270 lines
37 KiB
XML
270 lines
37 KiB
XML
<document id="EAF3CCF230F79C09AF47E21403D84F47" ID-ISSN="1175-5326" ID-ZooBank="2C674056-14B9-4948-9E6A-346D5ED8E4D5" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629832682804" checkinUser="felipe" docAuthor="Behounek, G. & V. S. Kononenko" docDate="2012" docId="4E00879C562EFA42FF5EFC99FC623D73" docLanguage="en" docName="zt03590p015.pdf" docOrigin="Zootaxa 3590" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anabelcia Behounek & V.S.Kononenko 2012, gen. n." docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="B239FFE4562FFA40FFC9FFE7FFAA3878" masterDocTitle="A revision of the “ Belciana ” kala Prout, 1924 species group with description of a new genus and two new species (Lepidoptera, Noctuidae: Pantheinae) from East Asia. Revision of Pantheinae, contribution VI" masterLastPageNumber="15" masterPageNumber="1" pageNumber="2" updateTime="1699118649507" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3AA477CDBEFD6B0F70540AA571B356C3">A revision of the “ Belciana ” kala Prout, 1924 species group with description of a new genus and two new species (Lepidoptera, Noctuidae: Pantheinae) from East Asia. Revision of Pantheinae, contribution VI</mods:title>
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<mods:name id="42F631E8502F03E128C46AA4B3774B33" type="personal">
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<mods:namePart id="271AB3DB93DF46E96FB90B8849099C69">Behounek, G.</mods:namePart>
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<mods:affiliation id="8254978ECAB5045304D86AF6E251139D">D- 85567 Grafing / Munich, Sudetenstrasse 6, Germany. E-mail: Gottfried. Behounek @ t-online. de</mods:affiliation>
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<mods:namePart id="625864DC20E33B5791FAC606F4E77F66">V. S. Kononenko</mods:namePart>
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<mods:affiliation id="8254C711A8BECA2CCD2629E6D561952F">Laboratory of Entomology, Institute of Biology and Soil Science Far Eastern Branch of Russian Academy of Sciences, RF- 690022 Vladivostok, Russia. E-mail: kononenko @ ibss. dvo. ru</mods:affiliation>
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<mods:title id="410AF7C1D45459080203809FB5151490">Zootaxa</mods:title>
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<mods:date id="B330AE38216ACAEB546C5F4B31B954B8">2012</mods:date>
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<mods:number id="3AF30F16ED81B2708130DBAB65F2765F">2012-12-18</mods:number>
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<mods:number id="605E6FEFB1DBA46B24DA284B233FF73B">3590</mods:number>
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<treatment id="4E00879C562EFA42FF5EFC99FC623D73" ID-DOI="http://doi.org/10.5281/zenodo.5259268" ID-Zenodo-Dep="5259268" LSID="urn:lsid:plazi:treatment:4E00879C562EFA42FF5EFC99FC623D73" httpUri="http://treatment.plazi.org/id/4E00879C562EFA42FF5EFC99FC623D73" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<subSubSection id="8EB36501562EFA41FF5EFC99FEC13BE0" box="[151,363,894,920]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="C616368A562EFA41FF5EFC99FEC13BE0" blockId="1.[151,363,894,920]" box="[151,363,894,920]" pageId="1" pageNumber="2">
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<heading id="9D5E81E6562EFA41FF5EFC99FEC13BE0" bold="true" box="[151,363,894,920]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="F4DDEA98562EFA41FF5EFC99FEC13BE0" bold="true" box="[151,363,894,920]" pageId="1" pageNumber="2">
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<taxonomicName id="01A94D09562EFA41FF5EFC99FEBB3BE0" authority="Behounek & V.S.Kononenko, 2012" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" box="[151,273,894,920]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="F4DDEA98562EFA41FF5EFC99FEBB3BE0" bold="true" box="[151,273,894,920]" italics="true" pageId="1" pageNumber="2">Anabelcia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EFEE57E3562EFA41FED1FC99FEC13BE0" box="[280,363,894,920]" pageId="1" pageNumber="2" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8EB36501562EFA41FF5EFC21FAD63BA3" box="[151,1404,965,987]" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="C616368A562EFA41FF5EFC21FAD63BA3" blockId="1.[151,1404,965,987]" box="[151,1404,965,987]" pageId="1" pageNumber="2">
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<materialsCitation id="76C13CD7562EFA41FF5EFC21FAD23BA3" box="[151,1400,965,987]" county="Bulletin" location="Witley" municipality="Hill Museum" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
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<typeStatus id="19128828562EFA41FF5EFC21FF613BA3" box="[151,203,966,987]" pageId="1" pageNumber="2">Type</typeStatus>
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species:
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<taxonomicName id="01A94D09562EFA41FEEFFC22FD9A3BA3" authority="Prout, 1924" authorityName="Prout" authorityYear="1924" box="[294,560,965,987]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="kala">
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<emphasis id="F4DDEA98562EFA41FEEFFC22FE183BA3" box="[294,434,965,987]" italics="true" pageId="1" pageNumber="2">Belciana kala</emphasis>
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<bibRefCitation id="A2384B7B562EFA41FE71FC21FD9A3BA3" author="Prout, A. E." box="[440,560,965,987]" pageId="1" pageNumber="2" pagination="400 - 412" refId="ref8424" refString="Prout, A. E. (1924) A preliminary revision of the genus Trisuloides Btlr. (Lep. Het. Noctuidae), with descriptions of new genera and new species and notes on the genitalia by G. Talbot F. E. S. Bulletin of the Hill Museum, Witley 1, 400 - 412, pls. 13, 16, 17." type="journal article" year="1924">Prout, 1924</bibRefCitation>
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</taxonomicName>
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||
,
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<emphasis id="F4DDEA98562EFA41FDF2FC22FC353BA3" box="[571,927,965,987]" italics="true" pageId="1" pageNumber="2">
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<collectingCounty id="2F774E06562EFA41FDF2FC22FD213BA3" box="[571,651,965,987]" pageId="1" pageNumber="2">Bulletin</collectingCounty>
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of the
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<collectingMunicipality id="2672ACF0562EFA41FD18FC22FCFD3BA3" box="[721,855,965,987]" pageId="1" pageNumber="2">Hill Museum</collectingMunicipality>
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,
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<location id="C3766051562EFA41FCA8FC22FC353BA3" LSID="urn:lsid:plazi:treatment:4E00879C562EFA42FF5EFC99FC623D73:C3766051562EFA41FCA8FC22FC353BA3" box="[865,927,965,987]" county="Bulletin" municipality="Hill Museum" name="Witley" pageId="1" pageNumber="2" stateProvince="Sumatera Utara">Witley</location>
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</emphasis>
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, 1: 442.
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<collectingRegion id="046DF868562EFA41FC31FC22FBE53BA3" box="[1016,1103,965,987]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="2">Sumatra</collectingRegion>
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,
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<location id="C3766051562EFA41FB90FC22FB453BA3" LSID="urn:lsid:plazi:treatment:4E00879C562EFA42FF5EFC99FC623D73:C3766051562EFA41FB90FC22FB453BA3" box="[1113,1263,965,987]" county="Bulletin" municipality="Hill Museum" name="Barisan Range" pageId="1" pageNumber="2" stateProvince="Sumatera Utara">Barisan Range</location>
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,
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<location id="C3766051562EFA41FB33FC21FAD23BA3" LSID="urn:lsid:plazi:treatment:4E00879C562EFA42FF5EFC99FC623D73:C3766051562EFA41FB33FC21FAD23BA3" box="[1274,1400,965,987]" county="Bulletin" municipality="Hill Museum" name="Mt. Korintji" pageId="1" pageNumber="2" stateProvince="Sumatera Utara">Mt. Korintji</location>
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="8EB36501562EFA42FF5EFBE0FDF739D4" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="diagnosis">
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<paragraph id="C616368A562EFA41FF5EFBE0FE923E88" blockId="1.[151,1436,1031,2028]" pageId="1" pageNumber="2">
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<emphasis id="F4DDEA98562EFA41FF5EFBE0FEBB3C58" bold="true" box="[151,273,1031,1056]" inLineHeading="true" pageId="1" pageNumber="2" reason="1">Diagnosis.</emphasis>
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Externally the new genus is similar to the genera
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<taxonomicName id="01A94D09562EFA41FC4BFBEFFB213C58" authority=", Diptheroides" authorityName="Diptheroides" box="[898,1163,1032,1056]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="F4DDEA98562EFA41FC4BFBEFFC4C3C58" box="[898,998,1032,1056]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
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,
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<emphasis id="F4DDEA98562EFA41FC33FBEFFB213C58" box="[1018,1163,1032,1056]" italics="true" pageId="1" pageNumber="2">Diptheroides</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="01A94D09562EFA41FB56FBEFFB413C58" authorityName="Moore" authorityYear="1882" box="[1183,1259,1032,1056]" class="Insecta" family="Noctuidae" genus="Donda" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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||
<emphasis id="F4DDEA98562EFA41FB56FBEFFB413C58" box="[1183,1259,1032,1056]" italics="true" pageId="1" pageNumber="2">Donda</emphasis>
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||
</taxonomicName>
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||
and
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<taxonomicName id="01A94D09562EFA41FAE6FBEFFE2F3C3C" authority="Kozhanchikov, 1950" authorityName="Kozhanchikov" authorityYear="1950" class="Insecta" family="Noctuidae" genus="Belciades" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="F4DDEA98562EFA41FAE6FBEFFA363C58" box="[1327,1436,1032,1056]" italics="true" pageId="1" pageNumber="2">Belciades</emphasis>
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Kozhanchikov, 1950
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</taxonomicName>
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, these closely related phyletic lineages form a common generic complex. Externally this group is characterized by bluish, bluish-green, green or green-brown ground colour of forewing and rather permanently stable main elements of wing pattern with blackish-brown subbasal, subterminal and tornal patches. The structures of the male genitalia show, however, clearly recognisable apomorphic features in the different genera. Compared with
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<taxonomicName id="01A94D09562EFA41FE7AFB5BFDBC3CAC" box="[435,534,1212,1236]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="F4DDEA98562EFA41FE7AFB5BFDBC3CAC" box="[435,534,1212,1236]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
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</taxonomicName>
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||
(
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||
<figureCitation id="5E922A0F562EFA41FDE2FB5BFDDD3CAC" box="[555,631,1212,1236]" captionStart="FIGURES 1–8" captionStartId="7.[151,263,1841,1864]" captionTargetBox="[151,1436,194,1821]" captionTargetId="figure-104@7.[151,1436,193,1821]" captionTargetPageId="7" captionText="FIGURES 1–8. Belciana and Anabelcia spp., adults. 1. B. biformis, female, Malaysia, Borneo (GB); 2. A. kala, male, Indonesia, Sumatra, genit. prep. 12018GB (ZMUC); 3. A. kala, male, Malaysia (Pahang) (AZ); 4. A. kala, female, Philippines, Palawan, genit. prep. 12063GB (ZFMK); 5. A. kala, male, Vietnam, genit. prep. 7348GB (GB); 6. A. nepalensis, male, Nepal, genit. prep. 4161GB (ZSM); 7. A. nepalensis, female, Nepal, genit. prep. 4162GB (ZSM); 8. A. nepalensis, female, N India, genit. prep. 7413GB (ZSM)." pageId="1" pageNumber="2">Figs 1</figureCitation>
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,
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<figureCitation id="5E922A0F562EFA41FD43FB5BFD0D3CAC" box="[650,679,1212,1236]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1841,1864]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-75@9.[151,1436,193,1821]" captionTargetPageId="9" captionText="FIGURES 17–22. Belciana and Anabelcia spp., structures of palps (17–20) and male genitalia (21, 22). The distinctive characters marked by lines. 17. B. bifornis; 18. A. kala; 19. A. nepalensis; 20. A. taiwana; 21. A. biformis, Indonesia, Sumatra, genit. prep. 4160GB (GB); 22. A. kala, Indonesia, Sumatra, genit. prep. 12018GB (GB)." pageId="1" pageNumber="2">17</figureCitation>
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) the new genus differs by more deeply green ground colour of forewing (bluish-green in
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<taxonomicName id="01A94D09562EFA41FE77FB07FD8B3C80" box="[446,545,1248,1272]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="F4DDEA98562EFA41FE77FB07FD8B3C80" box="[446,545,1248,1272]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
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</taxonomicName>
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) and more developed brown subbasal patch in costal area and more developed subterminal and tornal patches. Wing pattern formed with numerous black thin dentate lines (in most cases one blackish medial line in
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<taxonomicName id="01A94D09562EFA41FE52FACFFE543D38" box="[411,510,1320,1344]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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||
<emphasis id="F4DDEA98562EFA41FE52FACFFE543D38" box="[411,510,1320,1344]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
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||
</taxonomicName>
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||
). Hindwing usually bright, yellow or pale yellowish with prominent terminal band (fuscous in
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<taxonomicName id="01A94D09562EFA41FED4FAABFE2B3D1C" box="[285,385,1356,1380]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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||
<emphasis id="F4DDEA98562EFA41FED4FAABFE2B3D1C" box="[285,385,1356,1380]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
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||
</taxonomicName>
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||
, without clear terminal band).
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<bibRefCitation id="A2384B7B562EFA41FD2DFAABFC0C3D1C" author="Holloway, J. A." box="[740,934,1356,1380]" pageId="1" pageNumber="2" pagination="1 - 240" refId="ref7761" refString="Holloway, J. A. (2009) The Moths of Borneo, part 13, Family Noctuidae, subfamilies Pantheinae, Bagisarinae, Acontiinae, Aediinae, Eustrotiinae, Bryophilinae, Aeropteroninae, Aventiinae, Eublemminae. Malayan Nature Journal 62 (1 - 2), 1 - 240." type="journal article" year="2009">Holloway (2009)</bibRefCitation>
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||
noted that “
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||
<taxonomicName id="01A94D09562EFA41FBFFFAAAFB673D1B" authority="Prout" authorityName="Prout" authorityYear="1924" box="[1078,1229,1356,1380]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="kala">
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<emphasis id="F4DDEA98562EFA41FBFFFAAAFB2D3D1C" box="[1078,1159,1356,1380]" italics="true" pageId="1" pageNumber="2">B. kala</emphasis>
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Prout
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</taxonomicName>
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has atypical male genitalia sharing some features with
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<taxonomicName id="01A94D09562EFA41FDF6FA97FC8F3DFF" authority="Moore" authorityName="Moore" box="[575,805,1391,1415]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="striatovirens">
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||
<emphasis id="F4DDEA98562EFA41FDF6FA97FD613DFF" box="[575,715,1392,1415]" italics="true" pageId="1" pageNumber="2">striatovirens</emphasis>
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||
Moore
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||
</taxonomicName>
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||
”. The male genitalia of
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<taxonomicName id="01A94D09562EFA41FBF4FA97FB043DF0" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" box="[1085,1198,1392,1416]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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||
<emphasis id="F4DDEA98562EFA41FBF4FA97FB043DF0" box="[1085,1198,1392,1416]" italics="true" pageId="1" pageNumber="2">Anabelcia</emphasis>
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||
</taxonomicName>
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differ from those of
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<taxonomicName id="01A94D09562EFA41FF5EFA73FF513DD4" box="[151,251,1428,1452]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="F4DDEA98562EFA41FF5EFA73FF513DD4" box="[151,251,1428,1452]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
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||
</taxonomicName>
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||
(
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||
<figureCitation id="5E922A0F562EFA41FEC6FA73FECF3DD4" box="[271,357,1428,1452]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1841,1864]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-75@9.[151,1436,193,1821]" captionTargetPageId="9" captionText="FIGURES 17–22. Belciana and Anabelcia spp., structures of palps (17–20) and male genitalia (21, 22). The distinctive characters marked by lines. 17. B. bifornis; 18. A. kala; 19. A. nepalensis; 20. A. taiwana; 21. A. biformis, Indonesia, Sumatra, genit. prep. 4160GB (GB); 22. A. kala, Indonesia, Sumatra, genit. prep. 12018GB (GB)." pageId="1" pageNumber="2">Fig. 21</figureCitation>
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) by stronger apically tapered valva covered with strong setae, strong plate-like clasper, by presence of strong saccular extension, shape of juxta and uncus. Aedeagus stronger sclerotised than in
|
||
<taxonomicName id="01A94D09562EFA41FAF8FA5FFA3E3DA8" box="[1329,1428,1464,1488]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F4DDEA98562EFA41FAF8FA5FFA3E3DA8" box="[1329,1428,1464,1488]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
|
||
</taxonomicName>
|
||
, carina usually with conical spine, vesica tubular, with dorsal diverticula bearing thin but strong spine-like cornuti. In the male genitalia of
|
||
<taxonomicName id="01A94D09562EFA41FE7AF9E7FDBD3E60" box="[435,535,1536,1560]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F4DDEA98562EFA41FE7AF9E7FDBD3E60" box="[435,535,1536,1560]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
|
||
</taxonomicName>
|
||
valva broad, weak, rounded apically, sclerotised in costal part only; sacculus small, rounded, juxta amorphic, plate-like, uncus strongly arched, usually hooked, clasper and harpe very small; ventral margin of valva weakly sclerotised or membranous, the base of valva with large membranous gland bearing tuft of hairs. Aedeagus with carina armed by small spines, vesica bulbous, armed with stick-like or needle-like cornuti. The female genitalia in both genera are rather similar, but
|
||
<taxonomicName id="01A94D09562EFA41FC50F977FC573ED0" box="[921,1021,1680,1704]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F4DDEA98562EFA41FC50F977FC573ED0" box="[921,1021,1680,1704]" italics="true" pageId="1" pageNumber="2">Belciana</emphasis>
|
||
</taxonomicName>
|
||
has membranous ductus bursae and heavily sclerotised cervix bursae, while in new genus the ductus bursae usually sclerotised and cervix bursae often not expressed.
|
||
</paragraph>
|
||
<paragraph id="C616368A562EFA42FF0EF91BFDF739D4" blockId="1.[151,1436,1031,2028]" lastBlockId="2.[151,1437,151,1292]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
|
||
The genus
|
||
<taxonomicName id="01A94D09562EFA41FE85F91BFE173F6C" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" box="[332,445,1788,1812]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F4DDEA98562EFA41FE85F91BFE173F6C" box="[332,445,1788,1812]" italics="true" pageId="1" pageNumber="2">Anabelcia</emphasis>
|
||
</taxonomicName>
|
||
comprises three lineages or species-groups, which are easily distinguished by their external and genital features. The first group represents by
|
||
<taxonomicName id="01A94D09562EFA41FCF2F8C7FC213F40" authorityName="Prout" authorityYear="1924" box="[827,907,1824,1848]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="kala">
|
||
<emphasis id="F4DDEA98562EFA41FCF2F8C7FC213F40" box="[827,907,1824,1848]" italics="true" pageId="1" pageNumber="2">A. kala</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="01A94D09562EFA41FC55F8C7FB983F40" authorityName="Haruta" authorityYear="1993" box="[924,1074,1824,1848]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="nepalensis">
|
||
<emphasis id="F4DDEA98562EFA41FC55F8C7FB983F40" box="[924,1074,1824,1848]" italics="true" pageId="1" pageNumber="2">A. nepalensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="01A94D09562EFA41FBA5F8C7FB123F40" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" box="[1132,1208,1824,1848]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="thai">
|
||
<emphasis id="F4DDEA98562EFA41FBA5F8C7FB123F40" box="[1132,1208,1824,1848]" italics="true" pageId="1" pageNumber="2">A. thai</emphasis>
|
||
</taxonomicName>
|
||
. It characterized by rather uniform mossy-green with brown patches forewing pattern, weakly developed saccular extension and presence of diverticulum armed with row of spine-like cornuti in vesica. The female genitalia could be characterized by elongated split antrum and presence of sclerotised patch in joining of the ductus bursa with corpus bursa. The second group includes
|
||
<taxonomicName id="01A94D09562EFA41FDDCF857FD243FB0" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" baseAuthorityName="Remm" baseAuthorityYear="1983" box="[533,654,1968,1992]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="siitanae">
|
||
<emphasis id="F4DDEA98562EFA41FDDCF857FD243FB0" box="[533,654,1968,1992]" italics="true" pageId="1" pageNumber="2">A. siitanae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="F4DDEA98562EFA41FD0DF857FCEF3FB0" box="[708,837,1968,1992]" italics="true" pageId="1" pageNumber="2">
|
||
<taxonomicName id="01A94D09562EFA41FD0DF857FCEB3FB0" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" box="[708,833,1968,1992]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="taiwana">A. taiwana</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Both species have characteristic white mark between orbicular and reniform; in the male genitalia saccular extension is much stronger than in
|
||
<taxonomicName id="01A94D09562EFA41FBB0F833FB623F94" authorityName="Prout" authorityYear="1924" box="[1145,1224,2004,2028]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="kala">
|
||
<emphasis id="F4DDEA98562EFA41FBB0F833FB623F94" box="[1145,1224,2004,2028]" italics="true" pageId="1" pageNumber="2">A. kala</emphasis>
|
||
</taxonomicName>
|
||
species-group; the female genitalia have rounded split antevaginal plate, sclerotised patch in joining of the ductus with corpus bursa not expressed. The third group comprised by single species—
|
||
<taxonomicName id="01A94D09562DFA42FCA5FF5BFBA538AC" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" baseAuthorityName="Leech" baseAuthorityYear="1900" box="[876,1039,188,212]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="staudingeri">
|
||
<emphasis id="F4DDEA98562DFA42FCA5FF5BFBA538AC" box="[876,1039,188,212]" italics="true" pageId="2" pageNumber="3">A. staudingeri</emphasis>
|
||
</taxonomicName>
|
||
. The imagoes is characterized by bluish-green ground colour of forewing with brown patches and thin distinct transversal lines; in contrast to
|
||
<taxonomicName id="01A94D09562DFA42FA84FF07FA313880" authorityName="Prout" authorityYear="1924" box="[1357,1435,224,248]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kala">
|
||
<emphasis id="F4DDEA98562DFA42FA84FF07FA313880" box="[1357,1435,224,248]" italics="true" pageId="2" pageNumber="3">A. kala</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="01A94D09562DFA42FF03FEE3FEEF3964" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" baseAuthorityName="Remm" baseAuthorityYear="1983" box="[202,325,260,284]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="siitanae">
|
||
<emphasis id="F4DDEA98562DFA42FF03FEE3FEEF3964" box="[202,325,260,284]" italics="true" pageId="2" pageNumber="3">A. siitanae</emphasis>
|
||
</taxonomicName>
|
||
species-groups in
|
||
<taxonomicName id="01A94D09562DFA42FDE9FEE3FD683964" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" baseAuthorityName="Leech" baseAuthorityYear="1900" box="[544,706,260,284]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="staudingeri">
|
||
<emphasis id="F4DDEA98562DFA42FDE9FEE3FD683964" box="[544,706,260,284]" italics="true" pageId="2" pageNumber="3">A. staudingeri</emphasis>
|
||
</taxonomicName>
|
||
the hindwing are brownish-grey, paler in inner part. The male genitalia of
|
||
<taxonomicName id="01A94D09562DFA42FED6FECFFE153938" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" baseAuthorityName="Leech" baseAuthorityYear="1900" box="[287,447,296,320]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="staudingeri">
|
||
<emphasis id="F4DDEA98562DFA42FED6FECFFE153938" box="[287,447,296,320]" italics="true" pageId="2" pageNumber="3">A. staudingeri</emphasis>
|
||
</taxonomicName>
|
||
are characterize by presence of large basal extension of sacculus, the vesica differs by presence of large basal and small, sclerotised subapical diverticula, which are not presented in other species-group. The female genitalia differ from those in other groups by rather narrow, elongated antrum, with deep apical cut and long and thin membranous ductus bursa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8EB36501562DFA42FF0EFE50FC493CE7" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="C616368A562DFA42FF0EFE50FAB43ADF" blockId="2.[151,1437,151,1292]" pageId="2" pageNumber="3">
|
||
<emphasis id="F4DDEA98562DFA42FF0EFE50FE0239A8" bold="true" box="[199,424,439,464]" pageId="2" pageNumber="3">
|
||
<emphasis id="F4DDEA98562DFA42FF0EFE50FEF339A8" box="[199,345,439,464]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">Description.</emphasis>
|
||
Adult
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="5E922A0F562DFA42FE75FE50FD9F39B7" box="[444,565,439,463]" captionStart-0="FIGURES 1–8" captionStart-1="FIGURES 9–16" captionStartId-0="7.[151,263,1841,1864]" captionStartId-1="8.[151,263,1841,1864]" captionTargetBox-0="[151,1436,194,1821]" captionTargetBox-1="[154,1436,193,1821]" captionTargetId-0="figure-104@7.[151,1436,193,1821]" captionTargetId-1="figure-99@8.[151,1436,193,1821]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 1–8. Belciana and Anabelcia spp., adults. 1. B. biformis, female, Malaysia, Borneo (GB); 2. A. kala, male, Indonesia, Sumatra, genit. prep. 12018GB (ZMUC); 3. A. kala, male, Malaysia (Pahang) (AZ); 4. A. kala, female, Philippines, Palawan, genit. prep. 12063GB (ZFMK); 5. A. kala, male, Vietnam, genit. prep. 7348GB (GB); 6. A. nepalensis, male, Nepal, genit. prep. 4161GB (ZSM); 7. A. nepalensis, female, Nepal, genit. prep. 4162GB (ZSM); 8. A. nepalensis, female, N India, genit. prep. 7413GB (ZSM)." captionText-1="FIGURES 9–16. Anabelcia spp., adults. 9. A. thai, female, HT, Thailand genit. prep. 7414GB (GB); 10. A. thai, female, PT, Thailand (AB); 11. A. taiwana, male, HT, Taiwan, genit. prep. 7340GB (GB); 12. A. taiwana, male, HT, Taiwan, (GR); 13. A. taiwana, male, HT, Taiwan, (GR); 14. A. siitanae, male, Russia, Primorye (FMNH); 15. A. staudingeri, male, China (AB); 16. A. staudingeri, male, China (ZFMK)." pageId="2" pageNumber="3">Figs 2–16</figureCitation>
|
||
,
|
||
<figureCitation id="5E922A0F562DFA42FD8CFE50FD3A39B7" box="[581,656,439,463]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1841,1864]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-75@9.[151,1436,193,1821]" captionTargetPageId="9" captionText="FIGURES 17–22. Belciana and Anabelcia spp., structures of palps (17–20) and male genitalia (21, 22). The distinctive characters marked by lines. 17. B. bifornis; 18. A. kala; 19. A. nepalensis; 20. A. taiwana; 21. A. biformis, Indonesia, Sumatra, genit. prep. 4160GB (GB); 22. A. kala, Indonesia, Sumatra, genit. prep. 12018GB (GB)." pageId="2" pageNumber="3">18–20</figureCitation>
|
||
). Medium-sized moths, wingspan
|
||
<quantity id="01519B6F562DFA42FBEDFE50FB0F39B7" box="[1060,1189,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.0" metricValueMin="3.8" pageId="2" pageNumber="3" unit="mm" value="44.0" valueMax="50.0" valueMin="38.0">38–50 mm</quantity>
|
||
. Head with rounded, slightly bulging frons covered with scales; labial palps relatively long, extending over frons, with slightly curving upward with 3rd segment stick-like, large, about half as long as 2nd segment; proboscis developed; antennae filiform, fasciculate, scapus white; eye moderate, rounded; ocelli present; head and thorax covered with brown and green scales, thoracic and abdominal crests present, formed with green and black scales. Leg segments ringed with white. Forewing venation typically quadrifine with Cu vein appearing four branched; hindwing with typical for pantheine venation (i.e., M2 fully developed, arises near low angle of cell. Abdomen without coremata.
|
||
</paragraph>
|
||
<paragraph id="C616368A562DFA42FF0EFD53FEE03C4C" blockId="2.[151,1437,151,1292]" pageId="2" pageNumber="3">
|
||
<emphasis id="F4DDEA98562DFA42FF0EFD53FDB83AB4" box="[199,530,691,716]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">
|
||
<emphasis id="F4DDEA98562DFA42FF0EFD53FED83AB4" bold="true" box="[199,370,691,716]" pageId="2" pageNumber="3">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="5E922A0F562DFA42FE4BFD53FDAC3AB4" box="[386,518,692,716]" captionStart-0="FIGURES 17–22" captionStart-1="FIGURES 23–25" captionStart-2="FIGURES 26–28" captionStartId-0="9.[151,264,1841,1864]" captionStartId-1="10.[151,264,1841,1864]" captionStartId-2="11.[151,264,1841,1864]" captionTargetBox-0="[151,1436,193,1809]" captionTargetBox-1="[151,1424,230,1813]" captionTargetBox-2="[161,1416,195,1812]" captionTargetId-0="figure-75@9.[151,1436,193,1821]" captionTargetId-1="figure-62@10.[151,1436,193,1821]" captionTargetId-2="figure-52@11.[151,1436,193,1821]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURES 17–22. Belciana and Anabelcia spp., structures of palps (17–20) and male genitalia (21, 22). The distinctive characters marked by lines. 17. B. bifornis; 18. A. kala; 19. A. nepalensis; 20. A. taiwana; 21. A. biformis, Indonesia, Sumatra, genit. prep. 4160GB (GB); 22. A. kala, Indonesia, Sumatra, genit. prep. 12018GB (GB)." captionText-1="FIGURES 23–25. Anabelcia spp., male genitalia. 23. A. kala, North Vietnam, genit. prep. 7448GB (ZSM); 24. A. nepalensis, male, North India, genit. prep. 7413GB (ZSM); 25. A. nepalensis, Nepal, genit. prep. 4161GB (ZSM)." captionText-2="FIGURES 26–28. Anabelcia spp., male genitalia. 26. A. taiwana, Taiwan, genit. prep. 7250LR (HNHM); 27. A. siitanae, Russia, Primorye, genit. prep. 266VK (IBSS); 28. A. staudingeri, China, genit. prep. 7375GB (GB)." pageId="2" pageNumber="3">Figs 22–28</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Subscaphium sclerotised. Uncus moderate in length, rather thin, slightly arched apically, cylindrical, tapered to spine-like apex, tegumen moderately broad, about 2 times as wide ventrally as medially; vinculum shorter than tegumen, narrow with rather broad V-like saccus; paratergal sclerites broad, flat; juxta relatively small, heavily sclerotised plate-like, broader anteriorly with central sclerotised bar posteriorly or with central extension; valvae symmetrical, broader mesially, tapered apically, rounded at apex, cucullus not separated, corona not expressed; valvae surface covered with dense strong setae irremovable by maceration; sacculus moderate, elongate, with rather broad lobe and strong longitudinal saccular extension or fold; clasper positioned in basal half of valva, like triangular plate; harpe not expressed; costa strong, heavily sclerotised; ventral margin of valva in basal half weakly sclerotised. Aedeagus straight or somewhat curved, vesica tubular, extended medially, with medial diverticula, bearing row of medium-sized cornuti or with basal and medial diverticuli without cornuti.
|
||
</paragraph>
|
||
<paragraph id="C616368A562DFA42FF0EFBA7FC493CE7" blockId="2.[151,1437,151,1292]" pageId="2" pageNumber="3">
|
||
<emphasis id="F4DDEA98562DFA42FF0EFBA7FD923C2F" box="[199,568,1087,1112]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">
|
||
<emphasis id="F4DDEA98562DFA42FF0EFBA7FE3B3C20" bold="true" box="[199,401,1087,1112]" pageId="2" pageNumber="3">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="5E922A0F562DFA42FE6DFBD8FD863C2F" box="[420,556,1087,1111]" captionStart="FIGURES 29–34" captionStartId="12.[151,264,1841,1864]" captionTargetBox="[161,1420,208,1813]" captionTargetId="figure-86@12.[151,1436,193,1821]" captionTargetPageId="12" captionText="FIGURES 29–34. Belciana and Anabelcia spp., female genitalia. 29. B. biformis, Sumatra, genit. prep. 4183GB (ZSM); 30. A. kala, Sumatra, genit. prep. Z4178GB (ZSM); 31. A. nepalensis, Nepal, genit. prep. Z4162BG (ZSM); 32. A. thai, HT, genit. prep. 7414GB; 33. A. siitanae, Russia, Primorye, genit. prep. 654VK; 34. A. staudingeri, North Korea, genit. prep. 2165VK (ZFMK)." pageId="2" pageNumber="3">Figs 30–34</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Papillae anales small, quadrangular; apophyses anteriores and posteriores short, antrum shallow cup-like or deep, split; ductus bursae short, usually sclerotised, corpus bursae elongate, ovoid, in some species with sclerotised patch in joining with ductus bursae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8EB36501562DFA42FF0EFB4CFB7B3C9F" pageId="2" pageNumber="3" type="distribution">
|
||
<paragraph id="C616368A562DFA42FF0EFB4CFB7B3C9F" blockId="2.[151,1437,151,1292]" pageId="2" pageNumber="3">
|
||
<emphasis id="F4DDEA98562DFA42FF0EFB4CFECA3CBC" bold="true" box="[199,352,1195,1220]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">Distribution.</emphasis>
|
||
The genus occurs predominantly in the Oriental region from North
|
||
<collectingCountry id="BEBE761A562DFA42FB7AFB4BFB463CBC" box="[1203,1260,1196,1220]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="BEBE761A562DFA42FAFDFB4BFADD3CBC" box="[1332,1399,1196,1220]" name="Nepal" pageId="2" pageNumber="3">Nepal</collectingCountry>
|
||
to
|
||
<collectingCountry id="BEBE761A562DFA42FF5EFB28FEBF3C9F" box="[151,277,1231,1255]" name="Philippines" pageId="2" pageNumber="3">Philippines</collectingCountry>
|
||
and in the Eastern Palaearctic (two species live in the China-Manchurian subregion).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8EB36501562DFA42FF0EFB13FC623D73" box="[199,968,1268,1292]" pageId="2" pageNumber="3" type="etymology">
|
||
<paragraph id="C616368A562DFA42FF0EFB13FC623D73" blockId="2.[151,1437,151,1292]" box="[199,968,1268,1292]" pageId="2" pageNumber="3">
|
||
<emphasis id="F4DDEA98562DFA42FF0EFB13FEE73D74" bold="true" box="[199,333,1268,1292]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">Etymology.</emphasis>
|
||
The generic name
|
||
<taxonomicName id="01A94D09562DFA42FDEDFB13FD3F3D74" authorityName="Behounek & V.S.Kononenko" authorityYear="2012" box="[548,661,1268,1292]" class="Insecta" family="Noctuidae" genus="Anabelcia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F4DDEA98562DFA42FDEDFB13FD3F3D74" box="[548,661,1268,1292]" italics="true" pageId="2" pageNumber="3">Anabelcia</emphasis>
|
||
</taxonomicName>
|
||
is an anagram of
|
||
<taxonomicName id="01A94D09562DFA42FC94FB13FC6B3D74" box="[861,961,1268,1292]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F4DDEA98562DFA42FC94FB13FC6B3D74" box="[861,961,1268,1292]" italics="true" pageId="2" pageNumber="3">Belciana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |