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<document id="BE47648E7E44E45664FA68632CAA3E4C" ID-DOI="10.11646/zootaxa.3731.4.1" ID-GBIF-Dataset="fc05e511-82bb-4513-addd-3568e5c018f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="223206" ID-ZooBank="A785056E-CE4A-40CB-B2AA-8BCD869AD10A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460694450374" checkinUser="plazi" docAuthor="Serrano, Artur R. M. &amp; Capela, Rúben A." docDate="2013" docId="03D887FBFC77FFD2FF4F71977267F938" docLanguage="en" docName="zt03731p444.pdf" docOrigin="Zootaxa 3731 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Habrodera Motschulsky 1862" docType="treatment" docVersion="10" lastPageNumber="435" masterDocId="FFE1FF83FC56FFF0FFD8777B7366FFC2" masterDocTitle="The tiger beetles (Coleoptera: Carabidae, Cicindelinae) of Angola: a descriptive catalogue and designation of neotypes" masterLastPageNumber="444" masterPageNumber="401" pageNumber="434" updateTime="1698353693208" updateUser="plazi">
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<mods:title id="C89CE95C6FB26CB72C0E787474E9C310">The tiger beetles (Coleoptera: Carabidae, Cicindelinae) of Angola: a descriptive catalogue and designation of neotypes</mods:title>
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<mods:namePart id="5A6DBD1554FC06EB56DA02E8444E725D">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="46E601D9D55521734EA50D1F3C93C503">Capela, Rúben A.</mods:namePart>
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<mods:date id="244B918E978E79D55AD50377C796B664">2013</mods:date>
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<treatment id="03D887FBFC77FFD2FF4F71977267F938" ID-DOI="http://doi.org/10.5281/zenodo.5694350" ID-GBIF-Taxon="119581653" ID-Zenodo-Dep="5694350" LSID="urn:lsid:plazi:treatment:03D887FBFC77FFD2FF4F71977267F938" httpUri="http://treatment.plazi.org/id/03D887FBFC77FFD2FF4F71977267F938" lastPageId="34" lastPageNumber="435" pageId="33" pageNumber="434">
<subSubSection id="C36B6566FC77FFD2FF4F7197718FFF72" lastPageId="34" lastPageNumber="435" pageId="33" pageNumber="434" type="nomenclature">
<paragraph id="8BCE36EDFC77FFD1FF4F71977041F8C4" blockId="33.[151,807,1772,1798]" box="[151,807,1772,1798]" pageId="33" pageNumber="434">
<heading id="D0868181FC77FFD1FF4F71977041F8C4" bold="true" box="[151,807,1772,1798]" fontSize="11" level="1" pageId="33" pageNumber="434" reason="1">
<emphasis id="B905EAFFFC77FFD1FF4F71977041F8C4" bold="true" box="[151,807,1772,1798]" pageId="33" pageNumber="434">
<taxonomicName id="4C714D6EFC77FFD1FF4F7197716EF8C4" ID-CoL="8KX4X" authority="Motschulsky, 1862" authorityName="Motschulsky" authorityYear="1862" box="[151,520,1772,1798]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B905EAFFFC77FFD1FF4F7197727DF8C4" bold="true" box="[151,283,1772,1798]" italics="true" pageId="33" pageNumber="434">Habrodera</emphasis>
Motschulsky, 1862
</taxonomicName>
(
<emphasis id="B905EAFFFC77FFD1FDC2719771A0F8C4" bold="true" box="[538,710,1772,1798]" italics="true" pageId="33" pageNumber="434">Etud. entomol</emphasis>
. 11: 22)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36EDFC77FFD1FF4F704870B6F8AD" blockId="33.[151,1437,1842,1903]" pageId="33" pageNumber="434">
An African genus with five recognised valid species (Jaskula &amp; Cassola 2005). Three of them have been in
<collectingCountry id="F366767DFC77FFD1FA9B704976FFF889" box="[1347,1433,1842,1867]" name="Angola" pageId="33" pageNumber="434">Angola</collectingCountry>
. All species of this genus occur on sand beaches of oceans, rivers or lakes.
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F77EC718FFF72" blockId="34.[151,1436,151,717]" box="[151,745,151,176]" pageId="34" pageNumber="435">
<taxonomicName id="4C714D6EFC74FFD2FF4F77EC717EFF72" authority="Bates, 1878" authorityName="Bates" authorityYear="1878" box="[151,536,151,176]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="cabinda">
<emphasis id="B905EAFFFC74FFD2FF4F77EC717EFF72" bold="true" box="[151,536,151,176]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F77EC721EFF72" bold="true" box="[151,376,151,176]" italics="true" pageId="34" pageNumber="435">Habrodera cabinda</emphasis>
(Bates, 1878)
</emphasis>
</taxonomicName>
(
<emphasis id="B905EAFFFC74FFD2FDF177E371EDFF72" box="[553,651,152,176]" italics="true" pageId="34" pageNumber="435">Cist. Ent</emphasis>
.
<emphasis id="B905EAFFFC74FFD2FD4177EC71C1FF72" bold="true" box="[665,679,151,176]" pageId="34" pageNumber="435">2</emphasis>
: 331)
</paragraph>
</subSubSection>
<subSubSection id="C36B6566FC74FFD2FF4F77C777F1FE4A" pageId="34" pageNumber="435" type="distribution">
<paragraph id="8BCE36EDFC74FFD2FF4F77C772C5FF17" blockId="34.[151,1436,151,717]" box="[151,419,188,213]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F77C772C5FF17" bold="true" box="[151,419,188,213]" pageId="34" pageNumber="435">
Distribution in
<collectingCountry id="F366767DFC74FFD2FE9677C772C5FF17" box="[334,419,188,213]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F77A47248FF3A" blockId="34.[151,1436,151,717]" box="[151,302,223,248]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F77A47248FF3A" bold="true" box="[151,302,223,248]" pageId="34" pageNumber="435">Old records:</emphasis>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F767E70A5FEDE" blockId="34.[151,1436,151,717]" box="[151,963,260,285]" pageId="34" pageNumber="435">
—
<typeStatus id="54CA884FFC74FFD2FF6B767E738DFEDF" box="[179,235,261,285]" pageId="34" pageNumber="435">Type</typeStatus>
locality: Landana, Loango (U. More), several ♂ and ♀, BNHM?
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F765C7766FEA7" blockId="34.[151,1436,151,717]" pageId="34" pageNumber="435">
—Landana (U. More) (
<taxonomicName id="4C714D6EFC74FFD2FE7676537157FE82" box="[430,561,296,320]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FE7676537157FE82" box="[430,561,296,320]" name="Angola" pageId="34" pageNumber="435">CABINDA</collectingCountry>
</taxonomicName>
) (5), Luango (U. More) (
<taxonomicName id="4C714D6EFC74FFD2FCB476537089FE82" box="[876,1007,296,320]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FCB476537089FE82" box="[876,1007,296,320]" name="Angola" pageId="34" pageNumber="435">CABINDA</collectingCountry>
</taxonomicName>
) (5), Lobito (BENGUELA) (227/8) (Wellman &amp; Horn 1908 sub
<taxonomicName id="4C714D6EFC74FFD2FDDE763771B2FEA7" box="[518,724,332,357]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="cabinda">
<emphasis id="B905EAFFFC74FFD2FDDE763771B2FEA7" box="[518,724,332,357]" italics="true" pageId="34" pageNumber="435">Cicindela cabinda</emphasis>
</taxonomicName>
; Jaskula &amp; Cassola 2005).
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F761477F1FE4A" blockId="34.[151,1436,151,717]" box="[151,1175,367,392]" pageId="34" pageNumber="435">
—
<taxonomicName id="4C714D6EFC74FFD2FF6C76147275FE4A" box="[180,275,367,392]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FF6C76147275FE4A" box="[180,275,367,392]" name="Angola" pageId="34" pageNumber="435">Cabinda</collectingCountry>
</taxonomicName>
, without locality (
<taxonomicName id="4C714D6EFC74FFD2FE05760B7106FE4A" box="[477,608,368,392]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FE05760B7106FE4A" box="[477,608,368,392]" name="Angola" pageId="34" pageNumber="435">CABINDA</collectingCountry>
</taxonomicName>
) (Ferreira 1965 sub
<taxonomicName id="4C714D6EFC74FFD2FC9C761477EDFE4A" authority="Dejean" authorityName="Dejean" box="[836,1163,367,392]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="leucoptera">
<emphasis id="B905EAFFFC74FFD2FC9C76147756FE4A" box="[836,1072,367,392]" italics="true" pageId="34" pageNumber="435">Cicindela leucoptera</emphasis>
Dejean
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B6566FC74FFD2FF4F76EF71D5FE37" pageId="34" pageNumber="435" type="description">
<paragraph id="8BCE36EDFC74FFD2FF4F76EF71D5FE37" blockId="34.[151,1436,151,717]" pageId="34" pageNumber="435">
—
<taxonomicName id="4C714D6EFC74FFD2FF6C76EF7275FE6F" box="[180,275,404,429]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FF6C76EF7275FE6F" box="[180,275,404,429]" name="Angola" pageId="34" pageNumber="435">Cabinda</collectingCountry>
</taxonomicName>
, without locality (
<taxonomicName id="4C714D6EFC74FFD2FE3476EF7117FE6E" box="[492,625,404,428]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FE3476EF7117FE6E" box="[492,625,404,428]" name="Angola" pageId="34" pageNumber="435">CABINDA</collectingCountry>
</taxonomicName>
), Malongo (not Benguella) (
<taxonomicName id="4C714D6EFC74FFD2FC1976EF7722FE6E" box="[961,1092,404,428]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FC1976EF7722FE6E" box="[961,1092,404,428]" name="Angola" pageId="34" pageNumber="435">CABINDA</collectingCountry>
</taxonomicName>
) (5),
<collectingCountry id="F366767DFC74FFD2FB5376EF77D1FE6F" box="[1163,1207,404,429]" name="Iceland" pageId="34" pageNumber="435">Ilha</collectingCountry>
do Cabo (=Ilha de
<collectingCountry id="F366767DFC74FFD2FF1F76CC727BFE12" box="[199,285,439,464]" name="Rwanda" pageId="34" pageNumber="435">Luanda</collectingCountry>
) (
<collectingCountry id="F366767DFC74FFD2FEEE76C372D6FE12" box="[310,432,440,464]" name="Rwanda" pageId="34" pageNumber="435">LUANDA</collectingCountry>
) (89),
<collectingCountry id="F366767DFC74FFD2FDDB76CC7131FE12" box="[515,599,439,464]" name="Rwanda" pageId="34" pageNumber="435">Luanda</collectingCountry>
—Kwanza River (
<quantity id="4C899B08FC74FFD2FCEE76CC70E4FE12" box="[822,898,439,464]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="435" unit="km" value="25.0">25 km</quantity>
south
<collectingCountry id="F366767DFC74FFD2FC0D76CC774AFE12" box="[981,1068,439,464]" name="Rwanda" pageId="34" pageNumber="435">Luanda</collectingCountry>
, probably near Morro da Cruz) (
<collectingCountry id="F366767DFC74FFD2FF1676A7722EFE36" box="[206,328,476,500]" name="Rwanda" pageId="34" pageNumber="435">LUANDA</collectingCountry>
) (89) (Jaskula &amp; Cassola 2005).
</paragraph>
</subSubSection>
<subSubSection id="C36B6566FC74FFD2FF1F768470E5FCEC" pageId="34" pageNumber="435" type="discussion">
<paragraph id="8BCE36EDFC74FFD2FF1F768476FAFDDA" blockId="34.[151,1436,151,717]" box="[199,1436,511,536]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF1F7684725DFDDA" bold="true" box="[199,315,511,536]" pageId="34" pageNumber="435">Remarks.</emphasis>
A tiger beetle species with a distribution throughout the seashores of
<collectingCountry id="F366767DFC74FFD2FB9A768477F3FDDA" box="[1090,1173,511,536]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
and D. R. of the
<collectingCountry id="F366767DFC74FFD2FA89757B76FAFDDA" box="[1361,1436,512,536]" name="Republic of the Congo" pageId="34" pageNumber="435">Congo</collectingCountry>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F755F76FAFDFF" blockId="34.[151,1436,151,717]" box="[151,1436,548,573]" pageId="34" pageNumber="435">
(Werner 2000b).
<taxonomicName id="4C714D6EFC74FFD2FE8E755F7154FDFF" box="[342,562,548,573]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="cabinda">
<emphasis id="B905EAFFFC74FFD2FE8E755F7154FDFF" box="[342,562,548,573]" italics="true" pageId="34" pageNumber="435">Habrodera cabinda</emphasis>
</taxonomicName>
was described from
<collectingCountry id="F366767DFC74FFD2FCC3755F7008FDFF" box="[795,878,548,573]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
and it was latter synonymized with
<taxonomicName id="4C714D6EFC74FFD2FADA755E76FAFDFF" box="[1282,1436,548,573]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="leucoptera">
<emphasis id="B905EAFFFC74FFD2FADA755E76FAFDFF" box="[1282,1436,548,573]" italics="true" pageId="34" pageNumber="435">H. leucoptera</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F753C76FAFDA2" blockId="34.[151,1436,151,717]" box="[151,1436,583,608]" pageId="34" pageNumber="435">
by Fleutiaux (1892). Werner (2000b) named it as a subspecies of
<taxonomicName id="4C714D6EFC74FFD2FC5175327743FDA2" box="[905,1061,583,608]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="leucoptera">
<emphasis id="B905EAFFFC74FFD2FC5175327743FDA2" box="[905,1061,583,608]" italics="true" pageId="34" pageNumber="435">H. leucoptera</emphasis>
</taxonomicName>
, but it was elevated again to the
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F751776FAFD47" blockId="34.[151,1436,151,717]" box="[151,1436,620,645]" pageId="34" pageNumber="435">
level of full species by Jaskula &amp; Cassola (2005). Wellman &amp; Horn (1908) noticed that this species and
<taxonomicName id="4C714D6EFC74FFD2FAFB751676FAFD47" box="[1315,1436,620,645]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="B905EAFFFC74FFD2FAFB751676FAFD47" box="[1315,1436,620,645]" italics="true" pageId="34" pageNumber="435">H. nitidula</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F75EB76FAFD6A" blockId="34.[151,1436,151,717]" box="[151,1436,655,680]" pageId="34" pageNumber="435">are sympatric and were “taken on the beach (that part which is kept more or less wet by the tides)” and by its color</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F75CE7784FD0F" blockId="34.[151,1436,151,717]" box="[151,1250,692,717]" pageId="34" pageNumber="435">are “rendered inconspicuous as it sits on the sand”, and are “usually noticeable only when flying”.</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F746E70E5FCEC" blockId="34.[151,899,789,814]" box="[151,899,789,814]" pageId="34" pageNumber="435">
<taxonomicName id="4C714D6EFC74FFD2FF4F746E71E3FCEC" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" box="[151,645,789,814]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="subSpecies" species="nilotica" subSpecies="nilotica">
<emphasis id="B905EAFFFC74FFD2FF4F746E71E3FCEC" bold="true" box="[151,645,789,814]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F746E72B7FCEC" bold="true" box="[151,465,789,814]" italics="true" pageId="34" pageNumber="435">Habrodera nilotica nilotica</emphasis>
(Dejean, 1825)
</emphasis>
</taxonomicName>
(
<emphasis id="B905EAFFFC74FFD2FD4D746D704FFCEC" box="[661,809,789,814]" italics="true" pageId="34" pageNumber="435">Spec. Coleop</emphasis>
.
<emphasis id="B905EAFFFC74FFD2FCEF746E7024FCEC" bold="true" box="[823,834,789,814]" pageId="34" pageNumber="435">I</emphasis>
: 119)
</paragraph>
</subSubSection>
<subSubSection id="C36B6566FC74FFD2FF4F74267293FC20" pageId="34" pageNumber="435" type="distribution">
<paragraph id="8BCE36EDFC74FFD2FF4F742672C5FCB4" blockId="34.[151,1436,861,1174]" box="[151,419,861,886]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F742672C5FCB4" bold="true" box="[151,419,861,886]" pageId="34" pageNumber="435">
Distribution in
<collectingCountry id="F366767DFC74FFD2FE96742672C5FCB4" box="[334,419,861,886]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F74FA7248FC58" blockId="34.[151,1436,861,1174]" box="[151,302,897,922]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F74FA7248FC58" bold="true" box="[151,302,897,922]" pageId="34" pageNumber="435">Old records:</emphasis>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F74DE76FAFC7C" blockId="34.[151,1436,861,1174]" box="[151,1436,933,958]" pageId="34" pageNumber="435">
—Humbi (fronteira
<collectingCountry id="F366767DFC74FFD2FEA074DE72ACFC7C" box="[376,458,933,958]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
/
<collectingCountry id="F366767DFC74FFD2FE0B74DE7153FC7C" box="[467,565,933,958]" name="Namibia" pageId="34" pageNumber="435">Namíbia</collectingCountry>
) (=Humbe) (CUNENE) (422) (Horn 1935; Monard
<emphasis id="B905EAFFFC74FFD2FBA674DD77D6FC7C" box="[1150,1200,933,958]" italics="true" pageId="34" pageNumber="435">et al</emphasis>
. 1956; Ferreira 1965
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF1374B27293FC20" blockId="34.[151,1436,861,1174]" box="[203,501,969,994]" pageId="34" pageNumber="435">
all sub
<taxonomicName id="4C714D6EFC74FFD2FEC474B27283FC20" box="[284,485,969,994]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis id="B905EAFFFC74FFD2FEC474B27283FC20" box="[284,485,969,994]" italics="true" pageId="34" pageNumber="435">Cicindela nilotica</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B6566FC74FFD2FF1F749670ECFAC0" pageId="34" pageNumber="435" type="discussion">
<paragraph id="8BCE36EDFC74FFD2FF1F74967111FB54" blockId="34.[151,1436,861,1174]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF1F7496725DFBC4" bold="true" box="[199,315,1005,1030]" pageId="34" pageNumber="435">Remarks.</emphasis>
A species present throughout the African countries south of the Sahel (Werner 2000b). It occupies the shores and banks of rivers and lakes which conforms well to the Humbe locality near the Cunene River in
<collectingCountry id="F366767DFC74FFD2FF4F734E738BFB8C" box="[151,237,1077,1102]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
. Werner (2000b) in spite of the published records of this species in
<collectingCountry id="F366767DFC74FFD2FBC8734E7703FB8C" box="[1040,1125,1077,1102]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
, did not include it for this country. Unlike the majority of the Afrotropical tiger beetles, adults of
<taxonomicName id="4C714D6EFC74FFD2FC097321772BFBB0" box="[977,1101,1113,1138]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="B905EAFFFC74FFD2FC097321772BFBB0" box="[977,1101,1113,1138]" italics="true" pageId="34" pageNumber="435">H. nitidula</emphasis>
</taxonomicName>
are independent of the rainy seasons and are active throughout the year.
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F739270ECFAC0" blockId="34.[151,906,1257,1282]" box="[151,906,1257,1282]" pageId="34" pageNumber="435">
<taxonomicName id="4C714D6EFC74FFD2FF4F739271EDFAC0" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" box="[151,651,1257,1282]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="subSpecies" species="nitidula" subSpecies="nitidula">
<emphasis id="B905EAFFFC74FFD2FF4F739271EDFAC0" bold="true" box="[151,651,1257,1282]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F739272B1FAC0" bold="true" box="[151,471,1257,1282]" italics="true" pageId="34" pageNumber="435">Habrodera nitidula nitidula</emphasis>
(Dejean, 1825)
</emphasis>
</taxonomicName>
(
<emphasis id="B905EAFFFC74FFD2FD4373917049FAC0" box="[667,815,1257,1282]" italics="true" pageId="34" pageNumber="435">Spec. Coleop</emphasis>
.
<emphasis id="B905EAFFFC74FFD2FCE57392702EFAC0" bold="true" box="[829,840,1257,1282]" pageId="34" pageNumber="435">I</emphasis>
: 120)
</paragraph>
</subSubSection>
<subSubSection id="C36B6566FC74FFD2FF4F724A71DBFA3C" pageId="34" pageNumber="435" type="distribution">
<paragraph id="8BCE36EDFC74FFD2FF4F724A72C5FA88" blockId="34.[151,1436,1329,1786]" box="[151,419,1329,1354]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F724A72C5FA88" bold="true" box="[151,419,1329,1354]" pageId="34" pageNumber="435">
Distribution in
<collectingCountry id="F366767DFC74FFD2FE96724A72C5FA88" box="[334,419,1329,1354]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F722E7248FAAC" blockId="34.[151,1436,1329,1786]" box="[151,302,1365,1390]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF4F722E7248FAAC" bold="true" box="[151,302,1365,1390]" pageId="34" pageNumber="435">Old records:</emphasis>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F7202702DFA50" blockId="34.[151,1436,1329,1786]" box="[151,843,1401,1426]" pageId="34" pageNumber="435">
—
<collectingCountry id="F366767DFC74FFD2FF6B72027263FA50" box="[179,261,1401,1426]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
(Welwitsch) (Putzeys 1880 sub
<taxonomicName id="4C714D6EFC74FFD2FDA87202705DFA50" box="[624,827,1401,1426]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="B905EAFFFC74FFD2FDA87202705DFA50" box="[624,827,1401,1426]" italics="true" pageId="34" pageNumber="435">Cicindela nitidula</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F72E67661FA18" blockId="34.[151,1436,1329,1786]" pageId="34" pageNumber="435">
—
<collectingCountry id="F366767DFC74FFD2FF6B72E67263FA74" box="[179,261,1437,1462]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
(Welwitsch), Landana (U. More) (
<taxonomicName id="4C714D6EFC74FFD2FD6E72E5705FFA74" box="[694,825,1438,1462]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FD6E72E5705FFA74" box="[694,825,1438,1462]" name="Angola" pageId="34" pageNumber="435">CABINDA</collectingCountry>
</taxonomicName>
) (5), Luango (U. More) (
<taxonomicName id="4C714D6EFC74FFD2FBA772E57664FA74" box="[1151,1282,1438,1462]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="441" phylum="Arthropoda" rank="species" species="cabinda">
<collectingCountry id="F366767DFC74FFD2FBA772E57664FA74" box="[1151,1282,1438,1462]" name="Angola" pageId="34" pageNumber="435">CABINDA</collectingCountry>
</taxonomicName>
) (5), Lobito (BENGUELA) (227/8) (Wellman &amp; Horn 1908 sub
<taxonomicName id="4C714D6EFC74FFD2FCC872BA70BDFA18" box="[784,987,1473,1498]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="B905EAFFFC74FFD2FCC872BA70BDFA18" box="[784,987,1473,1498]" italics="true" pageId="34" pageNumber="435">Cicindela nitidula</emphasis>
</taxonomicName>
; Jaskula &amp; Cassola 2005).
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F729E71DBFA3C" blockId="34.[151,1436,1329,1786]" box="[151,701,1509,1534]" pageId="34" pageNumber="435">
—
<collectingCountry id="F366767DFC74FFD2FF6B729E7263FA3C" box="[179,261,1509,1534]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
(Ferreira 1965 sub
<taxonomicName id="4C714D6EFC74FFD2FE3A729E71CBFA3C" box="[482,685,1509,1534]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="B905EAFFFC74FFD2FE3A729E71CBFA3C" box="[482,685,1509,1534]" italics="true" pageId="34" pageNumber="435">Cicindela nitidula</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B6566FC74FFD2FF1F71727267F938" pageId="34" pageNumber="435" type="materials_examined">
<paragraph id="8BCE36EDFC74FFD2FF1F717276FDF984" blockId="34.[151,1436,1329,1786]" pageId="34" pageNumber="435">
<emphasis id="B905EAFFFC74FFD2FF1F717272D4F9E0" bold="true" box="[199,434,1545,1570]" pageId="34" pageNumber="435">Material examined:</emphasis>
<collectingCountry id="F366767DFC74FFD2FE617172716AF9E0" box="[441,524,1545,1570]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
(Welwitsch),
<specimenCount id="9D77FD64FC74FFD2FD71717171B3F9E3" box="[681,725,1546,1570]" pageId="34" pageNumber="435" type="generic">2 ♂</specimenCount>
, 1 ♀, reg. nº 1907, “Old Collection” MZUC.
<emphasis id="B905EAFFFC74FFD2FF1F7156725DF984" bold="true" box="[199,315,1581,1606]" pageId="34" pageNumber="435">Remarks.</emphasis>
A tiger beetle species with a distribution throughout the sandy seashores of the western African
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F712A76FAF9AB" blockId="34.[151,1436,1329,1786]" box="[151,1436,1617,1642]" pageId="34" pageNumber="435">
countries (Jaskula &amp; Cassola, 2005; Serrano, 2007). In spite of the several records given to
<collectingCountry id="F366767DFC74FFD2FB3D712A765CF9A8" box="[1253,1338,1617,1642]" name="Angola" pageId="34" pageNumber="435">Angola</collectingCountry>
, Werner
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F710E76FAF94C" blockId="34.[151,1436,1329,1786]" box="[151,1436,1653,1678]" pageId="34" pageNumber="435">(2000b) questioned its presence in this country. The three specimens examined here (MZUC) are much damaged</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F71E276FAF970" blockId="34.[151,1436,1329,1786]" box="[151,1436,1689,1714]" pageId="34" pageNumber="435">
and were erroneously identified as
<taxonomicName id="4C714D6EFC74FFD2FDEE71E27001F970" authority="Dejean." authorityName="Dejean." box="[566,871,1689,1714]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis id="B905EAFFFC74FFD2FDEE71E27062F970" box="[566,772,1689,1714]" italics="true" pageId="34" pageNumber="435">Cicindela nilotica</emphasis>
Dejean.
</taxonomicName>
<taxonomicName id="4C714D6EFC74FFD2FCAA71E27736F970" box="[882,1104,1689,1714]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="B905EAFFFC74FFD2FCAA71E27736F970" box="[882,1104,1689,1714]" italics="true" pageId="34" pageNumber="435">Habrodera nitidula</emphasis>
</taxonomicName>
is a sandy seashore species
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F71C676FAF917" blockId="34.[151,1436,1329,1786]" box="[151,1436,1725,1750]" pageId="34" pageNumber="435">
occurring simpatricaly with
<taxonomicName id="4C714D6EFC74FFD2FE0071C67069F914" authority="Bates" authorityName="Bates" box="[472,783,1725,1750]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="cabinda">
<emphasis id="B905EAFFFC74FFD2FE0071C671D0F914" box="[472,694,1725,1750]" italics="true" pageId="34" pageNumber="435">Habrodera cabinda</emphasis>
(Bates)
</taxonomicName>
in some Angolan localities (see above distribution on
<emphasis id="B905EAFFFC74FFD2FA5971C576FAF917" box="[1409,1436,1726,1749]" italics="true" pageId="34" pageNumber="435">H.</emphasis>
</paragraph>
<paragraph id="8BCE36EDFC74FFD2FF4F719A7267F938" blockId="34.[151,1436,1329,1786]" box="[151,257,1761,1786]" pageId="34" pageNumber="435">
<taxonomicName id="4C714D6EFC74FFD2FF4F719A7397F938" box="[151,241,1761,1786]" class="Insecta" family="Carabidae" genus="Habrodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="cabinda">
<emphasis id="B905EAFFFC74FFD2FF4F719A7397F938" box="[151,241,1761,1786]" italics="true" pageId="34" pageNumber="435">cabinda</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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