Tridium (= Fibularia), Lenicyamidia, and Leniechinus. On the left for each species is the oral view depicting tuberculation, sculpturing, and positions of buccal pores and sphaeridial chambers, with accompanying scale bar. On the right for each species is a sagittal section through the test (hatched) around the peristome, anterior to the left, with accompanying scale bar. In all cases, scale bar is 1.0 mm long." figureDoi="http://doi.org/10.5281/zenodo.4930064" httpUri="https://zenodo.org/record/4930064/files/figure.png" pageId="21" pageNumber="522">
|
||
Figs
|
||
<geoCoordinate id="EE0517F57214D50BFCB468D34107FAA2" box="[843,859,1257,1282]" degrees="4, 14" direction="east" orientation="longitude" pageId="21" pageNumber="522" precision="555" value="4.14">4</geoCoordinate>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="130A6DB77214D50BFC9668D341C6FAA2" box="[873,922,1257,1282]" captionStart="FIGURE 14" captionStartId="22.[151,250,1343,1365]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-305@22.[151,1436,181,1331]" captionTargetPageId="22" captionText="FIGURE 14. SEM images showing details on test surface of Lenicyamidia compta. A. Ornamentation of test just anterior of the peristome in specimen GA CPC 41773. B. Internal view of oral ambulacra and peristome (lower part of image) in specimen GA CPC 41775 showing perignatic girdle and sunken fields of accessory pores. C. Internal mould of specimen GA CPC 41776, showing sunken fields of accessory pores as elevated ridges and thickness of the test at the ambitus. D. Ornamentation of test just posterior of the peristome in specimen GA CPC 41774. E. Cross section through peristomial region of specimen GA CPC 41777 showing buccal pores and sphaeridial chamber. All scale bars 500 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930088" httpUri="https://zenodo.org/record/4930088/files/figure.png" pageId="21" pageNumber="522">14E</figureCitation>
|
||
); situated beneath distinct transverse bar just distal to buccal pores.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327214D50BFF38690B40FEFAEA" blockId="21.[151,1436,1148,2002]" box="[199,674,1329,1354]" pageId="21" pageNumber="522">
|
||
<emphasis id="B945AD207214D50BFF38690B404DFAEA" box="[199,529,1329,1354]" italics="true" pageId="21" pageNumber="522">Spines, pedicellariae, lantern</emphasis>
|
||
—Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B22B97214D50BFF38696F438CF872" pageId="21" pageNumber="522" type="discussion">
|
||
<paragraph id="8B8E71327214D50BFF38696F40ACFA5E" blockId="21.[151,1436,1148,2002]" pageId="21" pageNumber="522">
|
||
<emphasis id="B945AD207214D50BFF38696F4367FACE" bold="true" box="[199,315,1365,1390]" inLineHeading="true" pageId="21" pageNumber="522" reason="1">Remarks.</emphasis>
|
||
<bibRefCitation id="EFA00CC37214D50BFEB7696F400EFACE" author="Brunnschweiler, R. O." box="[328,594,1365,1390]" pageId="21" pageNumber="522" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler (1962)</bibRefCitation>
|
||
illustrated the apical disc as being composed of four genital plus an additional central plate. However,
|
||
<bibRefCitation id="EFA00CC37214D50BFDE8694340F7FA32" author="Philip, G. M." box="[535,683,1401,1426]" pageId="21" pageNumber="522" pagination="114 - 117" refId="ref13179" refString="Philip, G. M. (1966) Notes on three recently proposed Australian Tertiary echinoid genera. Proceedings of the Linnean Society of New South Wales, 91, 114 - 117." type="journal article" year="1966">Philip (1966)</bibRefCitation>
|
||
showed that the apical disc was of typical clypeasteroid monobasal structure. The illustration of an “internal mold” by
|
||
<bibRefCitation id="EFA00CC37214D50BFD1F69A74183FA16" author="Brunnschweiler, R. O." box="[736,991,1437,1462]" pageId="21" pageNumber="522" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler (1962</bibRefCitation>
|
||
: fig. 3) in reality represents an internal view of the oral surface (and thus shows a mirror image of the outside plating pattern). This accounts for the apparent deviation from Lovén’s Rule in his drawing.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327214D50BFF386A334347F92C" blockId="21.[151,1436,1148,2002]" pageId="21" pageNumber="522">
|
||
The peristomial region of
|
||
<taxonomicName id="4C310AB17214D50BFE0E6A30403EF981" authorityName="Brunnschweiler" authorityYear="1962" box="[497,610,1546,1569]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD207214D50BFE0E6A30403EF981" box="[497,610,1546,1569]" italics="true" pageId="21" pageNumber="522">L. compta</emphasis>
|
||
</taxonomicName>
|
||
is markedly different from that of other fibulariids studied here (
|
||
<figureCitation id="130A6DB77214D50BFAB76A3347D2F982" box="[1352,1422,1545,1570]" captionStart="FIGURE 4" captionStartId="9.[151,250,935,957]" captionTargetBox="[155,1430,415,893]" captionTargetId="figure-294@9.[151,1436,409,896]" captionTargetPageId="9" captionText="FIGURE 4. Morphology of the peristomial region in six species of fibulariids, including Cyamidia, Tridium (= Fibularia), Lenicyamidia, and Leniechinus. On the left for each species is the oral view depicting tuberculation, sculpturing, and positions of buccal pores and sphaeridial chambers, with accompanying scale bar. On the right for each species is a sagittal section through the test (hatched) around the peristome, anterior to the left, with accompanying scale bar. In all cases, scale bar is 1.0 mm long." figureDoi="http://doi.org/10.5281/zenodo.4930064" httpUri="https://zenodo.org/record/4930064/files/figure.png" pageId="21" pageNumber="522">Fig. 4</figureCitation>
|
||
). There are no clusters of tubercles on interambulacral basicoronals, the sides of the peristomial infundibulum are very steep and the circumferential stereom bars are very close to the peristomial edge, almost overhanging it in some cases.
|
||
</paragraph>
|
||
<paragraph id="8B8E71327214D50BFF386AA3438CF872" blockId="21.[151,1436,1148,2002]" pageId="21" pageNumber="522">
|
||
The oral tuberculation of
|
||
<taxonomicName id="4C310AB17214D50BFE116AA340DAF912" authorityName="Brunnschweiler" authorityYear="1962" box="[494,646,1689,1714]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207214D50BFE116AA340DAF912" box="[494,646,1689,1714]" italics="true" pageId="21" pageNumber="522">Lenicyamidia</emphasis>
|
||
</taxonomicName>
|
||
was compared with that of
|
||
<taxonomicName id="4C310AB17214D50BFC336AA04648F911" authorityName="Desor" authorityYear="1847" box="[972,1044,1690,1713]" class="Echinoidea" family="Scutellinidae" genus="Lenita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207214D50BFC336AA04648F911" box="[972,1044,1690,1713]" italics="true" pageId="21" pageNumber="522">Lenita</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFA00CC37214D50BFBBD6AA34716F912" author="Brunnschweiler, R. O." box="[1090,1354,1689,1714]" pageId="21" pageNumber="522" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler (1962)</bibRefCitation>
|
||
. While both possess a medial zone free from spine-bearing tubercles, there are two obvious differences: 1) in
|
||
<taxonomicName id="4C310AB17214D50BFAD66A84472DF975" authorityName="Desor" authorityYear="1847" box="[1321,1393,1726,1749]" class="Echinoidea" family="Scutellinidae" genus="Lenita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207214D50BFAD66A84472DF975" box="[1321,1393,1726,1749]" italics="true" pageId="21" pageNumber="522">Lenita</emphasis>
|
||
</taxonomicName>
|
||
the tubercles have deeply incised, asymmetrical areoles (these are shallow and near symmetrical in
|
||
<taxonomicName id="4C310AB17214D50BFB3C6ADB4707F95A" authorityName="Brunnschweiler" authorityYear="1962" box="[1219,1371,1761,1786]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207214D50BFB3C6ADB4707F95A" box="[1219,1371,1761,1786]" italics="true" pageId="21" pageNumber="522">Lenicyamidia</emphasis>
|
||
</taxonomicName>
|
||
); and 2) large glassy tubercles are lacking in this zone in
|
||
<taxonomicName id="4C310AB17214D50BFD276B3C4143F8BD" authorityName="Desor" authorityYear="1847" box="[728,799,1798,1821]" class="Echinoidea" family="Scutellinidae" genus="Lenita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207214D50BFD276B3C4143F8BD" box="[728,799,1798,1821]" italics="true" pageId="21" pageNumber="522">Lenita</emphasis>
|
||
</taxonomicName>
|
||
. It seems likely, therefore, that this tuberculation pattern is not homologous, but has adaptive significance causing it to have arisen independently in several neognathostome clades. In addition,
|
||
<taxonomicName id="4C310AB17214D50BFE836B744398F8C5" authorityName="Desor" authorityYear="1847" box="[380,452,1870,1893]" class="Echinoidea" family="Scutellinidae" genus="Lenita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207214D50BFE836B744398F8C5" box="[380,452,1870,1893]" italics="true" pageId="21" pageNumber="522">Lenita</emphasis>
|
||
</taxonomicName>
|
||
, which is an early scutelline, differs in many other respects, including possession of biserial interambulacra at the apex, paired basicoronals, and other features not found in laganiforms. In contrast,
|
||
<taxonomicName id="4C310AB17214D50BFF686BAF4373F80E" authorityName="Brunnschweiler" authorityYear="1962" box="[151,303,1941,1966]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="21" pageNumber="522" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207214D50BFF686BAF4373F80E" box="[151,303,1941,1966]" italics="true" pageId="21" pageNumber="522">Lenicyamidia</emphasis>
|
||
</taxonomicName>
|
||
, is clearly a laganiform, having two uniserial plates in the adapical interambulacra and a basicoronal circlet typical of fibulariids.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |