399 lines
57 KiB
XML
399 lines
57 KiB
XML
<document id="4E8762FED76D733938488B0A96257AF8" ID-CLB-Dataset="5071" ID-DOI="10.11646/zootaxa.3857.4.3" ID-GBIF-Dataset="b8178736-7493-4f74-bc85-0188fc50473d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4930056" ID-ZooBank="76021E0C-7542-455B-82F4-C670A3DC8806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623441838049" checkinUser="felipe" docAuthor="Mooi, Rich, Kroh, Andreas & Srivastava, Dinesh K." docDate="2014" docId="0398C024720DD50EFF6869F143A4FACD" docLanguage="en" docName="zt03857p526.pdf" docOrigin="Zootaxa 3857 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyamidia paucipora Brunnschweiler 1962" docType="treatment" docVersion="8" lastPageNumber="517" masterDocId="FFA1B85C7201D51EFFFF6C3A425CFFA0" masterDocTitle="Phylogenetic re-evaluation of fossil and extant micro-echinoids with revision of Tridium, Cyamidia, and Lenicyamidia (Echinoidea: Clypeasteroida)" masterLastPageNumber="526" masterPageNumber="501" pageNumber="513" updateTime="1699012996579" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="CB4F8F6A523FBD7F56ADDE3ECC4ADD82" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="22D8D46B656A916E03EAC6F03D76C215">
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<mods:title id="160C38DEEBF30F72CA784DB2B330126C">Phylogenetic re-evaluation of fossil and extant micro-echinoids with revision of Tridium, Cyamidia, and Lenicyamidia (Echinoidea: Clypeasteroida)</mods:title>
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<mods:name id="70C3FD23B268E47F504E77CD79D51B80" type="personal">
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<mods:roleTerm id="6735646080E187F703C7C69D43DFFC35">Author</mods:roleTerm>
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<mods:namePart id="39A0CC97F880F91E10C80C7C6F2633E8">Mooi, Rich</mods:namePart>
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<mods:affiliation id="1C1AAF952F5A7F120D0EE8E1325B30C4">Department of Invertebrate Zoology and Geology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118, USA. E-mail: rmooi @ calacademy. org</mods:affiliation>
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<mods:name id="C0C086F164B6B76324E03DB755EEB3D8" type="personal">
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<mods:roleTerm id="98BEC60438C317955B35E4E386E2B8E8">Author</mods:roleTerm>
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<mods:namePart id="474AB90E837540A75766D3335AC6C48D">Kroh, Andreas</mods:namePart>
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<mods:affiliation id="9F99C8F3AA89399C78083A353E1CED7C">Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria. E-mail: andreas. kroh @ nhm-wien. ac. at</mods:affiliation>
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<mods:name id="F49C56BBBCBAF0DB9EB99EDFB8E17340" type="personal">
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<mods:roleTerm id="DE77AA9EE70CF0BB0C522F485C8DA1CD">Author</mods:roleTerm>
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<mods:namePart id="4CC3B873A08F24C1ACBF760EA7819207">Srivastava, Dinesh K.</mods:namePart>
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<mods:affiliation id="800E0AF42EE56692CD9112B2377A4C3E">Centre of Advanced Study in Geology, University of Lucknow, Lucknow 226 007, India. E-mail: deardrdk @ gmail. com Corresponding author</mods:affiliation>
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<mods:title id="FE087808522D2A2DF0484205520B8F1C">Zootaxa</mods:title>
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<mods:part id="C04D85CC97504F4D61FFF5E2314C819B">
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<mods:date id="BC80579EAE23F59F26656EF06887AF83">2014</mods:date>
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<mods:number id="8BF0D129F2BF91A6EAEE4DD9D98559F4">2014-09-01</mods:number>
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<mods:number id="3C6B7AC38C46A56AE3978A6044CF66F7">3857</mods:number>
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<treatment id="0398C024720DD50EFF6869F143A4FACD" ID-DOI="http://doi.org/10.5281/zenodo.4929717" ID-GBIF-Taxon="183925371" ID-Zenodo-Dep="4929717" LSID="urn:lsid:plazi:treatment:0398C024720DD50EFF6869F143A4FACD" httpUri="http://treatment.plazi.org/id/0398C024720DD50EFF6869F143A4FACD" lastPageId="16" lastPageNumber="517" pageId="12" pageNumber="513">
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<subSubSection id="C32B22B9720DD512FF6869F140D5FA45" box="[151,649,1483,1509]" pageId="12" pageNumber="513" type="nomenclature">
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<paragraph id="8B8E7132720DD512FF6869F140D5FA45" blockId="12.[151,649,1483,1544]" box="[151,649,1483,1509]" pageId="12" pageNumber="513">
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<heading id="D0C6C65E720DD512FF6869F140D5FA45" bold="true" box="[151,649,1483,1509]" fontSize="10" level="2" pageId="12" pageNumber="513" reason="2">
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<taxonomicName id="4C310AB1720DD512FF6869F140D5FA45" authority="Brunnschweiler, 1962" authorityName="Brunnschweiler" authorityYear="1962" box="[151,649,1483,1509]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="12" pageNumber="513" phylum="Echinodermata" rank="species" species="paucipora">
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<emphasis id="B945AD20720DD512FF6869F140D5FA45" bold="true" box="[151,649,1483,1509]" pageId="12" pageNumber="513">
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<emphasis id="B945AD20720DD512FF6869F143DCFA44" bold="true" box="[151,384,1483,1508]" italics="true" pageId="12" pageNumber="513">Cyamidia paucipora</emphasis>
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<bibRefCitation id="EFA00CC3720DD512FE7769F640D5FA45" author="Brunnschweiler, R. O." box="[392,649,1484,1509]" pageId="12" pageNumber="513" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler, 1962</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B22B9720DD512FF6869D54174F9E7" pageId="12" pageNumber="513" type="description">
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<paragraph id="8B8E7132720DD512FF6869D54387F9A8" blockId="12.[151,649,1483,1544]" box="[151,475,1519,1544]" pageId="12" pageNumber="513">
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<figureCitation id="130A6DB7720DD512FF6869D5436BF9A8" box="[151,311,1519,1544]" captionStart="FIGURE 7" captionStartId="13.[151,250,1004,1026]" captionTargetBox="[228,1360,193,982]" captionTargetId="figure-107@13.[228,1360,193,982]" captionTargetPageId="13" captionText="FIGURE 7. Brunnschweiler's (1962) holotypes: A–C. Cyamidia paucipora (= juvenile Lenicyamidia compta), GA CPC 2825; D–F. Lenicyamidia compta, GA CPC 2827. Aboral (A, D), oral (B, E) and left lateral views (C, F)." figureDoi="http://doi.org/10.5281/zenodo.4930072" httpUri="https://zenodo.org/record/4930072/files/figure.png" pageId="12" pageNumber="513">Figures 7A–C</figureCitation>
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,
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<figureCitation id="130A6DB7720DD512FEBC69D54332F9A8" box="[323,366,1519,1544]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="13.[151,250,1864,1886]" captionStartId-1="14.[151,250,779,801]" captionTargetBox-0="[261,1326,1114,1836]" captionTargetBox-1="[151,1436,194,758]" captionTargetId-0="figure-116@13.[253,1334,1106,1842]" captionTargetId-1="figure-525@14.[151,1436,193,758]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 8. SEM images of the test of three specimens of Cyamidia paucipora (= juvenile Lenicyamidia compta). A. GA CPC 41767, aboral view. B–D: GA CPC 41768, aboral (B), oral (C) and left lateral view (D); E–G: GA CPC 41769, aboral (E), oral (F) and left lateral view (G)." captionText-1="FIGURE 9. SEM images showing details on test surface of Cyamidia paucipora (= juvenile Lenicyamidia compta). A. Apical region of specimen GA CPC 41767, pores of each respiratory tube foot linked by dotted lines, other apertures labelled: H = hydropore; G1, G2, G3, G4 = gonopores of interambulacra 1, 2, 3, and 4, respectively; ocular pores indicated by arrows. B. Ornamentation of test just anterior of the peristome in specimen GA CPC 41768. C. Peristomial region of specimen GA CPC 41769 showing peristomial region and periproct (bottom of image). All scale bars 200 µm long." figureDoi-0="http://doi.org/10.5281/zenodo.4930074" figureDoi-1="http://doi.org/10.5281/zenodo.4930076" httpUri-0="https://zenodo.org/record/4930074/files/figure.png" httpUri-1="https://zenodo.org/record/4930076/files/figure.png" pageId="12" pageNumber="513">8–9</figureCitation>
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,
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<figureCitation id="130A6DB7720DD512FE8469D543F1F9A8" box="[379,429,1519,1544]" captionStart="FIGURE 10" captionStartId="15.[151,250,1188,1210]" captionTargetBox="[160,1428,189,1167]" captionTargetId="figure-378@15.[151,1436,181,1173]" captionTargetPageId="15" captionText="FIGURE 10. Partial plate architecture of Lenicyamidia compta and Cyamidia paucipora (= juvenile Lenicyamidia compta). A–C. L. compta: A. aboral view of GA CPC 41774; B. oral view of the holotype GA CPC 2827; C. left lateral view of GA CPC 41774; D. C. paucipora, holotype CPC 2825, adapical part of interambulacrum 2. Labelling conventions as in Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.4930078" httpUri="https://zenodo.org/record/4930078/files/figure.png" pageId="12" pageNumber="513">10D</figureCitation>
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,
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<figureCitation id="130A6DB7720DD512FE4669D5438BF9A8" box="[441,471,1519,1544]" captionStart="FIGURE" captionStartId="16.[151,168,836,861]" captionTargetBox="[345,1243,152,809]" captionTargetId="figure-380@16.[341,1249,147,812]" captionTargetPageId="16" captionText="FIGURE 11. Scatter plot of test width (solid squares; n = 30), height (open triangles; n = 27) and respiratory pore number (solid circles; n = 17) vs. test length for the material attributed to Lenicyamidia compta and Cyamidia paucipora by Brunnschweiler (1962). Size range of C. paucipora highlighted by grey backfill. Thumbnail images show outline and profile for the smallest and largest complete specimens. Within profiles, scale bars 1.0 mm long, arrows indicate anterior end. APP = anterior paired petals (i.e. petal II or IV)." figureDoi="http://doi.org/10.5281/zenodo.4930082" httpUri="https://zenodo.org/record/4930082/files/figure.png" pageId="12" pageNumber="513">11</figureCitation>
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.
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</paragraph>
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<paragraph id="8B8E7132720DD512FF686A0B4174F9E7" blockId="12.[151,808,1585,1607]" box="[151,808,1585,1607]" pageId="12" pageNumber="513">
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1962
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<taxonomicName id="4C310AB1720DD512FF2F6A0B4017F9E7" authority="Brunnschweiler" authorityName="Brunnschweiler" authorityYear="1962" box="[208,587,1585,1607]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="12" pageNumber="513" phylum="Echinodermata" rank="species" species="paucipora">
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<emphasis id="B945AD20720DD512FF2F6A0B43FDF9E7" box="[208,417,1585,1607]" italics="true" pageId="12" pageNumber="513">Cyamidia paucipora</emphasis>
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Brunnschweiler
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</taxonomicName>
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: 162–164; fig. 1A–D.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C32B22B9720DD512FF686A57468FF822" pageId="12" pageNumber="513" type="materials_examined">
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<paragraph id="8B8E7132720DD512FF686A574165F926" blockId="12.[151,1436,1645,2030]" box="[151,825,1645,1670]" pageId="12" pageNumber="513">
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<emphasis id="B945AD20720DD512FF686A574319F926" bold="true" box="[151,325,1645,1670]" inLineHeading="true" pageId="12" pageNumber="513" reason="1">Type material.</emphasis>
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<materialsCitation id="3B597B6F720DD512FEB36A574169F926" ID-GBIF-Occurrence="3331843306" box="[332,821,1645,1670]" collectionCode="CPC" pageId="12" pageNumber="513" specimenCode="CPC 2825" specimenCount="1" typeStatus="holotype">
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GA
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<specimenCode id="DB97D949720DD512FE846A5743B3F926" box="[379,495,1645,1670]" collectionCode="CPC" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="12" pageNumber="513">CPC 2825</specimenCode>
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(
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<typeStatus id="548ACF90720DD512FE016A574039F926" box="[510,613,1645,1670]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
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;
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<figureCitation id="130A6DB7720DD512FD8F6A5740B1F926" box="[624,749,1645,1670]" captionStart="FIGURE 7" captionStartId="13.[151,250,1004,1026]" captionTargetBox="[228,1360,193,982]" captionTargetId="figure-107@13.[228,1360,193,982]" captionTargetPageId="13" captionText="FIGURE 7. Brunnschweiler's (1962) holotypes: A–C. Cyamidia paucipora (= juvenile Lenicyamidia compta), GA CPC 2825; D–F. Lenicyamidia compta, GA CPC 2827. Aboral (A, D), oral (B, E) and left lateral views (C, F)." figureDoi="http://doi.org/10.5281/zenodo.4930072" httpUri="https://zenodo.org/record/4930072/files/figure.png" pageId="12" pageNumber="513">Figs 7A–C</figureCitation>
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,
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<figureCitation id="130A6DB7720DD512FD076A574171F926" box="[760,813,1645,1670]" captionStart="FIGURE 10" captionStartId="15.[151,250,1188,1210]" captionTargetBox="[160,1428,189,1167]" captionTargetId="figure-378@15.[151,1436,181,1173]" captionTargetPageId="15" captionText="FIGURE 10. Partial plate architecture of Lenicyamidia compta and Cyamidia paucipora (= juvenile Lenicyamidia compta). A–C. L. compta: A. aboral view of GA CPC 41774; B. oral view of the holotype GA CPC 2827; C. left lateral view of GA CPC 41774; D. C. paucipora, holotype CPC 2825, adapical part of interambulacrum 2. Labelling conventions as in Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.4930078" httpUri="https://zenodo.org/record/4930078/files/figure.png" pageId="12" pageNumber="513">10D</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8E7132720DD512FF386AAB43FDF96E" blockId="12.[151,1436,1645,2030]" pageId="12" pageNumber="513">
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<emphasis id="B945AD20720DD512FF386AAB43FDF90A" bold="true" box="[199,417,1681,1706]" inLineHeading="true" pageId="12" pageNumber="513" reason="1">Material studied.</emphasis>
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<materialsCitation id="3B597B6F720DD512FE4A6AA843C1F96E" ID-GBIF-Occurrence="3331843307" collectionCode="CPC" pageId="12" pageNumber="513" specimenCode="CPC 2826, 41767-41772" specimenCount="1" typeStatus="holotype">
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In addition to the
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<typeStatus id="548ACF90720DD512FD4E6AAB414AF90A" box="[689,790,1681,1706]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
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, 7 “topotypes” (GA
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<specimenCode id="DB97D949720DD512FBCE6AAB46E9F90A" box="[1073,1205,1681,1706]" collectionCode="CPC" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="12" pageNumber="513">CPC 2826</specimenCode>
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||
,
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<specimenCode id="DB97D949720DD512FB326AAB4735F90A" box="[1229,1385,1681,1706]" collectionCode="CPC" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="12" pageNumber="513">41767–41772</specimenCode>
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) of
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<bibRefCitation id="EFA00CC3720DD512FF686A8F43C1F96E" author="Brunnschweiler, R. O." box="[151,413,1717,1742]" pageId="12" pageNumber="513" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler (1962)</bibRefCitation>
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</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="8B8E7132720DD512FF386AE340A6F89A" blockId="12.[151,1436,1645,2030]" pageId="12" pageNumber="513">
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<materialsCitation id="3B597B6F720DD512FF386AE340AAF89A" ID-GBIF-Occurrence="3331843308" latitude="-23.603334" location="Northwest Division" longLatPrecision="21" longitude="114.73722" municipality="Pleiades Hills" pageId="12" pageNumber="513" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<emphasis id="B945AD20720DD512FF386AE34339F952" bold="true" box="[199,357,1753,1778]" inLineHeading="true" pageId="12" pageNumber="513" reason="1">
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<typeStatus id="548ACF90720DD512FF386AE3435EF952" box="[199,258,1753,1778]" pageId="12" pageNumber="513">Type</typeStatus>
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||
locality.
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</emphasis>
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<collectorName id="26C414E4720DD512FE936AE343BCF952" box="[364,480,1753,1778]" pageId="12" pageNumber="513">Sample M</collectorName>
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24 from point 221 on airphoto
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<collectorName id="26C414E4720DD512FCC66AE0413FF951" box="[825,867,1754,1777]" pageId="12" pageNumber="513">No.</collectorName>
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5170 on
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<collectorName id="26C414E4720DD512FC336AE041A7F951" box="[972,1019,1754,1777]" pageId="12" pageNumber="513">Run</collectorName>
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2,
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<collectorName id="26C414E4720DD512FBE26AE3468FF952" box="[1053,1235,1753,1778]" pageId="12" pageNumber="513">Moogooloo Hill</collectorName>
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(hill crest located at
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,
|
||
<geoCoordinate id="EE0517F5720DD512FE966AC74047F8B5" box="[361,539,1789,1814]" degrees="114" direction="east" minutes="44" orientation="longitude" pageId="12" pageNumber="513" precision="15" seconds="14" value="114.73722">114° 44′ 14″ E</geoCoordinate>
|
||
; exact coordinates of sampling locality unknown); about
|
||
<quantity id="4CC9DCD7720DD512FB436AC7474EF8B6" box="[1212,1298,1789,1814]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="12" pageNumber="513" unit="mi" value="8.0">8 miles</quantity>
|
||
SSE of the
|
||
<collectingMunicipality id="6BEAEB48720DD512FF686B1B4368F89A" box="[151,308,1825,1850]" pageId="12" pageNumber="513">Pleiades Hills</collectingMunicipality>
|
||
,
|
||
<location id="8EEE27E9720DD512FEBF6B1B4042F89A" LSID="urn:lsid:plazi:treatment:0398C024720DD50EFF6869F143A4FACD:8EEE27E9720DD512FEBF6B1B4042F89A" box="[320,542,1825,1850]" latitude="-23.603334" longLatPrecision="21" longitude="114.73722" municipality="Pleiades Hills" name="Northwest Division" pageId="12" pageNumber="513" stateProvince="Western Australia">Northwest Division</location>
|
||
,
|
||
<collectingRegion id="49F5BFD0720DD512FDD56B1840AAF89A" box="[554,758,1825,1850]" country="Australia" name="Western Australia" pageId="12" pageNumber="513">Western Australia</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720DD512FF386B7F46ABF8FE" blockId="12.[151,1436,1645,2030]" box="[199,1271,1861,1886]" pageId="12" pageNumber="513">
|
||
<emphasis id="B945AD20720DD512FF386B7F4332F8FE" bold="true" box="[199,366,1861,1886]" inLineHeading="true" pageId="12" pageNumber="513" reason="1">
|
||
<typeStatus id="548ACF90720DD512FF386B7F435EF8FE" box="[199,258,1861,1886]" pageId="12" pageNumber="513">Type</typeStatus>
|
||
stratum.
|
||
</emphasis>
|
||
Merlinleigh Sandstone, Late Eocene (see
|
||
<bibRefCitation id="EFA00CC3720DD512FCB76B7F462FF8FE" author="Darragh, T. A. & Kendrick, G. W." box="[840,1139,1861,1886]" pageId="12" pageNumber="513" pagination="23 - 41" refId="ref11991" refString="Darragh, T. A. & Kendrick, G. W. (2010) Eocene molluscs from the Merlinleigh Sandstone, Carnarvon Basin, Western Australia. Records of the Western Australian Museum, 26, 23 - 41." type="journal article" year="2010">Darragh & Kendrick 2010</bibRefCitation>
|
||
: pp. 24–25)
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720DD512FF386B53468FF822" blockId="12.[151,1436,1645,2030]" box="[199,1235,1897,1922]" pageId="12" pageNumber="513">
|
||
<emphasis id="B945AD20720DD512FF386B5343D9F822" bold="true" box="[199,389,1897,1922]" pageId="12" pageNumber="513">
|
||
ZooBank
|
||
<collectionCode id="ED20E9F7720DD512FEC36B534323F822" box="[316,383,1897,1922]" pageId="12" pageNumber="513">LSID</collectionCode>
|
||
.
|
||
</emphasis>
|
||
<uri id="FFA07D30720DD512FE736B53468FF822" box="[396,1235,1897,1922]" pageId="12" pageNumber="513">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FF974BE7720DD512FD5C6B53468FF822" box="[675,1235,1897,1922]" pageId="12" pageNumber="513">AC48DF0B-22B0-4ED3-AB19-1B1EBAFB1799</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B22B9720DD511FF386BB740FEFA25" lastPageId="15" lastPageNumber="516" pageId="12" pageNumber="513" type="description">
|
||
<paragraph id="8B8E7132720DD512FF386BB747DDF84E" blockId="12.[151,1436,1645,2030]" pageId="12" pageNumber="513">
|
||
<emphasis id="B945AD20720DD512FF386BB74304F806" bold="true" box="[199,344,1933,1958]" inLineHeading="true" pageId="12" pageNumber="513" reason="1">Description.</emphasis>
|
||
<emphasis id="B945AD20720DD512FE9F6BB44054F806" box="[352,520,1933,1958]" italics="true" pageId="12" pageNumber="513">Size and shape</emphasis>
|
||
—Corona very small, up to 3.5 mm in TL in material originally attributed to this species; outline distinctly oval (antero-posteriorly elongated) in aboral view; slightly flattened in profile, maximum height decreasing with size, being up to 70% of TL in smallest and approximately 56% TL in largest specimens.
|
||
</paragraph>
|
||
<caption id="DF4E21BA720CD513FF686FD646DCFB80" ID-DOI="http://doi.org/10.5281/zenodo.4930072" ID-Zenodo-Dep="4930072" httpUri="https://zenodo.org/record/4930072/files/figure.png" pageId="13" pageNumber="514" startId="13.[151,250,1004,1026]" targetBox="[228,1360,193,982]" targetPageId="13">
|
||
<paragraph id="8B8E7132720CD513FF686FD646DCFB80" blockId="13.[151,1436,1004,1056]" pageId="13" pageNumber="514">
|
||
<emphasis id="B945AD20720CD513FF686FD6434EFBA1" bold="true" box="[151,274,1004,1026]" pageId="13" pageNumber="514">FIGURE 7.</emphasis>
|
||
<bibRefCitation id="EFA00CC3720CD513FEE66FD6404DFBA2" author="Brunnschweiler, R. O." box="[281,529,1004,1026]" pageId="13" pageNumber="514" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler's (1962)</bibRefCitation>
|
||
holotypes: A–C.
|
||
<taxonomicName id="4C310AB1720CD513FD3B6FD641C9FBA2" authorityName="Brunnschweiler" authorityYear="1962" box="[708,917,1004,1026]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="514" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD20720CD513FD3B6FD641C9FBA2" box="[708,917,1004,1026]" italics="true" pageId="13" pageNumber="514">Cyamidia paucipora</emphasis>
|
||
</taxonomicName>
|
||
(= juvenile
|
||
<taxonomicName id="4C310AB1720CD513FBEE6FD646B0FBA2" authorityName="Brunnschweiler" authorityYear="1962" box="[1041,1260,1004,1026]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="514" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720CD513FBEE6FD646B0FBA2" box="[1041,1260,1004,1026]" italics="true" pageId="13" pageNumber="514">Lenicyamidia compta</emphasis>
|
||
</taxonomicName>
|
||
), GA CPC 2825; D–F.
|
||
<taxonomicName id="4C310AB1720CD513FF3168304011FB80" authority=", GA CPC 2827" authorityName=", GA CPC" authorityYear="2827" box="[206,589,1034,1056]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="514" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720CD513FF31683043F5FB80" box="[206,425,1034,1056]" italics="true" pageId="13" pageNumber="514">Lenicyamidia compta</emphasis>
|
||
, GA CPC 2827
|
||
</taxonomicName>
|
||
. Aboral (A, D), oral (B, E) and left lateral views (C, F).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E21BA720CD513FF686B7243EAF83B" ID-DOI="http://doi.org/10.5281/zenodo.4930074" ID-Zenodo-Dep="4930074" httpUri="https://zenodo.org/record/4930074/files/figure.png" pageId="13" pageNumber="514" startId="13.[151,250,1864,1886]" targetBox="[261,1326,1114,1836]" targetPageId="13">
|
||
<paragraph id="8B8E7132720CD513FF686B7243EAF83B" blockId="13.[151,1436,1864,1947]" pageId="13" pageNumber="514">
|
||
<emphasis id="B945AD20720CD513FF686B72434EF8FD" bold="true" box="[151,274,1864,1886]" pageId="13" pageNumber="514">FIGURE 8.</emphasis>
|
||
SEM images of the test of three specimens of
|
||
<taxonomicName id="4C310AB1720CD513FD176B7241E5F8FE" authorityName="Brunnschweiler" authorityYear="1962" box="[744,953,1864,1886]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="514" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD20720CD513FD176B7241E5F8FE" box="[744,953,1864,1886]" italics="true" pageId="13" pageNumber="514">Cyamidia paucipora</emphasis>
|
||
</taxonomicName>
|
||
(= juvenile
|
||
<taxonomicName id="4C310AB1720CD513FBCB6B724752F8FE" authorityName=", GA CPC" authorityYear="2827" box="[1076,1294,1864,1886]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="13" pageNumber="514" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720CD513FBCB6B724752F8FE" box="[1076,1294,1864,1886]" italics="true" pageId="13" pageNumber="514">Lenicyamidia compta</emphasis>
|
||
</taxonomicName>
|
||
). A. GA CPC 41767, aboral view. B–D: GA CPC 41768, aboral (B), oral (C) and left lateral view (D); E–G: GA CPC 41769, aboral (E), oral (F) and left lateral view (G).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E21BA720FD510FF686F31463DFC3C" ID-DOI="http://doi.org/10.5281/zenodo.4930076" ID-Zenodo-Dep="4930076" httpUri="https://zenodo.org/record/4930076/files/figure.png" pageId="14" pageNumber="515" startId="14.[151,250,779,801]" targetBox="[151,1436,194,758]" targetPageId="14">
|
||
<paragraph id="8B8E7132720FD510FF686F31463DFC3C" blockId="14.[151,1436,779,924]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF686F31434FFC80" bold="true" box="[151,275,779,801]" pageId="14" pageNumber="515">FIGURE 9.</emphasis>
|
||
SEM images showing details on test surface of
|
||
<taxonomicName id="4C310AB1720FD510FD036F314191FC81" authorityName="Brunnschweiler" authorityYear="1962" box="[764,973,779,801]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="14" pageNumber="515" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD20720FD510FD036F314191FC81" box="[764,973,779,801]" italics="true" pageId="14" pageNumber="515">Cyamidia paucipora</emphasis>
|
||
</taxonomicName>
|
||
(= juvenile
|
||
<taxonomicName id="4C310AB1720FD510FBB66F314778FC81" authorityName=", GA CPC" authorityYear="2827" box="[1097,1316,779,801]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="14" pageNumber="515" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720FD510FBB66F314778FC81" box="[1097,1316,779,801]" italics="true" pageId="14" pageNumber="515">Lenicyamidia compta</emphasis>
|
||
</taxonomicName>
|
||
). A. Apical region of specimen GA CPC 41767, pores of each respiratory tube foot linked by dotted lines, other apertures labelled: H = hydropore; G1, G2, G3, G4 = gonopores of interambulacra 1, 2, 3, and 4, respectively; ocular pores indicated by arrows. B. Ornamentation of test just anterior of the peristome in specimen GA CPC 41768. C. Peristomial region of specimen GA CPC 41769 showing peristomial region and periproct (bottom of image). All scale bars 200 µm long.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E7132720FD510FF386FF2404AFC41" blockId="14.[151,1437,968,2038]" box="[199,534,968,993]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF386FF243FFFC41" box="[199,419,968,993]" italics="true" pageId="14" pageNumber="515">Internal buttressing</emphasis>
|
||
—Absent.
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720FD510FF386FD64094FBD1" blockId="14.[151,1437,968,2038]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF386FD64315FBA5" box="[199,329,1004,1029]" italics="true" pageId="14" pageNumber="515">Apical disc</emphasis>
|
||
—Situated centrally, at apex of corona; monobasal, with four gonopores and single hydropore which lies close to genital pore 2 and not situated in pit or groove; ocular pores small and located outside area enclosed by gonopores (
|
||
<figureCitation id="130A6DB7720FD510FE55680E4059FBED" box="[426,517,1076,1101]" captionStart="FIGURE 9" captionStartId="14.[151,250,779,801]" captionTargetBox="[151,1436,194,758]" captionTargetId="figure-525@14.[151,1436,193,758]" captionTargetPageId="14" captionText="FIGURE 9. SEM images showing details on test surface of Cyamidia paucipora (= juvenile Lenicyamidia compta). A. Apical region of specimen GA CPC 41767, pores of each respiratory tube foot linked by dotted lines, other apertures labelled: H = hydropore; G1, G2, G3, G4 = gonopores of interambulacra 1, 2, 3, and 4, respectively; ocular pores indicated by arrows. B. Ornamentation of test just anterior of the peristome in specimen GA CPC 41768. C. Peristomial region of specimen GA CPC 41769 showing peristomial region and periproct (bottom of image). All scale bars 200 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930076" httpUri="https://zenodo.org/record/4930076/files/figure.png" pageId="14" pageNumber="515">Fig. 9A</figureCitation>
|
||
); gonopores open in specimens as small as 2.55 mm, but may still be completely closed in slightly larger specimens (2.69 mm TL).
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720FD510FF386846471EFA15" blockId="14.[151,1437,968,2038]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF386846431FFB35" box="[199,323,1148,1173]" italics="true" pageId="14" pageNumber="515">Ambulacra</emphasis>
|
||
—Plating not easily discernable, but ambulacra appear to expand towards ambitus; petals short, with up to five non-conjugate respiratory pore pairs in each ambulacrum; pore pairs strongly oblique, with obliqueness increasing distally; in small specimens with well-preserved surface detail, interporal areas raised, with glossy surface reminiscent of glassy tubercles, in slightly larger specimens this area is less raised but still appears to be composed of massive, unperforated stereom; interporiferous zones narrow and of constant width along petal; petaloid region large, extending about 60% of TL; food grooves absent; buccal pores not visible in oral view, apparently lying close to peristome and facing horizontally into it; accessory pores crowded in small oblique clusters in centre of each plate; widespread isolated accessory pores present between these clusters, including in interporiferous zones; ambitus initiating at approximately fourth pair of ambulacral postbasicoronal plates.
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720FD510FF3869FA40C7F981" blockId="14.[151,1437,968,2038]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF3869FA432BFA79" box="[199,375,1472,1497]" italics="true" pageId="14" pageNumber="515">Interambulacra</emphasis>
|
||
—Adapically, two unpaired plates lie in tandem adjacent to apical system (
|
||
<figureCitation id="130A6DB7720FD510FAE169FA47D3FA79" box="[1310,1423,1472,1497]" captionStart="FIGURE 10" captionStartId="15.[151,250,1188,1210]" captionTargetBox="[160,1428,189,1167]" captionTargetId="figure-378@15.[151,1436,181,1173]" captionTargetPageId="15" captionText="FIGURE 10. Partial plate architecture of Lenicyamidia compta and Cyamidia paucipora (= juvenile Lenicyamidia compta). A–C. L. compta: A. aboral view of GA CPC 41774; B. oral view of the holotype GA CPC 2827; C. left lateral view of GA CPC 41774; D. C. paucipora, holotype CPC 2825, adapical part of interambulacrum 2. Labelling conventions as in Fig. 6." figureDoi="http://doi.org/10.5281/zenodo.4930078" httpUri="https://zenodo.org/record/4930078/files/figure.png" pageId="14" pageNumber="515">Fig. 10D</figureCitation>
|
||
); basicoronal plates stout and quadrangular; ambital plating not discernable; a few scattered accessory pores present both in aboral and oral interambulacral plates.
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720FD510FF386A164717F911" blockId="14.[151,1437,968,2038]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF386A16433FF9E5" box="[199,355,1580,1605]" italics="true" pageId="14" pageNumber="515">Tuberculation</emphasis>
|
||
—Primary tubercles crenulate, perforate; homogeneously distributed on aboral surface; on oral surface, towards the peristome, tubercles up to 1.5 times larger and more widely scattered, separated by ridges bearing glassy tubercles and granules (
|
||
<figureCitation id="130A6DB7720FD510FDB96A4E40C1F92D" box="[582,669,1652,1677]" captionStart="FIGURE 9" captionStartId="14.[151,250,779,801]" captionTargetBox="[151,1436,194,758]" captionTargetId="figure-525@14.[151,1436,193,758]" captionTargetPageId="14" captionText="FIGURE 9. SEM images showing details on test surface of Cyamidia paucipora (= juvenile Lenicyamidia compta). A. Apical region of specimen GA CPC 41767, pores of each respiratory tube foot linked by dotted lines, other apertures labelled: H = hydropore; G1, G2, G3, G4 = gonopores of interambulacra 1, 2, 3, and 4, respectively; ocular pores indicated by arrows. B. Ornamentation of test just anterior of the peristome in specimen GA CPC 41768. C. Peristomial region of specimen GA CPC 41769 showing peristomial region and periproct (bottom of image). All scale bars 200 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930076" httpUri="https://zenodo.org/record/4930076/files/figure.png" pageId="14" pageNumber="515">Fig. 9B</figureCitation>
|
||
); tubercles entirely missing on basicoronal plate of interambulacrum 5 and most of plate surface of plates 5.a.2 and 5.b.2; here large glassy tubercles are found instead (
|
||
<figureCitation id="130A6DB7720FD510FB196AA24762F911" box="[1254,1342,1688,1713]" captionStart="FIGURE 9" captionStartId="14.[151,250,779,801]" captionTargetBox="[151,1436,194,758]" captionTargetId="figure-525@14.[151,1436,193,758]" captionTargetPageId="14" captionText="FIGURE 9. SEM images showing details on test surface of Cyamidia paucipora (= juvenile Lenicyamidia compta). A. Apical region of specimen GA CPC 41767, pores of each respiratory tube foot linked by dotted lines, other apertures labelled: H = hydropore; G1, G2, G3, G4 = gonopores of interambulacra 1, 2, 3, and 4, respectively; ocular pores indicated by arrows. B. Ornamentation of test just anterior of the peristome in specimen GA CPC 41768. C. Peristomial region of specimen GA CPC 41769 showing peristomial region and periproct (bottom of image). All scale bars 200 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930076" httpUri="https://zenodo.org/record/4930076/files/figure.png" pageId="14" pageNumber="515">Fig. 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720FD510FF386A874352F8E1" blockId="14.[151,1437,968,2038]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF386A874365F974" box="[199,313,1725,1748]" italics="true" pageId="14" pageNumber="515">Peristome</emphasis>
|
||
—Relatively large, about 25% TL; facing directly downwards; infundibulum shallow, but with down-turned, almost vertical lip; incomplete ring of slightly elevated ambulacral stereom bars framing peristome; distinct perradial bulge containing the sphaeridium in centre of each bar; in oral view, buccal pores are hidden by these bars.
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720FD510FF386B7741FFF80D" blockId="14.[151,1437,968,2038]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF386B77436DF8C4" box="[199,305,1869,1892]" italics="true" pageId="14" pageNumber="515">Periproct</emphasis>
|
||
—Smaller than peristome, approximately 12% TL; positioned inframarginally and facing obliquely down-/backwards in oral view; distinctly elongated along anterior-posterior axis; anterior end bounded by first pair of post-basicoronal plates (5.a.2, 5.b.2), other sutures not discernable.
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720FD510FF386B824289F856" blockId="14.[151,1437,968,2038]" pageId="14" pageNumber="515">
|
||
<emphasis id="B945AD20720FD510FF386B8243C4F871" box="[199,408,1976,2001]" italics="true" pageId="14" pageNumber="515">Perignathic girdle</emphasis>
|
||
—Consisting of five small processes (auricles), one on each interambulacral basicoronal plate.
|
||
</paragraph>
|
||
<caption id="DF4E21BA720ED511FF68689E4764FB57" ID-DOI="http://doi.org/10.5281/zenodo.4930078" ID-Zenodo-Dep="4930078" httpUri="https://zenodo.org/record/4930078/files/figure.png" pageId="15" pageNumber="516" startId="15.[151,250,1188,1210]" targetBox="[160,1428,189,1167]" targetPageId="15">
|
||
<paragraph id="8B8E7132720ED511FF68689E4764FB57" blockId="15.[151,1436,1188,1271]" pageId="15" pageNumber="516">
|
||
<emphasis id="B945AD20720ED511FF68689E437DFB19" bold="true" box="[151,289,1188,1210]" pageId="15" pageNumber="516">FIGURE 10.</emphasis>
|
||
Partial plate architecture of
|
||
<taxonomicName id="4C310AB1720ED511FDB4689E4174FB1A" authorityName=", GA CPC" authorityYear="2827" box="[587,808,1188,1210]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720ED511FDB4689E4174FB1A" box="[587,808,1188,1210]" italics="true" pageId="15" pageNumber="516">Lenicyamidia compta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C310AB1720ED511FCA3689E4673FB1A" authorityName="Brunnschweiler" authorityYear="1962" box="[860,1071,1188,1210]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD20720ED511FCA3689E4673FB1A" box="[860,1071,1188,1210]" italics="true" pageId="15" pageNumber="516">Cyamidia paucipora</emphasis>
|
||
</taxonomicName>
|
||
(= juvenile
|
||
<taxonomicName id="4C310AB1720ED511FB4F689E47D1FB1A" authorityName=", GA CPC" authorityYear="2827" box="[1200,1421,1188,1210]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720ED511FB4F689E47D1FB1A" box="[1200,1421,1188,1210]" italics="true" pageId="15" pageNumber="516">Lenicyamidia compta</emphasis>
|
||
</taxonomicName>
|
||
). A–C.
|
||
<taxonomicName id="4C310AB1720ED511FF2C68FE4364FB79" authorityName=", GA CPC" authorityYear="2827" box="[211,312,1220,1241]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720ED511FF2C68FE4364FB79" box="[211,312,1220,1241]" italics="true" pageId="15" pageNumber="516">L. compta</emphasis>
|
||
</taxonomicName>
|
||
: A. aboral view of GA CPC 41774; B. oral view of the holotype GA CPC 2827; C. left lateral view of GA CPC 41774; D.
|
||
<taxonomicName id="4C310AB1720ED511FEFD68DB43D4FB57" authorityName="Brunnschweiler" authorityYear="1962" box="[258,392,1249,1271]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD20720ED511FEFD68DB43D4FB57" box="[258,392,1249,1271]" italics="true" pageId="15" pageNumber="516">C. paucipora</emphasis>
|
||
</taxonomicName>
|
||
, holotype CPC 2825, adapical part of interambulacrum 2. Labelling conventions as in Fig. 6.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E7132720ED511FF38691E4287FAC0" blockId="15.[151,1436,1316,1989]" pageId="15" pageNumber="516">
|
||
<taxonomicName id="4C310AB1720ED511FF38691E431EFA9D" box="[199,322,1316,1341]" class="Entognatha" family="Sminthurididae" genus="Sphaeridia" kingdom="Animalia" order="Collembola" pageId="15" pageNumber="516" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945AD20720ED511FF38691E431EFA9D" box="[199,322,1316,1341]" italics="true" pageId="15" pageNumber="516">Sphaeridia</emphasis>
|
||
</taxonomicName>
|
||
—One per ambulacrum; fully enclosed; situated beneath distinct transverse bar just distal to buccal pores.
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720ED511FF38695740FEFA25" blockId="15.[151,1436,1316,1989]" box="[199,674,1388,1413]" pageId="15" pageNumber="516">
|
||
<emphasis id="B945AD20720ED511FF386957404DFA25" box="[199,529,1388,1413]" italics="true" pageId="15" pageNumber="516">Spines, pedicellariae, lantern</emphasis>
|
||
—Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B22B9720ED50EFF3869AA43A4FACD" lastPageId="16" lastPageNumber="517" pageId="15" pageNumber="516" type="discussion">
|
||
<paragraph id="8B8E7132720ED511FF3869AA4689F905" blockId="15.[151,1436,1316,1989]" pageId="15" pageNumber="516">
|
||
<emphasis id="B945AD20720ED511FF3869AA4367FA09" bold="true" box="[199,315,1424,1449]" inLineHeading="true" pageId="15" pageNumber="516" reason="1">Remarks.</emphasis>
|
||
<bibRefCitation id="EFA00CC3720ED511FEBD69AA401AFA09" author="Brunnschweiler, R. O." box="[322,582,1424,1449]" pageId="15" pageNumber="516" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler (1962)</bibRefCitation>
|
||
established a new species for the specimens described above and attributed it to the genus
|
||
<taxonomicName id="4C310AB1720ED511FEC2698E43F6FA6D" authorityName="Lambert & Thiery" authorityYear="1914" box="[317,426,1460,1485]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD20720ED511FEC2698E43F6FA6D" box="[317,426,1460,1485]" italics="true" pageId="15" pageNumber="516">Cyamidia</emphasis>
|
||
</taxonomicName>
|
||
on the basis of its elongate periproct and lack of internal buttressing. It was differentiated from previously described species of
|
||
<taxonomicName id="4C310AB1720ED511FDBC69E240ECFA51" authorityName="Lambert & Thiery" authorityYear="1914" box="[579,688,1496,1521]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD20720ED511FDBC69E240ECFA51" box="[579,688,1496,1521]" italics="true" pageId="15" pageNumber="516">Cyamidia</emphasis>
|
||
</taxonomicName>
|
||
by its larger and more marginal periproct, the smaller number of petaloid pores and the glassy tubercles between the periproct and peristome. Brunnschweiler, however, did not take allometric growth into account. In most clypeasteroids, and indeed most echinoids, the periproct and peristome are proportionally larger in smaller specimens and the petaloid pore pair number is lower. The main difference from other
|
||
<taxonomicName id="4C310AB1720ED511FF266A5243C3F921" box="[217,415,1640,1665]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="undetermined">
|
||
<emphasis id="B945AD20720ED511FF266A52431AF921" box="[217,326,1640,1665]" italics="true" pageId="15" pageNumber="516">Cyamidia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
is therefore the presence of glassy tubercles between the periproct and peristome and the more marginal periproct (although periproct position too tends to change during early ontogeny).
|
||
</paragraph>
|
||
<paragraph id="8B8E7132720ED50EFF386A8A4741FB35" blockId="15.[151,1436,1316,1989]" lastBlockId="16.[151,1436,1040,1389]" lastPageId="16" lastPageNumber="517" pageId="15" pageNumber="516">
|
||
<taxonomicName id="4C310AB1720ED511FF386A8A43F2F968" authorityName="Brunnschweiler" authorityYear="1962" box="[199,430,1712,1737]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD20720ED511FF386A8A43F2F968" box="[199,430,1712,1737]" italics="true" pageId="15" pageNumber="516">Cyamidia paucipora</emphasis>
|
||
</taxonomicName>
|
||
co-occurs with the larger species
|
||
<taxonomicName id="4C310AB1720ED511FCD66A8A4647F968" authorityName=", GA CPC" authorityYear="2827" box="[809,1051,1712,1737]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720ED511FCD66A8A4647F968" box="[809,1051,1712,1737]" italics="true" pageId="15" pageNumber="516">Lenicyamidia compta</emphasis>
|
||
</taxonomicName>
|
||
at the
|
||
<typeStatus id="548ACF90720ED511FB9A6A8B46C9F969" box="[1125,1173,1713,1737]" pageId="15" pageNumber="516">type</typeStatus>
|
||
locality. At first glance the two species appear very different—the former is very small, high, and ovoid in outline, while the latter is larger, more flattened and rounded. It also has a distinct oral tuberculation pattern. Upon closer examination, however, the two forms appear to be part of a single ontogenetic series, with the main differences being related to differential growth. Test height, for example, increases only slowly during growth (
|
||
<figureCitation id="130A6DB7720ED511FC406B7A464DF8F9" box="[959,1041,1856,1881]" captionStart="FIGURE" captionStartId="16.[151,168,836,861]" captionTargetBox="[345,1243,152,809]" captionTargetId="figure-380@16.[341,1249,147,812]" captionTargetPageId="16" captionText="FIGURE 11. Scatter plot of test width (solid squares; n = 30), height (open triangles; n = 27) and respiratory pore number (solid circles; n = 17) vs. test length for the material attributed to Lenicyamidia compta and Cyamidia paucipora by Brunnschweiler (1962). Size range of C. paucipora highlighted by grey backfill. Thumbnail images show outline and profile for the smallest and largest complete specimens. Within profiles, scale bars 1.0 mm long, arrows indicate anterior end. APP = anterior paired petals (i.e. petal II or IV)." figureDoi="http://doi.org/10.5281/zenodo.4930082" httpUri="https://zenodo.org/record/4930082/files/figure.png" pageId="15" pageNumber="516">Fig. 11</figureCitation>
|
||
), causing the end members to look very different. When included in the same graph, specimens attributed to
|
||
<taxonomicName id="4C310AB1720ED511FC276B5F4630F8DC" authorityName="Brunnschweiler" authorityYear="1962" box="[984,1132,1893,1916]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD20720ED511FC276B5F4630F8DC" box="[984,1132,1893,1916]" italics="true" pageId="15" pageNumber="516">C. paucipora</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFA00CC3720ED511FB676B5E47C0F8DD" author="Brunnschweiler, R. O." box="[1176,1436,1892,1917]" pageId="15" pageNumber="516" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler (1962)</bibRefCitation>
|
||
fall along the same regression lines as
|
||
<taxonomicName id="4C310AB1720ED511FDB96BB340E9F800" authorityName=", GA CPC" authorityYear="2827" box="[582,693,1929,1952]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720ED511FDB96BB340E9F800" box="[582,693,1929,1952]" italics="true" pageId="15" pageNumber="516">L. compta</emphasis>
|
||
</taxonomicName>
|
||
. The size gap between
|
||
<taxonomicName id="4C310AB1720ED511FC436BB34613F800" authorityName="Brunnschweiler" authorityYear="1962" box="[956,1103,1929,1952]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD20720ED511FC436BB34613F800" box="[956,1103,1929,1952]" italics="true" pageId="15" pageNumber="516">C. paucipora</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C310AB1720ED511FB796BB346AAF800" authorityName=", GA CPC" authorityYear="2827" box="[1158,1270,1929,1952]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="15" pageNumber="516" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD20720ED511FB796BB346AAF800" box="[1158,1270,1929,1952]" italics="true" pageId="15" pageNumber="516">L. compta</emphasis>
|
||
</taxonomicName>
|
||
apparently led Brunnschweiler to believe that he was dealing with two different taxa. This discontinuity in the size distribution might have other reasons including, but not limited to: a) collection bias (the small specimens probably being picked from sieving residues, while the larger ones might have been hand-picked in the field); b) year-cohorts (many marine invertebrates reproduce only once annually, leading to different size cohorts within populations particularly among miniaturized, paedomorphic forms in which these size differences are exaggerated).
|
||
</paragraph>
|
||
<caption id="DF4E21BA7211D50EFF686F7E403CFC48" ID-DOI="http://doi.org/10.5281/zenodo.4930082" ID-Zenodo-Dep="4930082" httpUri="https://zenodo.org/record/4930082/files/figure.png" pageId="16" pageNumber="517" startId="16.[151,168,836,861]" targetBox="[345,1243,152,809]" targetPageId="16">
|
||
<paragraph id="8B8E71327211D50EFF686F7E403CFC48" blockId="16.[151,1436,836,1000]" pageId="16" pageNumber="517">
|
||
<emphasis id="B945AD207211D50EFF686F7E437DFCFB" bold="true" box="[151,289,836,861]" pageId="16" pageNumber="517">FIGURE 11.</emphasis>
|
||
Scatter plot of test width (solid squares; n = 30), height (open triangles; n = 27) and respiratory pore number (solid circles; n = 17) vs. test length for the material attributed to
|
||
<taxonomicName id="4C310AB17211D50EFC806F52463DFCDE" authorityName=", GA CPC" authorityYear="2827" box="[895,1121,872,894]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="16" pageNumber="517" phylum="Echinodermata" rank="species" species="compta">
|
||
<emphasis id="B945AD207211D50EFC806F52463DFCDE" box="[895,1121,872,894]" italics="true" pageId="16" pageNumber="517">Lenicyamidia compta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C310AB17211D50EFB616F524729FCDE" authorityName="Brunnschweiler" authorityYear="1962" box="[1182,1397,872,894]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="16" pageNumber="517" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD207211D50EFB616F524729FCDE" box="[1182,1397,872,894]" italics="true" pageId="16" pageNumber="517">Cyamidia paucipora</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFA00CC37211D50EFF686FB043DAFC00" author="Brunnschweiler, R. O." box="[151,390,906,928]" pageId="16" pageNumber="517" pagination="159 - 169" refId="ref11830" refString="Brunnschweiler, R. O. (1962) On echinoids in the Tertiary of Western Australia with a description of two new Eocene Fibulariidae. Journal of the Geological Society of Australia, 8, 159 - 169. http: // dx. doi. org / 10.1080 / 14400956208527871" type="journal article" year="1962">Brunnschweiler (1962)</bibRefCitation>
|
||
. Size range of
|
||
<taxonomicName id="4C310AB17211D50EFDDF6FB040FBFC00" authorityName="Brunnschweiler" authorityYear="1962" box="[544,679,906,928]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="16" pageNumber="517" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD207211D50EFDDF6FB040FBFC00" box="[544,679,906,928]" italics="true" pageId="16" pageNumber="517">C. paucipora</emphasis>
|
||
</taxonomicName>
|
||
highlighted by grey backfill. Thumbnail images show outline and profile for the smallest and largest complete specimens. Within profiles, scale bars 1.0 mm long, arrows indicate anterior end. APP = anterior paired petals (i.e. petal II or IV).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E71327211D50EFF38689A43A4FACD" blockId="16.[151,1436,1040,1389]" pageId="16" pageNumber="517">
|
||
The only other difference between
|
||
<taxonomicName id="4C310AB17211D50EFDAD689B40BAFB18" authorityName="Brunnschweiler" authorityYear="1962" box="[594,742,1185,1208]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="16" pageNumber="517" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD207211D50EFDAD689B40BAFB18" box="[594,742,1185,1208]" italics="true" pageId="16" pageNumber="517">C. paucipora</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C310AB17211D50EFCE1689A41EAFB19" authorityName="Brunnschweiler" authorityYear="1962" box="[798,950,1184,1209]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="16" pageNumber="517" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207211D50EFCE1689A41EAFB19" box="[798,950,1184,1209]" italics="true" pageId="16" pageNumber="517">Lenicyamidia</emphasis>
|
||
</taxonomicName>
|
||
, apart from shape and pore pair number, is that in tuberculation pattern. However, the oral, medial area of glassy tubercles in
|
||
<taxonomicName id="4C310AB17211D50EFB9F68FE46A5FB7D" authorityName="Brunnschweiler" authorityYear="1962" box="[1120,1273,1220,1245]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="16" pageNumber="517" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B945AD207211D50EFB9F68FE46A5FB7D" box="[1120,1273,1220,1245]" italics="true" pageId="16" pageNumber="517">Lenicyamidia</emphasis>
|
||
</taxonomicName>
|
||
can easily be derived from the corresponding area in
|
||
<taxonomicName id="4C310AB17211D50EFDA768D2411CFAA0" authorityName="Brunnschweiler" authorityYear="1962" box="[600,832,1256,1281]" class="Echinoidea" family="Fibulariidae" genus="Cyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="16" pageNumber="517" phylum="Echinodermata" rank="species" species="paucipora">
|
||
<emphasis id="B945AD207211D50EFDA768D2411CFAA0" box="[600,832,1256,1281]" italics="true" pageId="16" pageNumber="517">Cyamidia paucipora</emphasis>
|
||
</taxonomicName>
|
||
. The latter also largely lacks spine-bearing tubercles, and features glassy ones in the oral, medial area instead (
|
||
<figureCitation id="130A6DB77211D50EFCED69364137FA85" box="[786,875,1292,1317]" captionStart="FIGURE 9" captionStartId="14.[151,250,779,801]" captionTargetBox="[151,1436,194,758]" captionTargetId="figure-525@14.[151,1436,193,758]" captionTargetPageId="14" captionText="FIGURE 9. SEM images showing details on test surface of Cyamidia paucipora (= juvenile Lenicyamidia compta). A. Apical region of specimen GA CPC 41767, pores of each respiratory tube foot linked by dotted lines, other apertures labelled: H = hydropore; G1, G2, G3, G4 = gonopores of interambulacra 1, 2, 3, and 4, respectively; ocular pores indicated by arrows. B. Ornamentation of test just anterior of the peristome in specimen GA CPC 41768. C. Peristomial region of specimen GA CPC 41769 showing peristomial region and periproct (bottom of image). All scale bars 200 µm long." figureDoi="http://doi.org/10.5281/zenodo.4930076" httpUri="https://zenodo.org/record/4930076/files/figure.png" pageId="16" pageNumber="517">Fig. 9C</figureCitation>
|
||
). Based on these observations, the two species are here considered synonymous and
|
||
<emphasis id="B945AD207211D50EFDDD690A414CFAE8" box="[546,784,1328,1353]" italics="true" pageId="16" pageNumber="517">
|
||
Lenicyamida
|
||
<taxonomicName id="4C310AB17211D50EFD42690B414CFAE8" authorityName=", GA CPC" authorityYear="2827" box="[701,784,1329,1352]" class="Echinoidea" family="Fibulariidae" genus="Lenicyamidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="16" pageNumber="517" phylum="Echinodermata" rank="species" species="compta">compta</taxonomicName>
|
||
</emphasis>
|
||
is chosen as the senior synonym (Principle of the First Reviser; ICZN 1999 Art. 24.2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |