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<document id="0B3AC8DB270733CA1CCA5C1EB33EEA93" ID-DOI="10.5281/zenodo.198881" ID-GBIF-Dataset="66ad2a9f-d785-4cc2-b637-ab82e061eaa7" ID-ISSN="1175-5326" ID-Zenodo-Dep="198881" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460316322591" checkinUser="plazi" docAuthor="Wolski, Andrzej" docDate="2010" docId="03EB772EFFEDFF85FF4B939FFD195635" docLanguage="en" docName="zt02653p036.pdf" docOrigin="Zootaxa 2653" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Rhinocylapus scutatus Hsiao" docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="FFD20F56FFF6FF99FFDC9275FFEC5519" masterDocTitle="Revision of the Rhinocylapus - group (Hemiptera: Heteroptera: Miridae: Cylapinae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="28" updateTime="1698255163851" updateUser="plazi">
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<mods:title id="FA66C22E4AA1C60287829B7641EC2902">Revision of the Rhinocylapus - group (Hemiptera: Heteroptera: Miridae: Cylapinae)</mods:title>
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<mods:date id="BE485F67C13DDC7F58D9687366C18DC8">2010</mods:date>
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<mods:identifier id="0D2655594209EDC2504C4001D0FF3276" type="DOI">10.5281/zenodo.198881</mods:identifier>
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<treatment id="03EB772EFFEDFF85FF4B939FFD195635" ID-DOI="http://doi.org/10.5281/zenodo.6197310" ID-GBIF-Taxon="119401758" ID-Zenodo-Dep="6197310" LSID="urn:lsid:plazi:treatment:03EB772EFFEDFF85FF4B939FFD195635" httpUri="http://treatment.plazi.org/id/03EB772EFFEDFF85FF4B939FFD195635" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="C35895B3FFEDFF82FF4B939FFE02573E" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BFDC638FFEDFF82FF4B939FFE1A571D" blockId="27.[151,502,490,551]" box="[151,502,490,516]" pageId="27" pageNumber="28">
<heading id="D0B57154FFEDFF82FF4B939FFE1A571D" bold="true" box="[151,502,490,516]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<taxonomicName id="4C42BDBBFFEDFF82FF4B939FFE1A571D" ID-CoL="4S8SD" authority="Hsiao" authorityName="Hsiao" box="[151,502,490,516]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B9361A2AFFEDFF82FF4B939FFE1A571D" bold="true" box="[151,502,490,516]" pageId="27" pageNumber="28">
<emphasis id="B9361A2AFFEDFF82FF4B939FFE4B571D" bold="true" box="[151,423,490,516]" italics="true" pageId="27" pageNumber="28">Rhinocylapus scutatus</emphasis>
Hsiao
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFDC638FFEDFF82FF4B9078FE02573E" blockId="27.[151,502,490,551]" box="[151,494,525,551]" pageId="27" pageNumber="28">
(
<figureCitation id="1379DABDFFEDFF82FF439078FF1D573E" box="[159,241,525,551]" captionStart="FIGURE 2" captionStartId="5.[151,255,1845,1869]" captionTargetBox="[183,1430,209,1812]" captionTargetId="figure@5.[156,1430,194,1823]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus color photographs of the Rhinocylapus - group: A – B, Proamblia cunealis: A—paralectotype (Ψ), B—lectotype; C – D Proamblia elongata: C—holotype, D— ♂: Sandakan Borneo; E— Mycetocylapus minor (♂: Los Banos, P. I.); F— Mycetocylapus major (Ψ: Los Banos, P. I.); G— Mycetocylapus pacificus (holotype); H – I, Rhinocylapidius velocipedoides: H—lectotype, I— ♂: Formosa; Fuhosho; J— Rhinocylapus acutangulus (holotype); K— Rh. redeii (holotype); L— Rh. scutatus (♂: paratype); M— Rh. simplicicollis (holotype); N— Rh. sumatranus (holotype); O – P, Rh. pallescens: O—Ψ: paratype, P—holotype; Q-R, Rh. vittatus: Q—holotype, R— ♂: Sandakan, Borneo." httpUri="https://zenodo.org/record/198883/files/figure.png" pageId="27" pageNumber="28">Figs. 2</figureCitation>
L, 3D, 4E, 9A–B, 13)
</paragraph>
</subSubSection>
<subSubSection id="C35895B3FFEDFF82FF4B902FFB615788" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BFDC638FFEDFF82FF4B902FFDD15768" blockId="27.[151,1165,602,657]" box="[151,573,602,625]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB52E116FFEDFF82FF4B902FFDD15768" box="[151,573,602,625]" pageId="27" pageNumber="28">
<taxonomicName id="4C42BDBBFFEDFF82FF4B902FFDD15768" ID-CoL="4S8SD" authority="Hsiao 1944: 380" authorityName="Hsiao" authorityPageNumber="380" authorityYear="1944" box="[151,573,602,625]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B9361A2AFFEDFF82FF4B902FFE695768" box="[151,389,602,625]" italics="true" pageId="27" pageNumber="28">Rhinocylapus scutatus</emphasis>
<treatmentCitation id="0AE3E029FFEDFF82FE50902FFDD15768" author="Hsiao" box="[396,573,602,625]" page="380" pageId="27" pageNumber="28" year="1944">
<bibRefCitation id="EFD3BBC9FFEDFF82FE50902FFDE65768" author="Hsiao" box="[396,522,602,625]" pageId="27" pageNumber="28" refString="Hsiao, T-y. (1944) New genera and species of Oriental and Australian plant bugs in the United States National Museum. Proceedings of the United States National Museum, 95, 369 - 396." type="journal article" year="1944">Hsiao 1944</bibRefCitation>
: 380
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFDC638FFEDFF82FF4B900FFB615788" blockId="27.[151,1165,602,657]" box="[151,1165,634,657]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB52E116FFEDFF82FF4B900FFB615788" box="[151,1165,634,657]" pageId="27" pageNumber="28">
<taxonomicName id="4C42BDBBFFEDFF82FF4B900FFCEC5788" authority="Carvalho 1955b: 221" authorityName="Carvalho" authorityPageNumber="221" authorityYear="1955" box="[151,768,634,657]" class="Insecta" family="Miridae" genus="Rhinocylapidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B9361A2AFFEDFF82FF4B900FFE475788" box="[151,427,634,657]" italics="true" pageId="27" pageNumber="28">Rhinocylapidius scutatus:</emphasis>
<treatmentCitation id="0AE3E029FFEDFF82FE6E900FFD795788" author="Carvalho" box="[434,661,634,657]" page="221" pageId="27" pageNumber="28" year="1955">
<bibRefCitation id="EFD3BBC9FFEDFF82FE6E900FFD8C5788" author="Carvalho" box="[434,608,634,657]" pageId="27" pageNumber="28" refString="Carvalho, J. C. M. (1955 b) Analecta Miridologica: Miscellaneous observations in some American museums and bibliography. Revista Chilena de Entomologia, 4, 221 - 227." type="journal article" year="1955" yearSuffix="b">Carvalho 1955b</bibRefCitation>
: 221
</treatmentCitation>
, 1957: 23
</taxonomicName>
;
<treatmentCitation id="0AE3E029FFEDFF82FCD0900FFC5A5788" author="Schuh" box="[780,950,634,657]" page="35" pageId="27" pageNumber="28" year="1995">
<bibRefCitation id="EFD3BBC9FFEDFF82FCD0900FFC625788" author="Schuh" box="[780,910,634,657]" pageId="27" pageNumber="28" refString="Schuh, R. T. (1995) Plant bugs of the World (Insecta: Heteroptera: Miridae). New York Entomological Society, New York City, N. Y. 1329 pp." type="book" year="1995">Schuh 1995</bibRefCitation>
: 35
</treatmentCitation>
;
<treatmentCitation id="0AE3E029FFEDFF82FC1D900FFB615788" author="Gorczyca" box="[961,1165,634,657]" page="72" pageId="27" pageNumber="28" year="2006">
<bibRefCitation id="EFD3BBC9FFEDFF82FC1D900FFB8B5788" author="Gorczyca" box="[961,1127,634,657]" pageId="27" pageNumber="28" refString="Gorczyca, J. (2006) The Catalogue of the subfamily Cylapinae Kirkaldy, 1903 of the World (Hemiptera, Heteroptera, Miridae). Monographs of the Upper Silesian Museum, 5, 1 - 100." type="journal article" year="2006">Gorczyca 2006</bibRefCitation>
: 72
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35895B3FFEDFF82FF4B90B7FB1156FB" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8BFDC638FFEDFF82FF4B90B7FB1156FB" blockId="27.[151,1437,706,2002]" pageId="27" pageNumber="28">
<emphasis id="B9361A2AFFEDFF82FF4B90B7FEFB57C5" bold="true" box="[151,279,706,732]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
Recognized by the following set of characters: spines on labrum present (
<figureCitation id="1379DABDFFEDFF82FB5F90B7FB2457C5" box="[1155,1224,706,732]" captionStart="FIGURE 3" captionStartId="6.[151,257,1125,1149]" captionTargetBox="[203,1387,194,1103]" captionTargetId="figure@6.[199,1387,194,1103]" captionTargetPageId="6" captionText="FIGURE 3. Left lateral view: A, Rhinocylapus acutangulus; B, Rh. redeii; C, Rh. simplicicollis; D, Rh. scutatus; E, Rhinocylapidius velocipedoides. SP—spines on labrum." httpUri="https://zenodo.org/record/198884/files/figure.png" pageId="27" pageNumber="28">Fig. 3</figureCitation>
D); lateral margin of anterior lobe of pronotum gently carinate, with row of punctures, each bearing single, decumbent seta; scutellum flattened, with large, yellowish patch medially (
<figureCitation id="1379DABDFFEDFF82FCBE9178FC55563E" box="[866,953,781,807]" captionStart="FIGURE 2" captionStartId="5.[151,255,1845,1869]" captionTargetBox="[183,1430,209,1812]" captionTargetId="figure@5.[156,1430,194,1823]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus color photographs of the Rhinocylapus - group: A – B, Proamblia cunealis: A—paralectotype (Ψ), B—lectotype; C – D Proamblia elongata: C—holotype, D— ♂: Sandakan Borneo; E— Mycetocylapus minor (♂: Los Banos, P. I.); F— Mycetocylapus major (Ψ: Los Banos, P. I.); G— Mycetocylapus pacificus (holotype); H – I, Rhinocylapidius velocipedoides: H—lectotype, I— ♂: Formosa; Fuhosho; J— Rhinocylapus acutangulus (holotype); K— Rh. redeii (holotype); L— Rh. scutatus (♂: paratype); M— Rh. simplicicollis (holotype); N— Rh. sumatranus (holotype); O – P, Rh. pallescens: O—Ψ: paratype, P—holotype; Q-R, Rh. vittatus: Q—holotype, R— ♂: Sandakan, Borneo." httpUri="https://zenodo.org/record/198883/files/figure.png" pageId="27" pageNumber="28">Figs. 2</figureCitation>
L, 3D); endosoma with large, distinctly curved, and twisted medially sclerite (
<figureCitation id="1379DABDFFEDFF82FDBF9147FD5E5655" box="[611,690,818,844]" captionStart="FIGURE 13" captionStartId="28.[151,255,1939,1963]" captionTargetBox="[280,1311,886,1915]" captionTargetId="figure@28.[274,1312,869,1918]" captionTargetPageId="28" captionText="FIGURE 13. Male genitalia of Rhinocylapus scutatus: A, endosoma, B, left paramere, left lateral view; C, left paramere, dorsal view; D, right paramere. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/198894/files/figure.png" pageId="27" pageNumber="28">Fig.13</figureCitation>
A). The species is most closely related to
<taxonomicName id="4C42BDBBFFEDFF82FB709147FA915652" box="[1196,1405,818,843]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="simplicicollis">
<emphasis id="B9361A2AFFEDFF82FB709147FA915652" box="[1196,1405,818,843]" italics="true" pageId="27" pageNumber="28">Rh. simplicicollis</emphasis>
</taxonomicName>
in sharing similar body coloration, a flattened anterior lobe of pronotum, and non-rugose posterior lobe. It can be readily distinguished, however, by having a large, yellowish patch on the scutellum medially and by short and decumbent setae.
<taxonomicName id="4C42BDBBFFEDFF82FEBB91D7FE1A56A2" box="[359,502,930,955]" class="Insecta" family="Miridae" genus="Rhinocylapidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B9361A2AFFEDFF82FEBB91D7FE1A56A2" box="[359,502,930,955]" italics="true" pageId="27" pageNumber="28">Rh. scutatus</emphasis>
</taxonomicName>
shares a row of punctures on the lateral margin of the anterior lobe of pronotum with
<taxonomicName id="4C42BDBBFFEDFF82FF0E91BDFEBB56F8" box="[210,343,968,994]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vittatus">
<emphasis id="B9361A2AFFEDFF82FF0E91BDFF1E56F8" box="[210,242,968,993]" italics="true" pageId="27" pageNumber="28">Rh</emphasis>
.
<emphasis id="B9361A2AFFEDFF82FEDD91BDFEBB56F8" box="[257,343,968,993]" italics="true" pageId="27" pageNumber="28">vittatus</emphasis>
</taxonomicName>
, but can be easily distinguished by a distinctly darker coloration of the dorsum.
</paragraph>
</subSubSection>
<subSubSection id="C35895B3FFEDFF85FF199198FD8655CE" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="8BFDC638FFEDFF82FF199198FD1553BE" blockId="27.[151,1437,706,2002]" pageId="27" pageNumber="28">
<emphasis id="B9361A2AFFEDFF82FF199198FE22511E" bold="true" box="[197,462,1005,1031]" pageId="27" pageNumber="28">Redescription. Male</emphasis>
.
<emphasis id="B9361A2AFFEDFF82FE039198FD43511E" bold="true" box="[479,687,1005,1031]" pageId="27" pageNumber="28">COLORATION</emphasis>
(
<figureCitation id="1379DABDFFEDFF82FD1D9198FCF5511E" box="[705,793,1005,1031]" captionStart="FIGURE 2" captionStartId="5.[151,255,1845,1869]" captionTargetBox="[183,1430,209,1812]" captionTargetId="figure@5.[156,1430,194,1823]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus color photographs of the Rhinocylapus - group: A – B, Proamblia cunealis: A—paralectotype (Ψ), B—lectotype; C – D Proamblia elongata: C—holotype, D— ♂: Sandakan Borneo; E— Mycetocylapus minor (♂: Los Banos, P. I.); F— Mycetocylapus major (Ψ: Los Banos, P. I.); G— Mycetocylapus pacificus (holotype); H – I, Rhinocylapidius velocipedoides: H—lectotype, I— ♂: Formosa; Fuhosho; J— Rhinocylapus acutangulus (holotype); K— Rh. redeii (holotype); L— Rh. scutatus (♂: paratype); M— Rh. simplicicollis (holotype); N— Rh. sumatranus (holotype); O – P, Rh. pallescens: O—Ψ: paratype, P—holotype; Q-R, Rh. vittatus: Q—holotype, R— ♂: Sandakan, Borneo." httpUri="https://zenodo.org/record/198883/files/figure.png" pageId="27" pageNumber="28">Figs. 2</figureCitation>
L). Dorsal surface dark castaneous.
<emphasis id="B9361A2AFFEDFF82FB0B9198FAF7511E" bold="true" box="[1239,1307,1005,1031]" italics="true" pageId="27" pageNumber="28">Head</emphasis>
. Antennal segment I brown; segment II brown, with yellowish red, wide ring beyond medial part; segment III pale yellow basally, then dark yellow; segment IV dark yellow; rostrum brown.
<emphasis id="B9361A2AFFEDFF82FB849642FB5B5148" bold="true" box="[1112,1207,1079,1105]" italics="true" pageId="27" pageNumber="28">Thorax</emphasis>
.
<emphasis id="B9361A2AFFEDFF82FB15964DFEE2516F" italics="true" pageId="27" pageNumber="28">Mesoscutum and scutellum</emphasis>
. Scutellum with large, yellowish patch medially.
<emphasis id="B9361A2AFFEDFF82FCA19628FBAB516F" box="[893,1095,1117,1142]" italics="true" pageId="27" pageNumber="28">Thoracic pleura</emphasis>
. Dark brown.
<emphasis id="B9361A2AFFEDFF82FB229628FA79516E" bold="true" box="[1278,1429,1117,1143]" italics="true" pageId="27" pageNumber="28">Hemelytron</emphasis>
. Cuneus with inner angle yellowish red.
<emphasis id="B9361A2AFFEDFF82FD5E96F6FD575185" box="[642,699,1155,1180]" italics="true" pageId="27" pageNumber="28">Legs</emphasis>
. Brownish yellow; coxa and trochanters dark brown; femur brown; tibia brown; meso- and metatibia paler, yellowish apically.
<emphasis id="B9361A2AFFEDFF82FC6296D2FBDF51D8" bold="true" box="[958,1075,1191,1217]" italics="true" pageId="27" pageNumber="28">Abdomen</emphasis>
. Ranging from brown to dark brown.
<emphasis id="B9361A2AFFEDFF82FF2896B8FD4751FE" bold="true" box="[244,683,1229,1255]" pageId="27" pageNumber="28">STRUCTURE AND VESTITURE</emphasis>
(
<figureCitation id="1379DABDFFEDFF82FD6196B8FCEB51FE" box="[701,775,1229,1255]" captionStart="FIGURE 2" captionStartId="5.[151,255,1845,1869]" captionTargetBox="[183,1430,209,1812]" captionTargetId="figure@5.[156,1430,194,1823]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus color photographs of the Rhinocylapus - group: A – B, Proamblia cunealis: A—paralectotype (Ψ), B—lectotype; C – D Proamblia elongata: C—holotype, D— ♂: Sandakan Borneo; E— Mycetocylapus minor (♂: Los Banos, P. I.); F— Mycetocylapus major (Ψ: Los Banos, P. I.); G— Mycetocylapus pacificus (holotype); H – I, Rhinocylapidius velocipedoides: H—lectotype, I— ♂: Formosa; Fuhosho; J— Rhinocylapus acutangulus (holotype); K— Rh. redeii (holotype); L— Rh. scutatus (♂: paratype); M— Rh. simplicicollis (holotype); N— Rh. sumatranus (holotype); O – P, Rh. pallescens: O—Ψ: paratype, P—holotype; Q-R, Rh. vittatus: Q—holotype, R— ♂: Sandakan, Borneo." httpUri="https://zenodo.org/record/198883/files/figure.png" pageId="27" pageNumber="28">Fig. 2</figureCitation>
L, 3D, 9A–B). Dorsal surface covered with fine, dark, relatively long setae.
<emphasis id="B9361A2AFFEDFF82FE789687FE045015" bold="true" box="[420,488,1266,1292]" italics="true" pageId="27" pageNumber="28">Head</emphasis>
. Shining, covered with very sparse, fine, short setae; gula mixed with long, semidecumbent setae and with very long and protruding setae; frons slightly rugose medially; labrum with relative long, thick, flattened, protruding spines; segment II covered with sparse, semidecumbent setae, outer portion covered with very dense and short, apically curved setae, segment III sparsely covered with semidecumbent setae; segment IV covered with setae denser than those on segment III; rostrum thin, long, reaching abdomen apically.
<emphasis id="B9361A2AFFEDFF82FE0197D8FDDB50DE" bold="true" box="[477,567,1453,1479]" italics="true" pageId="27" pageNumber="28">Thorax</emphasis>
. Pronotum covered with sparse, fine setae; pronotal collar rugose, covered with fine setae; anterior lobe rather flattened, impunctate, slightly rugose, lateral margin gently carinate, with row of punctures, each bearing single, decumbent seta.
<emphasis id="B9361A2AFFEDFF82FCF8978DFB8E5308" box="[804,1122,1528,1553]" italics="true" pageId="27" pageNumber="28">Mesoscutum and scutellum</emphasis>
. Covered with short setae, somewhat rugose.
<emphasis id="B9361A2AFFEDFF82FEAB9468FDD5532F" box="[375,569,1565,1590]" italics="true" pageId="27" pageNumber="28">Thoracic pleura</emphasis>
. Propleuron impunctate anteriorly.
<emphasis id="B9361A2AFFEDFF82FC399468FB9C532F" box="[997,1136,1565,1590]" italics="true" pageId="27" pageNumber="28">Hemelytron</emphasis>
. Covered with relatively dense, short, decumbent, dark setae; embolium covered with setae longer than those on corium; cuneus covered with dense, relatively long setae.
<emphasis id="B9361A2AFFEDFF82FD5B941DFD525398" box="[647,702,1640,1665]" italics="true" pageId="27" pageNumber="28">Legs</emphasis>
. Femora and tibiae covered with rather short, semidecumbent setae.
<emphasis id="B9361A2AFFEDFF82FF3C94F8FEB853BE" bold="true" box="[224,340,1677,1703]" italics="true" pageId="27" pageNumber="28">Abdomen</emphasis>
. Covered with pale, reclining setae.
</paragraph>
<paragraph id="8BFDC638FFEDFF82FF1994C7FBBB520E" blockId="27.[151,1437,706,2002]" pageId="27" pageNumber="28">
<emphasis id="B9361A2AFFEDFF82FF1994C7FE9A53D5" bold="true" box="[197,374,1714,1740]" italics="true" pageId="27" pageNumber="28">Male genitalia</emphasis>
(
<figureCitation id="1379DABDFFEDFF82FE5494C7FE0D53D5" box="[392,481,1714,1740]" captionStart="FIGURE 13" captionStartId="28.[151,255,1939,1963]" captionTargetBox="[280,1311,886,1915]" captionTargetId="figure@28.[274,1312,869,1918]" captionTargetPageId="28" captionText="FIGURE 13. Male genitalia of Rhinocylapus scutatus: A, endosoma, B, left paramere, left lateral view; C, left paramere, dorsal view; D, right paramere. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/198894/files/figure.png" pageId="27" pageNumber="28">Fig. 13</figureCitation>
). Endosoma with two sclerites: one, smaller, situated beneath second gonopore, distinctly enlarged and folded proximally, remainder of this sclerite thin, sharply terminated; second sclerite very large, distinctly curved and twisted medially, its proximal portion bottle-like.
</paragraph>
<paragraph id="8BFDC638FFEDFF82FF199557FBCC527B" blockId="27.[151,1437,706,2002]" pageId="27" pageNumber="28">
<emphasis id="B9361A2AFFEDFF82FF199557FECA5225" bold="true" box="[197,294,1826,1852]" pageId="27" pageNumber="28">Female</emphasis>
. Similar to male but slightly larger. Labrum without spines; antennal segment II not bent, somewhat less stout, not covered with small setae, which are curved apically.
</paragraph>
<paragraph id="8BFDC638FFEDFF82FF199518FA7052CB" blockId="27.[151,1437,706,2002]" pageId="27" pageNumber="28">
<emphasis id="B9361A2AFFEDFF82FF199518FE68529E" bold="true" box="[197,388,1901,1927]" pageId="27" pageNumber="28">Measurements.</emphasis>
Ψ/ɗ: body length 8.8–9.5/7.4, width 2.5–2.75/2.2. Head. Length 1.75–2.0/1.5, width 1.1– 1.3/1.35, diameter of eye in dorsal view 0.3/0.3. Antenna. Length of segment I 1.55–1.65/1.3, II 2.9/2.15, III 5.1/4.0, IV 2.6/2.45. Labium. Length of segment I 3.2–3.3/2.55, II 1.8–1.85/1.75, III 2.2–2.55/2.0, IV 1.7/1.4.
</paragraph>
<paragraph id="8BFDC638FFEAFF85FF4B92EDFD8655CE" blockId="28.[151,1437,152,812]" pageId="28" pageNumber="29">Pronotum. Length 1.75–1.8/1.3. width of anterior margin 1.0/0.7–0.9, length of lateral margin 1.8–1.9/1.5, width posterior margin 1.7–1.8/2.2–2.3.</paragraph>
</subSubSection>
<subSubSection id="C35895B3FFEAFF85FF199297FE4055E5" box="[197,428,226,252]" pageId="28" pageNumber="29" type="biology_ecology">
<paragraph id="8BFDC638FFEAFF85FF199297FE4055E5" blockId="28.[151,1437,152,812]" box="[197,428,226,252]" pageId="28" pageNumber="29">
<emphasis id="B9361A2AFFEAFF85FF199297FEC555E5" bold="true" box="[197,297,226,252]" pageId="28" pageNumber="29">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35895B3FFEAFF85FF199372FCC8543B" box="[197,804,263,290]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8BFDC638FFEAFF85FF199372FCC8543B" blockId="28.[151,1437,152,812]" box="[197,804,263,290]" pageId="28" pageNumber="29">
<emphasis id="B9361A2AFFEAFF85FF199372FE895438" bold="true" box="[197,357,263,289]" pageId="28" pageNumber="29">Distribution.</emphasis>
<collectingCountry id="F35586A8FFEAFF85FEB0937DFE3B543B" box="[364,471,264,290]" name="Malaysia" pageId="28" pageNumber="29">Malaysia</collectingCountry>
(Borneo, Malay Peninsula).
</paragraph>
</subSubSection>
<subSubSection id="C35895B3FFEAFF85FF199358FEA3573E" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BFDC638FFEAFF85FF199358FA7A548B" blockId="28.[151,1437,152,812]" pageId="28" pageNumber="29">
<emphasis id="B9361A2AFFEAFF85FF199358FE90545E" bold="true" box="[197,380,301,327]" pageId="28" pageNumber="29">
<typeStatus id="54F9789AFFEAFF85FF199358FEE9545E" box="[197,261,301,327]" pageId="28" pageNumber="29">Type</typeStatus>
material
</emphasis>
:
<emphasis id="B9361A2AFFEAFF85FE529358FDE9545E" bold="true" box="[398,517,301,327]" pageId="28" pageNumber="29">
<typeStatus id="54F9789AFFEAFF85FE529358FDE9545E" box="[398,517,301,327]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
ɗ: Sandakan Borneo
<collectingCountry id="F35586A8FFEAFF85FCC99358FC8E545E" box="[789,866,301,327]" name="United States Pacific Island Wildlife Refuges" pageId="28" pageNumber="29">Baker</collectingCountry>
;
<typeStatus id="54F9789AFFEAFF85FCAC9358FC0E545E" box="[880,994,301,327]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C42BDBBFFEAFF85FC379358FA7D545E" authority="T. Y. Hsiao" authorityName="T. Y. Hsiao" box="[1003,1425,301,327]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B9361A2AFFEAFF85FC379358FAEC545F" box="[1003,1280,301,326]" italics="true" pageId="28" pageNumber="29">Rhinocylapus scutatus</emphasis>
T. Y. Hsiao
</taxonomicName>
;
<typeStatus id="54F9789AFFEAFF85FF4B9327FEE85475" box="[151,260,338,364]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
No. 56723
<collectingCountry id="F35586A8FFEAFF85FE509327FE525475" box="[396,446,338,364]" name="United States of America" pageId="28" pageNumber="29">U.S.</collectingCountry>
N.M.
<emphasis id="B9361A2AFFEAFF85FDD99327FD6D5475" bold="true" box="[517,641,338,364]" pageId="28" pageNumber="29">
<typeStatus id="54F9789AFFEAFF85FDD99327FD6D5475" box="[517,641,338,364]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
</emphasis>
:
<emphasis id="B9361A2AFFEAFF85FD4C9327FD725475" bold="true" box="[656,670,338,364]" pageId="28" pageNumber="29">1</emphasis>
Ψ: Sandakan Borneo
<collectingCountry id="F35586A8FFEAFF85FC4A9327FC0C5475" box="[918,992,338,364]" name="United States Pacific Island Wildlife Refuges" pageId="28" pageNumber="29">Baker</collectingCountry>
;
<typeStatus id="54F9789AFFEAFF85FC379327FBBE5475" box="[1003,1106,338,364]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
<taxonomicName id="4C42BDBBFFEAFF85FB849327FF33548B" authority="T. Y. Hsiao" authorityName="T. Y. Hsiao" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B9361A2AFFEAFF85FB849327FAB35472" box="[1112,1375,338,363]" italics="true" pageId="28" pageNumber="29">Rhinocylapus scutatus</emphasis>
T. Y. Hsiao
</taxonomicName>
;
<typeStatus id="54F9789AFFEAFF85FF36930DFEBD548B" box="[234,337,376,402]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
No. 56723
<collectingCountry id="F35586A8FFEAFF85FE07930DFDE2548B" box="[475,526,376,402]" name="United States of America" pageId="28" pageNumber="29">U.S.</collectingCountry>
N.M.;
<emphasis id="B9361A2AFFEAFF85FD819302FD805488" bold="true" box="[605,620,375,401]" pageId="28" pageNumber="29">1</emphasis>
ɗ: Sandakan Borneo
<collectingCountry id="F35586A8FFEAFF85FCB9930DFC43548B" box="[869,943,376,402]" name="United States Pacific Island Wildlife Refuges" pageId="28" pageNumber="29">Baker</collectingCountry>
;
<typeStatus id="54F9789AFFEAFF85FC67930DFBCD548B" box="[955,1057,376,402]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
No. 56723
<collectingCountry id="F35586A8FFEAFF85FB77930DFB31548B" box="[1195,1245,376,402]" name="United States of America" pageId="28" pageNumber="29">U.S.</collectingCountry>
N.M. (
<emphasis id="B9361A2AFFEAFF85FAF39302FA695488" bold="true" box="[1327,1413,375,401]" pageId="28" pageNumber="29">
<collectionCode id="ED535EFDFFEAFF85FAF39302FA695488" LSID="urn:lsid:biocol.org:col:34871" box="[1327,1413,375,401]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29">USNM</collectionCode>
</emphasis>
).
</paragraph>
<paragraph id="8BFDC638FFEAFF85FF1993E8FEA3573E" blockId="28.[151,1437,152,812]" pageId="28" pageNumber="29">
<emphasis id="B9361A2AFFEAFF85FF1993E8FDA254AE" bold="true" box="[197,590,413,439]" pageId="28" pageNumber="29">Additional examined material</emphasis>
.
<emphasis id="B9361A2AFFEAFF85FD8293E8FD8154AE" bold="true" box="[606,621,413,439]" pageId="28" pageNumber="29">1</emphasis>
Ψ: Sandakan Borneo
<collectingCountry id="F35586A8FFEAFF85FCAA93E8FC2F54AE" box="[886,963,413,439]" name="United States Pacific Island Wildlife Refuges" pageId="28" pageNumber="29">Baker</collectingCountry>
; Allotype
<taxonomicName id="4C42BDBBFFEAFF85FB9893E8FF3354C5" authority="T. Y. Hsiao" authorityName="T. Y. Hsiao" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B9361A2AFFEAFF85FB9893E8FAB554AF" box="[1092,1369,413,438]" italics="true" pageId="28" pageNumber="29">Rhinocylapus scutatus</emphasis>
T. Y. Hsiao
</taxonomicName>
; Allotype No. 56723
<collectingCountry id="F35586A8FFEAFF85FE0493B7FDE754C5" box="[472,523,450,476]" name="United States of America" pageId="28" pageNumber="29">U.S.</collectingCountry>
N.M.;
<emphasis id="B9361A2AFFEAFF85FD8593B7FD8454C5" bold="true" box="[601,616,450,476]" pageId="28" pageNumber="29">1</emphasis>
ɗ: N. Borneo, Samawang, nr. Sandakan, jungle, July, 1927; Exp. P. M. S. Museum. B. M. 1955-354; 1Ψ:
<collectingCountry id="F35586A8FFEAFF85FDF1939DFD7B571B" box="[557,663,488,514]" name="Malaysia" pageId="28" pageNumber="29">Sarawak</collectingCountry>
: Mt. Kalulong, 400–650 Pt. l.
<date id="FFFCE0F8FFEAFF85FBDD939DFB89571B" box="[1025,1125,488,514]" pageId="28" pageNumber="29" value="1932-11">xi. 1932</date>
; Oxford Univ. Exp. B. M, 1933-254 (
<collectingCountry id="F35586A8FFEAFF85FECB9078FEAD573E" box="[279,321,525,551]" name="United States of America" pageId="28" pageNumber="29">US</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C35895B3FFEAFF85FF199047FD195635" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BFDC638FFEAFF85FF199047FD195635" blockId="28.[151,1437,152,812]" pageId="28" pageNumber="29">
<emphasis id="B9361A2AFFEAFF85FF199047FEBD5755" bold="true" box="[197,337,562,588]" pageId="28" pageNumber="29">Discussion.</emphasis>
<bibRefCitation id="EFD3BBC9FFEAFF85FE859047FDCD5755" author="Gorczyca" box="[345,545,562,588]" pageId="28" pageNumber="29" refString="Gorczyca, J. (2006) The Catalogue of the subfamily Cylapinae Kirkaldy, 1903 of the World (Hemiptera, Heteroptera, Miridae). Monographs of the Upper Silesian Museum, 5, 1 - 100." type="journal article" year="2006">Gorczyca (2006)</bibRefCitation>
considered the data provided by
<bibRefCitation id="EFD3BBC9FFEAFF85FC719047FA705755" author="Kerzhner" box="[941,1436,562,588]" pageId="28" pageNumber="29" refString="Kerzhner, I. M. &amp; Konstantinov, F. V. (1999) Structure of the aedeagus in Miridae (Heteroptera) and its bearing to suprageneric classification. Acta Societatis Zoologicae Bohemicae, 63, 117 - 137." type="journal article" year="1999">
Kerzhner &amp; Konstantinov (1999,
<figureCitation id="1379DABDFFEAFF85FAE29047FA7E5755" box="[1342,1426,562,588]" captionStart="FIGURE 14" captionStartId="30.[151,258,1284,1308]" captionTargetBox="[212,1372,441,1276]" captionTargetId="figure@30.[210,1377,440,1278]" captionTargetPageId="30" captionText="FIGURE 14. Male genitalia of Rhinocylapus sumatranus: A, endosoma, B, left paramere, left lateral view; C, left paramere, dorsal view; D, right paramere. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/198895/files/figure.png" pageId="28" pageNumber="29">Fig. 14</figureCitation>
)
</bibRefCitation>
for
<taxonomicName id="4C42BDBBFFEAFF85FF1C902DFE925768" box="[192,382,600,625]" class="Insecta" family="Miridae" genus="Rhinocylapidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9361A2AFFEAFF85FF1C902DFE925768" box="[192,382,600,625]" italics="true" pageId="28" pageNumber="29">Rhinocylapidius</emphasis>
</taxonomicName>
sp. as referring to
<taxonomicName id="4C42BDBBFFEAFF85FD84902DFD0B5768" box="[600,743,600,626]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B9361A2AFFEAFF85FD84902DFD945768" box="[600,632,600,625]" italics="true" pageId="28" pageNumber="29">Rh</emphasis>
.
<emphasis id="B9361A2AFFEAFF85FD5B902DFD0B5768" box="[647,743,600,625]" italics="true" pageId="28" pageNumber="29">scutatus</emphasis>
</taxonomicName>
. However, the shape of the aedeagus of all known males of
<taxonomicName id="4C42BDBBFFEAFF85FF4B9008FEBA578F" box="[151,342,637,662]" class="Insecta" family="Miridae" genus="Rhinocylapidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9361A2AFFEAFF85FF4B9008FEBA578F" box="[151,342,637,662]" italics="true" pageId="28" pageNumber="29">Rhinocylapidius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C42BDBBFFEAFF85FE519008FDC0578F" box="[397,556,637,662]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9361A2AFFEAFF85FE519008FDC0578F" box="[397,556,637,662]" italics="true" pageId="28" pageNumber="29">Rhinocylapus</emphasis>
</taxonomicName>
(except for
<taxonomicName id="4C42BDBBFFEAFF85FD659008FC17578F" box="[697,1019,637,662]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="simplicicollis">
<emphasis id="B9361A2AFFEAFF85FD659008FC17578F" box="[697,1019,637,662]" italics="true" pageId="28" pageNumber="29">Rhinocylapus simplicicollis</emphasis>
</taxonomicName>
) is not consistent with that pictured by
<bibRefCitation id="EFD3BBC9FFEAFF85FF6790D7FDAC57A5" author="Kerzhner" box="[187,576,674,700]" pageId="28" pageNumber="29" refString="Kerzhner, I. M. &amp; Konstantinov, F. V. (1999) Structure of the aedeagus in Miridae (Heteroptera) and its bearing to suprageneric classification. Acta Societatis Zoologicae Bohemicae, 63, 117 - 137." type="journal article" year="1999">Kerzhner &amp; Konstantinov (1999)</bibRefCitation>
and the drawing most likely presents an undescribed taxon. Based on the single endosomal sclerite depicted in the drawing, which is unlike all known species of
<taxonomicName id="4C42BDBBFFEAFF85FB1890BDFA8457F8" box="[1220,1384,712,737]" class="Insecta" family="Miridae" genus="Rhinocylapus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9361A2AFFEAFF85FB1890BDFA8457F8" box="[1220,1384,712,737]" italics="true" pageId="28" pageNumber="29">Rhinocylapus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C42BDBBFFEAFF85FF4B9098FE8A561F" box="[151,358,749,774]" class="Insecta" family="Miridae" genus="Rhinocylapidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9361A2AFFEAFF85FF4B9098FE8A561F" box="[151,358,749,774]" italics="true" pageId="28" pageNumber="29">Rhinocylapidius</emphasis>
</taxonomicName>
where the endosoma is complex, with numerous sclerites and sclerotized lobes, the illustration may even present an undescribed genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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