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<document id="4F220BDFE43904880A1889BDB45FBEA6" ID-CLB-Dataset="46468" ID-DOI="10.11646/zootaxa.3764.4.3" ID-GBIF-Dataset="9dc5d2f3-c7e3-425b-9f51-0100b5bd2455" ID-ISSN="1175-5326" ID-Zenodo-Dep="230623" ID-ZooBank="04036FAE-6FF1-4ADB-AA0F-4C60DBF55BCE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460382799073" checkinUser="plazi" docAuthor="Chan, Kin Onn, Wood Jr, Perry L., Anuar, Shahrul, Muin, Mohd Abdul, Quah, Evan S. H., Sumarli, Alexandra X. Y. &amp; Grismer, L. Lee" docDate="2014" docId="03D887A5FFE0A067CB8DFA54FEE53837" docLanguage="en" docName="zt03764p440.pdf" docOrigin="Zootaxa 3764 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ansonia lumut Chan, Jr, Anuar, Muin, Quah, Sumarli &amp; Grismer, 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="437" masterDocId="FFE1FFDDFFE4A06DCB1AFFBAFF9B3E3F" masterDocTitle="A new species of upland Stream Toad of the genus Ansonia Stoliczka, 1870 (Anura: Bufonidae) from northeastern Peninsular Malaysia" masterLastPageNumber="440" masterPageNumber="427" pageNumber="431" updateTime="1698297051380" updateUser="ExternalLinkService">
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<mods:title id="1DAE57E4BB615C996E85EF955B24138B">A new species of upland Stream Toad of the genus Ansonia Stoliczka, 1870 (Anura: Bufonidae) from northeastern Peninsular Malaysia</mods:title>
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<mods:roleTerm id="7825B7A53ECD254BA92035DC53FF36CC">Author</mods:roleTerm>
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<mods:namePart id="30BC81B0D03C3323B09DF79E75FCE15F">Chan, Kin Onn</mods:namePart>
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<mods:namePart id="0FD5808D50D2CA2189A7F336C30C363A">Wood Jr, Perry L.</mods:namePart>
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<mods:name id="D3383454578D02D141B9990F72A1D293" type="personal">
<mods:role id="B6E465BDA02C9FFA57B438F65D131CB3">
<mods:roleTerm id="3C5BEBA2E7B8D4C9D0E69C8C51E7FBF3">Author</mods:roleTerm>
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<mods:namePart id="21E0F0A99F28D940B05E2C5CF191EA20">Anuar, Shahrul</mods:namePart>
</mods:name>
<mods:name id="96CAE47D17D6A493881D1D34A12BAEBA" type="personal">
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<mods:roleTerm id="5B50DA7C1DB5FD6D8E8485B68BA4CDB5">Author</mods:roleTerm>
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<mods:namePart id="B7135A6C5B0C9BAEC31587AC40C4CA3A">Muin, Mohd Abdul</mods:namePart>
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<mods:roleTerm id="44E8F81B3B3109AE577957F11F1DA1E3">Author</mods:roleTerm>
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<mods:namePart id="4D8FC40389483902FBF65D0EAFA86DD3">Quah, Evan S. H.</mods:namePart>
</mods:name>
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<mods:role id="ABFD4D63C50FB2A897C20E7352D65378">
<mods:roleTerm id="F048FFB5BB43D37EA9C83F9DEF850909">Author</mods:roleTerm>
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<mods:namePart id="2AF273F997C647198A399168D9CEB45D">Sumarli, Alexandra X. Y.</mods:namePart>
</mods:name>
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<mods:role id="3B95383A81A7A055A961751500349CBE">
<mods:roleTerm id="B8114BF24F4E79E5A8B44108C4C4DE3F">Author</mods:roleTerm>
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<mods:namePart id="CAFC64C2BEBCE4FB1060715201256C75">Grismer, L. Lee</mods:namePart>
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<mods:title id="A31DE9533ED9EEBDC770CCC6C4C1A3DF">Zootaxa</mods:title>
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<mods:date id="CD52EDC44A53E20124FE45C9066980C6">2014</mods:date>
<mods:detail id="9BF3D50B86C3F4F2ACA0B791FF1057A7" type="volume">
<mods:number id="BB1A02B2FC69CA5FC72DD3D204A3208C">3764</mods:number>
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<treatment id="03D887A5FFE0A067CB8DFA54FEE53837" ID-DOI="http://doi.org/10.5281/zenodo.5629443" ID-GBIF-Taxon="119414583" ID-Zenodo-Dep="5629443" LSID="urn:lsid:plazi:treatment:03D887A5FFE0A067CB8DFA54FEE53837" httpUri="http://treatment.plazi.org/id/03D887A5FFE0A067CB8DFA54FEE53837" lastPageId="10" lastPageNumber="437" pageId="4" pageNumber="431">
<subSubSection id="C36B6538FFE0A069CB8DFA54FF423814" pageId="4" pageNumber="431" type="nomenclature">
<paragraph id="8BCE36B3FFE0A069CB8DFA54FE383838" blockId="4.[151,419,1518,1579]" box="[151,419,1518,1543]" pageId="4" pageNumber="431">
<heading id="D08681DFFFE0A069CB8DFA54FE383838" bold="true" box="[151,419,1518,1543]" fontSize="10" level="2" pageId="4" pageNumber="431" reason="2">
<emphasis id="B905EAA1FFE0A069CB8DFA54FE383838" bold="true" box="[151,419,1518,1543]" pageId="4" pageNumber="431">
<taxonomicName id="4C714D30FFE0A069CB8DFA54FED93838" ID-CoL="66Y8L" box="[151,322,1518,1543]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="4" pageNumber="431" phylum="Chordata" rank="species" species="lumut" status="sp. nov.">
<emphasis id="B905EAA1FFE0A069CB8DFA54FED93838" bold="true" box="[151,322,1518,1543]" italics="true" pageId="4" pageNumber="431">Ansonia lumut</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657DAFFE0A069CA53FA54FE383838" box="[329,419,1518,1543]" pageId="4" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36B3FFE0A069CB8DF9A8FF423814" blockId="4.[151,419,1518,1579]" box="[151,217,1554,1579]" pageId="4" pageNumber="431">
<figureCitation id="134A2A36FFE0A069CB8DF9A8FF423814" box="[151,217,1554,1579]" captionStart="FIGURE 2" captionStartId="7.[151,250,1068,1090]" captionTargetBox="[151,1436,193,1045]" captionTargetId="figure@7.[151,1436,193,1046]" captionTargetPageId="7" captionText="FIGURE 2. Live coloration of female paratype LSUHC 11211." httpUri="https://zenodo.org/record/230625/files/figure.png" pageId="4" pageNumber="431">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B6538FFE0A069CB8DF9E1FE60384F" box="[151,507,1626,1648]" pageId="4" pageNumber="431" type="reference_group">
<paragraph id="8BCE36B3FFE0A069CB8DF9E1FE60384F" blockId="4.[151,507,1626,1648]" box="[151,507,1626,1648]" pageId="4" pageNumber="431">
<treatmentCitationGroup id="AB61119DFFE0A069CB8DF9E1FE60384F" box="[151,507,1626,1648]" pageId="4" pageNumber="431">
<taxonomicName id="4C714D30FFE0A069CB8DF9E1FE60384F" ID-CoL="679WL" authority="Dring 1979:184" authorityName="Dring" authorityPageNumber="184" authorityYear="1979" box="[151,507,1626,1648]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="4" pageNumber="431" phylum="Chordata" rank="species" species="malayana">
<emphasis id="B905EAA1FFE0A069CB8DF9E1FECF384F" box="[151,340,1627,1648]" italics="true" pageId="4" pageNumber="431">Ansonia malayana</emphasis>
<treatmentCitation id="0AD010A2FFE0A069CA40F9E0FE60384F" author="Dring" box="[346,507,1626,1648]" page="184" pageId="4" pageNumber="431" year="1979">
<bibRefCitation id="EFE04B42FFE0A069CA40F9E0FE4B384F" author="Dring" box="[346,464,1626,1648]" pageId="4" pageNumber="431" refString="Dring, J. C. M. (1979) Amphibians and reptiles from northern Terengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum Natural History (Zoology), 34, 181 - 241." type="journal article" year="1979">Dring 1979</bibRefCitation>
:184
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36B6538FFE0A069CB8DF927FD05398D" pageId="4" pageNumber="431" type="materials_examined">
<paragraph id="8BCE36B3FFE0A069CB8DF927FD3938C1" blockId="4.[151,1436,1693,1970]" pageId="4" pageNumber="431">
<emphasis id="B905EAA1FFE0A069CB8DF927FE903889" bold="true" box="[151,267,1693,1718]" pageId="4" pageNumber="431">
<typeStatus id="54CA8811FFE0A069CB8DF927FE9D3889" box="[151,262,1693,1718]" pageId="4" pageNumber="431" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult female (
<collectionCode id="ED60AE76FFE0A069CAA2F924FE6B3889" box="[440,496,1694,1718]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="431">ZRC</collectionCode>
1.12503) collected on
<date id="FFCF1073FFE0A069C9E3F924FC343889" box="[761,943,1694,1718]" pageId="4" pageNumber="431" value="2012-08-31">31 August 2012</date>
from Gunung Tebu, Terengganu,
<collectingCountry id="F3667623FFE0A069CE2FF927FA073889" box="[1333,1436,1693,1718]" name="Malaysia" pageId="4" pageNumber="431">Malaysia</collectingCountry>
(
<geoCoordinate id="EE455074FFE0A069CB85F97BFEB738E6" box="[159,300,1729,1754]" direction="north" orientation="latitude" pageId="4" pageNumber="431" precision="9" value="5.6018333">05°36.11’ N</geoCoordinate>
<geoCoordinate id="EE455074FFE0A069CA2FF97BFE5C38E5" box="[309,455,1729,1754]" direction="east" orientation="longitude" pageId="4" pageNumber="431" precision="9" value="102.603165">102°36.19’E</geoCoordinate>
;
<quantity id="4C899B56FFE0A069CACEF97BFD8638E6" box="[468,541,1729,1754]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="4" pageNumber="431" unit="m" value="610.0">610 m</quantity>
a.s.l) by Shahrul Anuar, Mohd. A. Muin, E. Quah, L. Grismer, B. Beltran, A. Cobos, A. Alonso, C. Thompson, and C. Ogle.
</paragraph>
<paragraph id="8BCE36B3FFE0A069CBDDF8B3FD05398D" blockId="4.[151,1436,1693,1970]" pageId="4" pageNumber="431">
<emphasis id="B905EAA1FFE0A069CBDDF8B3FEDD391D" bold="true" box="[199,326,1801,1826]" pageId="4" pageNumber="431">
<typeStatus id="54CA8811FFE0A069CBDDF8B3FEDA391D" box="[199,321,1801,1826]" pageId="4" pageNumber="431" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Adult male (
<collectionCode id="ED60AE76FFE0A069CAF1F8B0FDD2391D" box="[491,585,1802,1826]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="4" pageNumber="431">LSUHC</collectionCode>
10899) collected on
<date id="FFCF1073FFE0A069C854F8B0FB96391D" box="[846,1037,1802,1826]" pageId="4" pageNumber="431" value="2012-08-31">31 August 2012</date>
from Gunung Tebu, Terengganu,
<collectingCountry id="F3667623FFE0A069CB8DF897FF653979" box="[151,254,1837,1862]" name="Malaysia" pageId="4" pageNumber="431">Malaysia</collectingCountry>
(
<geoCoordinate id="EE455074FFE0A069CA0AF897FE05397A" box="[272,414,1837,1862]" direction="north" orientation="latitude" pageId="4" pageNumber="431" precision="9" value="5.6018333">05°36.11’ N</geoCoordinate>
<geoCoordinate id="EE455074FFE0A069CAB2F897FDA13979" box="[424,570,1837,1862]" direction="east" orientation="longitude" pageId="4" pageNumber="431" precision="9" value="102.603165">102°36.19’E</geoCoordinate>
;
<quantity id="4C899B56FFE0A069C953F897FD08397A" box="[585,659,1837,1862]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="4" pageNumber="431" unit="m" value="610.0">610 m</quantity>
a.s.l) by Shahrul Anuar, Mohd. A. Muin, E. Quah, L. Grismer, B. Beltran, A. Cobos, A. Alonso, C. Thompson, and C. Ogle. Adult male (
<collectionCode id="ED60AE76FFE0A069C8A9F8E8FB8A3955" box="[947,1041,1874,1898]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="4" pageNumber="431">LSUHC</collectionCode>
11172) and adult females (
<collectionCode id="ED60AE76FFE0A069CE24F8E8FA073955" box="[1342,1436,1874,1898]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="4" pageNumber="431">LSUHC</collectionCode>
11211, 11214) collected on
<date id="FFCF1073FFE0A069CAD4F8CCFDC939B1" box="[462,594,1909,1934]" pageId="4" pageNumber="431" value="2013-07-01">1 July 2013</date>
from the same locality by Chan Kin Onn, Shahrul Anuar, Mohd, A. Muin, A. Sumarli, J. Chan, H. Heinz and L. Grismer.
</paragraph>
</subSubSection>
<caption id="DF0E663BFFE2A06BCB8DF81AFD7C39CC" httpUri="https://zenodo.org/record/230624/files/figure.png" pageId="6" pageNumber="433" targetBox="[268,1283,204,1924]" targetPageId="6">
<paragraph id="8BCE36B3FFE2A06BCB8DF81AFD7C39CC" blockId="6.[151,1436,1952,2035]" pageId="6" pageNumber="433">
<emphasis id="B905EAA1FFE2A06BCB8DF81AFE89398A" bold="true" box="[151,274,1952,1974]" pageId="6" pageNumber="433">FIGURE 1.</emphasis>
Bayesian phylogeny of
<taxonomicName id="4C714D30FFE2A06BC91CF81BFDC23989" box="[518,601,1953,1974]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="6" pageNumber="433" phylum="Chordata" rank="genus">
<emphasis id="B905EAA1FFE2A06BC91CF81BFDC23989" box="[518,601,1953,1974]" italics="true" pageId="6" pageNumber="433">Ansonia</emphasis>
</taxonomicName>
based on 2685 bp fragment of the mitochondrial genes 12S, 16S rRNA and tRNAval. Nodal support are indicated by Bayesian posterior probabilitiy (BPP) / Maximum Likelihood bootstrap (MLBS). Dashes indicate nodes that were not supported in the ML analysis.
</paragraph>
</caption>
<caption id="DF0E663BFFE3A06ACB8DFB96FC873A7D" box="[151,796,1068,1090]" httpUri="https://zenodo.org/record/230625/files/figure.png" pageId="7" pageNumber="434" targetBox="[151,1436,193,1045]" targetPageId="7">
<paragraph id="8BCE36B3FFE3A06ACB8DFB96FC873A7D" blockId="7.[151,796,1068,1090]" box="[151,796,1068,1090]" pageId="7" pageNumber="434">
<emphasis id="B905EAA1FFE3A06ACB8DFB96FE883A7E" bold="true" box="[151,275,1068,1090]" pageId="7" pageNumber="434">FIGURE 2.</emphasis>
Live coloration of female paratype LSUHC 11211.
</paragraph>
</caption>
<subSubSection id="C36B6538FFE3A06ACBDDFBD3FBFE3B9D" pageId="7" pageNumber="434" type="diagnosis">
<paragraph id="8BCE36B3FFE3A06ACBDDFBD3FBFE3B9D" blockId="7.[151,1436,1129,2018]" pageId="7" pageNumber="434">
<emphasis id="B905EAA1FFE3A06ACBDDFBD3FEDA3ABD" bold="true" box="[199,321,1129,1154]" pageId="7" pageNumber="434">Diagnosis.</emphasis>
<taxonomicName id="4C714D30FFE3A06ACA53FBD3FE763ABE" box="[329,493,1129,1153]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="434" phylum="Chordata" rank="species" species="lumut" status="sp. nov.">
<emphasis id="B905EAA1FFE3A06ACA53FBD3FE763ABE" box="[329,493,1129,1153]" italics="true" pageId="7" pageNumber="434">Ansonia lumut</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657DAFFE3A06ACAEFFBD3FDCB3ABD" box="[501,592,1129,1154]" pageId="7" pageNumber="434" rank="species">
<emphasis id="B905EAA1FFE3A06ACAEFFBD3FDCB3ABD" bold="true" box="[501,592,1129,1154]" pageId="7" pageNumber="434">sp. nov.</emphasis>
</taxonomicNameLabel>
is assigned to the genus
<taxonomicName id="4C714D30FFE3A06AC877FBD3FC523ABF" box="[877,969,1129,1152]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="434" phylum="Chordata" rank="genus">
<emphasis id="B905EAA1FFE3A06AC877FBD3FC523ABF" box="[877,969,1129,1152]" italics="true" pageId="7" pageNumber="434">Ansonia</emphasis>
</taxonomicName>
based on its phylogenetic placement and the following morphological characters: small body size; slender limbs; no parotoid glands; weak subarticular tubercles; and membranous foot webbing (
<bibRefCitation id="EFE04B42FFE3A06AC96EFB0BFD683AF5" author="Inger" box="[628,755,1201,1226]" pageId="7" pageNumber="434" refString="Inger, R. F. (1960) A review of the Oriental toads of the genus Ansonia Stoliczka. Fieldiana Zoology, 39, 473 - 503." type="journal article" year="1960">Inger 1960</bibRefCitation>
,
<bibRefCitation id="EFE04B42FFE3A06AC9E4FB0BFCA13AF6" author="Inger" box="[766,826,1201,1225]" pageId="7" pageNumber="434" refString="Inger, R. F. (1992) Variation of apomorphic characters in stream-dwelling tadpoles of the bufonid genus Ansonia (Amphibia: Anura). Zoological Journal of the Linnaean Society, 105, 225 - 237. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1992. tb 01230. x" type="journal article" year="1992">1992</bibRefCitation>
).
<taxonomicName id="4C714D30FFE3A06AC857FB0BFC743AF6" box="[845,1007,1201,1225]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="434" phylum="Chordata" rank="species" species="lumut" status="sp. nov.">
<emphasis id="B905EAA1FFE3A06AC857FB0BFC743AF6" box="[845,1007,1201,1225]" italics="true" pageId="7" pageNumber="434">Ansonia lumut</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657DAFFE3A06AC8ECFB0BFBD43AF5" box="[1014,1103,1201,1226]" pageId="7" pageNumber="434" rank="species">
<emphasis id="B905EAA1FFE3A06AC8ECFB0BFBD43AF5" bold="true" box="[1014,1103,1201,1226]" pageId="7" pageNumber="434">sp. nov.</emphasis>
</taxonomicNameLabel>
can be differentiated from all other congeners by the following combination of characters: SVL
<quantity id="4C899B56FFE3A06AC863FB6EFB9C3AD2" box="[889,1031,1236,1261]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" metricValueMax="2.36" metricValueMin="2.1" pageId="7" pageNumber="434" unit="mm" value="22.3" valueMax="23.6" valueMin="21.0">21–23.6 mm</quantity>
in males,
<quantity id="4C899B56FFE3A06ACF62FB6EFA803AD2" box="[1144,1307,1236,1261]" metricMagnitude="-2" metricUnit="m" metricValue="2.965" metricValueMax="3.16" metricValueMin="2.77" pageId="7" pageNumber="434" unit="mm" value="29.65" valueMax="31.6" valueMin="27.7">27.7–31.6 mm</quantity>
in females; first finger shorter than second; absence of interorbital and tarsal ridges; distinct dorsal tubercles and dorsolateral row of enlarged tubercles; finger tips not expanded into discs; slightly less than two phalanges free of web on fifth toe; light interscapular spot and light patch below eye absent; presence of large, yellow rictal tubercle; dorsum black with greenish-yellow reticulations; flanks with small yellow spots; fore and hind limbs with yellow crossbars; venter light gray with fine, white spotting; males with nuptial pads on first finger.
</paragraph>
</subSubSection>
<subSubSection id="C36B6538FFE3A064CBD5FA17FDEA3963" lastPageId="9" lastPageNumber="436" pageId="7" pageNumber="434" type="description">
<paragraph id="8BCE36B3FFE3A06ACBD5FA17FECB3911" blockId="7.[151,1436,1129,2018]" pageId="7" pageNumber="434">
<emphasis id="B905EAA1FFE3A06ACBD5FA17FE6B3BF9" bold="true" box="[207,496,1453,1478]" pageId="7" pageNumber="434">
Description of
<typeStatus id="54CA8811FFE3A06ACA99FA17FE773BF9" box="[387,492,1453,1478]" pageId="7" pageNumber="434" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female, SVL
<quantity id="4C899B56FFE3A06AC9C1FA17FCDF3BFA" box="[731,836,1453,1477]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="7" pageNumber="434" unit="mm" value="29.3">29.3 mm</quantity>
; head longer than wide (HL/HW=1.12); snout wider than long (SW/SL=1.18), longer than eye diameter (SL/ED=1.46), slightly projecting beyond lower jaw, dorsally convex with a midline depression, truncated with slight median point in dorsal view, truncated and caudoventrally sloping in lateral view; canthus rostralis distinct, lores vertical, slightly concave; nares open laterally, just below canthus, nearly terminal on snout, distance between nares smaller than snout length (
<collectingCountry id="F3667623FFE3A06ACF5DF987FBE03869" box="[1095,1147,1597,1622]" name="India" pageId="7" pageNumber="434">IND</collectingCountry>
/SL=0.58), approximately half of snout width (
<collectingCountry id="F3667623FFE3A06ACA9EF9DBFE233846" box="[388,440,1633,1657]" name="India" pageId="7" pageNumber="434">IND</collectingCountry>
/SW=0.49); eyes large, slightly protruding beyond labials in dorsal view, diameter less than snout length (ED/SL=0.68) and interorbital distance (ED/IOD=0.87), pupils circular; interorbital region flat, distance smaller than snout width (IOD/SW=0.67) and snout length (IOD/SL=0.79); tympanum distinct, oval, taller than wide, vertical diameter smaller than eye diameter (TD/ED=0.65); choanae subcircular, separated by distance larger than their diameter; vomerine ridge and teeth absent; tongue narrow, ending in median point, posterior ⅓ free.
</paragraph>
<paragraph id="8BCE36B3FFE3A06ACBDDF883FE5A3982" blockId="7.[151,1436,1129,2018]" pageId="7" pageNumber="434">
Forelimbs and fingers long and slender; finger length from shortest to longest: I&lt;II&lt;IV&lt;III; basal webbing not extending beyond proximal subarticular tubercle; tips rounded, slightly dilated but not forming discs; subarticular tubercles indistinct; inner and outer metacarpal tubercles weak, oval, flat, inner smaller than outer; supernumerary tubercles absent (
<figureCitation id="134A2A36FFE3A06ACA40F81FFE073982" box="[346,412,1957,1982]" captionStart="FIGURE 3" captionStartId="8.[151,250,1605,1627]" captionTargetBox="[160,1435,304,1578]" captionTargetId="figure@8.[151,1436,303,1583]" captionTargetPageId="8" captionText="FIGURE 3. Holotype of Ansonia lumut sp. nov. (ZRC 1.12503). A: Ventral view of right manus; B: Ventral view of right pes; C: Dorsal surface; D: Ventral surface." httpUri="https://zenodo.org/record/230626/files/figure.png" pageId="7" pageNumber="434">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="8BCE36B3FFE3A065CBDDF873FC6D3EC4" blockId="7.[151,1436,1129,2018]" lastBlockId="8.[151,1436,151,251]" lastPageId="8" lastPageNumber="435" pageId="7" pageNumber="434">
Hindlimbs and toes long and slender (CL/SVL=0.47), foot shorter than tibia (PL/CL=0.76); toe length from shortest to longest: I&lt;II&lt;III≤V&lt;IV; webbing formula: I ½–
<date id="FFCF1073FFECA065C82DFF20FCFB3E8D" box="[823,864,154,179]" pageId="8" pageNumber="435">2 II</date>
½–
<date id="FFCF1073FFECA065C894FF20FC5F3E8D" box="[910,964,151,179]" bridgedPair="-" pageId="8" pageNumber="435">3- III</date>
1–3½
<date id="FFCF1073FFECA065CF0FFF21FBDC3E8C" box="[1045,1095,154,179]" pageId="8" pageNumber="435">IV 3</date>
½–
<date id="FFCF1073FFECA065CF76FF20FB053E8D" box="[1132,1182,151,179]" bridgedPair="-" pageId="8" pageNumber="435">2- V</date>
; tips rounded, slightly dilated but not forming discs; subarticular tubercles indistinct; inner metatarsal tubercle elongate, flat; outer metatarsal tubercle slightly raised, oval, slightly smaller than inner (
<figureCitation id="134A2A36FFECA065C88BFF58FC483EC4" box="[913,979,226,251]" captionStart="FIGURE 3" captionStartId="8.[151,250,1605,1627]" captionTargetBox="[160,1435,304,1578]" captionTargetId="figure@8.[151,1436,303,1583]" captionTargetPageId="8" captionText="FIGURE 3. Holotype of Ansonia lumut sp. nov. (ZRC 1.12503). A: Ventral view of right manus; B: Ventral view of right pes; C: Dorsal surface; D: Ventral surface." httpUri="https://zenodo.org/record/230626/files/figure.png" pageId="8" pageNumber="435">Fig. 3</figureCitation>
B).
</paragraph>
<caption id="DF0E663BFFECA065CB8DF9FFFD8E3845" httpUri="https://zenodo.org/record/230626/files/figure.png" pageId="8" pageNumber="435" targetBox="[160,1435,304,1578]" targetPageId="8">
<paragraph id="8BCE36B3FFECA065CB8DF9FFFD8E3845" blockId="8.[151,1436,1605,1658]" pageId="8" pageNumber="435">
<emphasis id="B905EAA1FFECA065CB8DF9FFFE883865" bold="true" box="[151,275,1605,1627]" pageId="8" pageNumber="435">FIGURE 3.</emphasis>
Holotype of
<taxonomicName id="4C714D30FFECA065CA80F9FCFDB63865" box="[410,557,1605,1627]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="435" phylum="Chordata" rank="species" species="lumut" status="sp. nov.">
<emphasis id="B905EAA1FFECA065CA80F9FCFDB63865" box="[410,557,1605,1627]" italics="true" pageId="8" pageNumber="435">Ansonia lumut</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657DAFFECA065C929F9FCFD1F3864" box="[563,644,1606,1627]" pageId="8" pageNumber="435" rank="species">
<emphasis id="B905EAA1FFECA065C929F9FCFD1F3864" bold="true" box="[563,644,1606,1627]" pageId="8" pageNumber="435">sp. nov.</emphasis>
</taxonomicNameLabel>
(ZRC 1.12503). A: Ventral view of right manus; B: Ventral view of right pes; C: Dorsal surface; D: Ventral surface.
</paragraph>
</caption>
<paragraph id="8BCE36B3FFECA065CBDDF918FB2A39E5" blockId="8.[151,1436,1697,2010]" pageId="8" pageNumber="435">
Upper eyelid, interorbital region, dorsal part of snout and canthus covered with small, flat tubercles bearing brown, keratinized tips; interorbital ridges absent; tubercles absent on lores; single row of small spinules on upper lip and outer margin of upper eyelid; large tubercle at posterior end of upper lip, level with anterior margin of tympanum and a larger tubercle just above rictus, posterior to tympanum; supratympanic fold and parotoid gland absent; slight scapular swelling; back, flanks and dorsal part of limbs with irregularly spaced large and small tubercles bearing brown keratinized spinules, larger tubercles may have more than one spinule; dorsal tubercles largest around nuchal, scapular and dorsolateral region, smallest on dorsal part of limbs (
<figureCitation id="134A2A36FFECA065CF8EF8C3FB4239AD" box="[1172,1241,1913,1938]" captionStart="FIGURE 3" captionStartId="8.[151,250,1605,1627]" captionTargetBox="[160,1435,304,1578]" captionTargetId="figure@8.[151,1436,303,1583]" captionTargetPageId="8" captionText="FIGURE 3. Holotype of Ansonia lumut sp. nov. (ZRC 1.12503). A: Ventral view of right manus; B: Ventral view of right pes; C: Dorsal surface; D: Ventral surface." httpUri="https://zenodo.org/record/230626/files/figure.png" pageId="8" pageNumber="435">Fig. 3</figureCitation>
C); entire ventral surface except for manus and pes with fine, evenly spaced spinules that are most dense around rictal and pectoral region (
<figureCitation id="134A2A36FFECA065CBF4F87BFEAB39E5" box="[238,304,1985,2010]" captionStart="FIGURE 3" captionStartId="8.[151,250,1605,1627]" captionTargetBox="[160,1435,304,1578]" captionTargetId="figure@8.[151,1436,303,1583]" captionTargetPageId="8" captionText="FIGURE 3. Holotype of Ansonia lumut sp. nov. (ZRC 1.12503). A: Ventral view of right manus; B: Ventral view of right pes; C: Dorsal surface; D: Ventral surface." httpUri="https://zenodo.org/record/230626/files/figure.png" pageId="8" pageNumber="435">Fig. 3</figureCitation>
D). Complete measurements of
<typeStatus id="54CA8811FFECA065C98FF87BFD6C39E5" box="[661,759,1985,2010]" pageId="8" pageNumber="435" type="holotype">holotype</typeStatus>
and
<typeStatus id="54CA8811FFECA065C837F878FC0339E5" box="[813,920,1986,2010]" pageId="8" pageNumber="435" type="paratype">paratypes</typeStatus>
are presented in
<tableCitation id="C6F30308FFECA065CF42F87BFB3639E5" box="[1112,1197,1985,2010]" captionStart="TABLE 4" captionStartId="9.[151,239,152,175]" captionTargetBox="[151,1411,305,1279]" captionTargetPageId="9" captionText="TABLE 4. Measurements of the type series summarized by gender. Ranges follow mean ± standard error." httpUri="http://table.plazi.org/id/DF0E663BFFEDA064CB8DFF22FB643E90" pageId="8" pageNumber="435" tableUuid="DF0E663BFFEDA064CB8DFF22FB643E90">Table 4</tableCitation>
.
</paragraph>
<caption id="DF0E663BFFEDA064CB8DFF22FB643E90" ID-Table-UUID="DF0E663BFFEDA064CB8DFF22FB643E90" box="[151,1279,151,176]" httpUri="http://table.plazi.org/id/DF0E663BFFEDA064CB8DFF22FB643E90" pageId="9" pageNumber="436" targetBox="[151,1411,305,1279]" targetIsTable="true" targetPageId="9">
<paragraph id="8BCE36B3FFEDA064CB8DFF22FB643E90" blockId="9.[151,1411,151,286]" box="[151,1279,151,176]" pageId="9" pageNumber="436">
<emphasis id="B905EAA1FFEDA064CB8DFF22FE913E90" bold="true" box="[151,266,152,175]" pageId="9" pageNumber="436">TABLE 4.</emphasis>
Measurements of the type series summarized by gender. Ranges follow mean ± standard error.
</paragraph>
</caption>
<paragraph id="8BCE36B3FFEDA064CBE1FF70FB093F21" blockId="9.[151,1411,151,286]" pageId="9" pageNumber="436">
ZRC 1.12503 LSUHC LSUHC Mean ± Std Error LSUHC LSUHC Mean ± Std Error
<typeStatus id="54CA8811FFEDA064CBE1FF53FEC23EC0" box="[251,345,233,255]" pageId="9" pageNumber="436" type="holotype">Holotype</typeStatus>
11214 11211 Min–Max 11172 10899 Min–Max
<typeStatus id="54CA8811FFEDA064CA86FEB3FB093F21" box="[412,1170,265,286]" pageId="9" pageNumber="436" type="paratype">Paratype Paratype Paratype Paratype</typeStatus>
</paragraph>
<paragraph id="8BCE36B3FFEDA064CB85FE88FA853AC0" pageId="9" pageNumber="436">
<table id="F971C413FFED5F92CB8DFE8BFA183AC0" box="[151,1411,305,1279]" gridcols="6" gridrows="12" pageId="9" pageNumber="436">
<tr id="354134F1FFED5F92CB8DFE8BFA183FB0" box="[151,1411,305,399]" gridrow="0" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFE8BFEBC3FB0" box="[151,295,305,399]" gridcol="0" gridrow="0" pageId="9" pageNumber="436">sex f SVL 29.3</th>
<th id="76905D8DFFED5F92CA86FE8BFE533FB0" box="[412,456,305,399]" gridcol="1" gridrow="0" pageId="9" pageNumber="436">f 27.7</th>
<th id="76905D8DFFED5F92C92FFE8BFDFA3FB0" box="[565,609,305,399]" gridcol="2" gridrow="0" pageId="9" pageNumber="436">f 31.6</th>
<th id="76905D8DFFED5F92C9D4FE8BFC503FB0" box="[718,971,305,399]" gridcol="3" gridrow="0" pageId="9" pageNumber="436">Female (n=3) m 29.5 ± 1.1 23.6 27.7–31.6</th>
<th id="76905D8DFFED5F92CF23FE8BFBFE3FB0" box="[1081,1125,305,399]" gridcol="4" gridrow="0" pageId="9" pageNumber="436">m 21.0</th>
<th id="76905D8DFFED5F92CFC8FE8BFA183FB0" box="[1234,1411,305,399]" gridcol="5" gridrow="0" pageId="9" pageNumber="436">Male (n=2) 22.3 ± 1.3 21.0–23.6</th>
</tr>
<tr id="354134F1FFED5F92CB8DFE10FA183FE0" box="[151,1411,426,479]" gridrow="1" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFE10FEBC3FE0" box="[151,295,426,479]" gridcol="0" gridrow="1" pageId="9" pageNumber="436">HL 8.5</th>
<td id="76905D8DFFED5F92CA86FE10FE533FE0" box="[412,456,426,479]" gridcol="1" gridrow="1" pageId="9" pageNumber="436">8.2</td>
<td id="76905D8DFFED5F92C92FFE10FDFA3FE0" box="[565,609,426,479]" gridcol="2" gridrow="1" pageId="9" pageNumber="436">10.0</td>
<td id="76905D8DFFED5F92C9D4FE10FC503FE0" box="[718,971,426,479]" gridcol="3" gridrow="1" pageId="9" pageNumber="436">8.9 ± 0.6 7.1 8.2–10.0</td>
<td id="76905D8DFFED5F92CF23FE10FBFE3FE0" box="[1081,1125,426,479]" gridcol="4" gridrow="1" pageId="9" pageNumber="436">6.9</td>
<td id="76905D8DFFED5F92CFC8FE10FA183FE0" box="[1234,1411,426,479]" gridcol="5" gridrow="1" pageId="9" pageNumber="436">7.0 ± 0.1 6.9–7.1</td>
</tr>
<tr id="354134F1FFED5F92CB8DFE40FA183C10" box="[151,1411,506,559]" gridrow="2" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFE40FEBC3C10" box="[151,295,506,559]" gridcol="0" gridrow="2" pageId="9" pageNumber="436">HW 7.6</th>
<td id="76905D8DFFED5F92CA86FE40FE533C10" box="[412,456,506,559]" gridcol="1" gridrow="2" pageId="9" pageNumber="436">7.4</td>
<td id="76905D8DFFED5F92C92FFE40FDFA3C10" box="[565,609,506,559]" gridcol="2" gridrow="2" pageId="9" pageNumber="436">8.5</td>
<td id="76905D8DFFED5F92C9D4FE40FC503C10" box="[718,971,506,559]" gridcol="3" gridrow="2" pageId="9" pageNumber="436">7.8 ± 0.3 6.6 7.4–8.5</td>
<td id="76905D8DFFED5F92CF23FE40FBFE3C10" box="[1081,1125,506,559]" gridcol="4" gridrow="2" pageId="9" pageNumber="436">6.3</td>
<td id="76905D8DFFED5F92CFC8FE40FA183C10" box="[1234,1411,506,559]" gridcol="5" gridrow="2" pageId="9" pageNumber="436">6.5 ± 0.2 6.3–6.6</td>
</tr>
<tr id="354134F1FFED5F92CB8DFDF0FA183C40" box="[151,1411,586,639]" gridrow="3" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFDF0FEBC3C40" box="[151,295,586,639]" gridcol="0" gridrow="3" pageId="9" pageNumber="436">SL 3.8</th>
<td id="76905D8DFFED5F92CA86FDF0FE533C40" box="[412,456,586,639]" gridcol="1" gridrow="3" pageId="9" pageNumber="436">3.3</td>
<td id="76905D8DFFED5F92C92FFDF0FDFA3C40" box="[565,609,586,639]" gridcol="2" gridrow="3" pageId="9" pageNumber="436">4.0</td>
<td id="76905D8DFFED5F92C9D4FDF0FC503C40" box="[718,971,586,639]" gridcol="3" gridrow="3" pageId="9" pageNumber="436">3.7 ± 0.2 3.3 3.3–4.0</td>
<td id="76905D8DFFED5F92CF23FDF0FBFE3C40" box="[1081,1125,586,639]" gridcol="4" gridrow="3" pageId="9" pageNumber="436">3.1</td>
<td id="76905D8DFFED5F92CFC8FDF0FA183C40" box="[1234,1411,586,639]" gridcol="5" gridrow="3" pageId="9" pageNumber="436">3.2 ± 0.1 3.1–3.3</td>
</tr>
<tr id="354134F1FFED5F92CB8DFD20FA183CF0" box="[151,1411,666,719]" gridrow="4" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFD20FEBC3CF0" box="[151,295,666,719]" gridcol="0" gridrow="4" pageId="9" pageNumber="436">SW 4.5</th>
<td id="76905D8DFFED5F92CA86FD20FE533CF0" box="[412,456,666,719]" gridcol="1" gridrow="4" pageId="9" pageNumber="436">4.2</td>
<td id="76905D8DFFED5F92C92FFD20FDFA3CF0" box="[565,609,666,719]" gridcol="2" gridrow="4" pageId="9" pageNumber="436">4.6</td>
<td id="76905D8DFFED5F92C9D4FD20FC503CF0" box="[718,971,666,719]" gridcol="3" gridrow="4" pageId="9" pageNumber="436">4.4 ± 0.1 3.8 4.2–4.6</td>
<td id="76905D8DFFED5F92CF23FD20FBFE3CF0" box="[1081,1125,666,719]" gridcol="4" gridrow="4" pageId="9" pageNumber="436">3.3</td>
<td id="76905D8DFFED5F92CFC8FD20FA183CF0" box="[1234,1411,666,719]" gridcol="5" gridrow="4" pageId="9" pageNumber="436">3.6 ± 0.3 3.3–3.8</td>
</tr>
<tr id="354134F1FFED5F92CB8DFD50FA183D20" box="[151,1411,746,799]" gridrow="5" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFD50FEBC3D20" box="[151,295,746,799]" gridcol="0" gridrow="5" pageId="9" pageNumber="436">IND 2.2</th>
<td id="76905D8DFFED5F92CA86FD50FE533D20" box="[412,456,746,799]" gridcol="1" gridrow="5" pageId="9" pageNumber="436">2.3</td>
<td id="76905D8DFFED5F92C92FFD50FDFA3D20" box="[565,609,746,799]" gridcol="2" gridrow="5" pageId="9" pageNumber="436">2.4</td>
<td id="76905D8DFFED5F92C9D4FD50FC503D20" box="[718,971,746,799]" gridcol="3" gridrow="5" pageId="9" pageNumber="436">2.3 ± 0.1 2 2.2–2.4</td>
<td id="76905D8DFFED5F92CF23FD50FBFE3D20" box="[1081,1125,746,799]" gridcol="4" gridrow="5" pageId="9" pageNumber="436">1.8</td>
<td id="76905D8DFFED5F92CFC8FD50FA183D20" box="[1234,1411,746,799]" gridcol="5" gridrow="5" pageId="9" pageNumber="436">1.9 ± 0.1 1.8–2.0</td>
</tr>
<tr id="354134F1FFED5F92CB8DFC80FA183D50" box="[151,1411,826,879]" gridrow="6" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFC80FEBC3D50" box="[151,295,826,879]" gridcol="0" gridrow="6" pageId="9" pageNumber="436">IOD 3.0</th>
<td id="76905D8DFFED5F92CA86FC80FE533D50" box="[412,456,826,879]" gridcol="1" gridrow="6" pageId="9" pageNumber="436">3.0</td>
<td id="76905D8DFFED5F92C92FFC80FDFA3D50" box="[565,609,826,879]" gridcol="2" gridrow="6" pageId="9" pageNumber="436">3.3</td>
<td id="76905D8DFFED5F92C9D4FC80FC503D50" box="[718,971,826,879]" gridcol="3" gridrow="6" pageId="9" pageNumber="436">3.1 ± 0.1 2.7 3.0–3.3</td>
<td id="76905D8DFFED5F92CF23FC80FBFE3D50" box="[1081,1125,826,879]" gridcol="4" gridrow="6" pageId="9" pageNumber="436">2.2</td>
<td id="76905D8DFFED5F92CFC8FC80FA183D50" box="[1234,1411,826,879]" gridcol="5" gridrow="6" pageId="9" pageNumber="436">2.5 ± 0.3 2.2–2.7</td>
</tr>
<tr id="354134F1FFED5F92CB8DFC30FA183D80" box="[151,1411,906,959]" gridrow="7" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFC30FEBC3D80" box="[151,295,906,959]" gridcol="0" gridrow="7" pageId="9" pageNumber="436">ED 2.6</th>
<td id="76905D8DFFED5F92CA86FC30FE533D80" box="[412,456,906,959]" gridcol="1" gridrow="7" pageId="9" pageNumber="436">2.6</td>
<td id="76905D8DFFED5F92C92FFC30FDFA3D80" box="[565,609,906,959]" gridcol="2" gridrow="7" pageId="9" pageNumber="436">3.1</td>
<td id="76905D8DFFED5F92C9D4FC30FC503D80" box="[718,971,906,959]" gridcol="3" gridrow="7" pageId="9" pageNumber="436">2.8 ± 0.2 2.4 2.6–3.1</td>
<td id="76905D8DFFED5F92CF23FC30FBFE3D80" box="[1081,1125,906,959]" gridcol="4" gridrow="7" pageId="9" pageNumber="436">2.3</td>
<td id="76905D8DFFED5F92CFC8FC30FA183D80" box="[1234,1411,906,959]" gridcol="5" gridrow="7" pageId="9" pageNumber="436">2.4 ± 0.1 2.3–2.4</td>
</tr>
<tr id="354134F1FFED5F92CB8DFC60FA183A30" box="[151,1411,986,1039]" gridrow="8" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFC60FEBC3A30" box="[151,295,986,1039]" gridcol="0" gridrow="8" pageId="9" pageNumber="436">TD 1.7</th>
<td id="76905D8DFFED5F92CA86FC60FE533A30" box="[412,456,986,1039]" gridcol="1" gridrow="8" pageId="9" pageNumber="436">1.6</td>
<td id="76905D8DFFED5F92C92FFC60FDFA3A30" box="[565,609,986,1039]" gridcol="2" gridrow="8" pageId="9" pageNumber="436">1.8</td>
<td id="76905D8DFFED5F92C9D4FC60FC503A30" box="[718,971,986,1039]" gridcol="3" gridrow="8" pageId="9" pageNumber="436">1.7 ± 0.1 1.6 1.6–1.8</td>
<td id="76905D8DFFED5F92CF23FC60FBFE3A30" box="[1081,1125,986,1039]" gridcol="4" gridrow="8" pageId="9" pageNumber="436">1.0</td>
<td id="76905D8DFFED5F92CFC8FC60FA183A30" box="[1234,1411,986,1039]" gridcol="5" gridrow="8" pageId="9" pageNumber="436">1.3 ± 0.3 1.0–1.6</td>
</tr>
<tr id="354134F1FFED5F92CB8DFB90FA183A60" box="[151,1411,1066,1119]" gridrow="9" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFB90FEBC3A60" box="[151,295,1066,1119]" gridcol="0" gridrow="9" pageId="9" pageNumber="436">ML 7.8</th>
<td id="76905D8DFFED5F92CA86FB90FE533A60" box="[412,456,1066,1119]" gridcol="1" gridrow="9" pageId="9" pageNumber="436">7.4</td>
<td id="76905D8DFFED5F92C92FFB90FDFA3A60" box="[565,609,1066,1119]" gridcol="2" gridrow="9" pageId="9" pageNumber="436">8.6</td>
<td id="76905D8DFFED5F92C9D4FB90FC503A60" box="[718,971,1066,1119]" gridcol="3" gridrow="9" pageId="9" pageNumber="436">7.9 ± 0.4 6.2 7.4–8.6</td>
<td id="76905D8DFFED5F92CF23FB90FBFE3A60" box="[1081,1125,1066,1119]" gridcol="4" gridrow="9" pageId="9" pageNumber="436">5.5</td>
<td id="76905D8DFFED5F92CFC8FB90FA183A60" box="[1234,1411,1066,1119]" gridcol="5" gridrow="9" pageId="9" pageNumber="436">5.9 ± 0.4 5.5–6.2</td>
</tr>
<tr id="354134F1FFED5F92CB8DFBC0FA183A90" box="[151,1411,1146,1199]" gridrow="10" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFBC0FEBC3A90" box="[151,295,1146,1199]" gridcol="0" gridrow="10" pageId="9" pageNumber="436">CL 13.9</th>
<td id="76905D8DFFED5F92CA86FBC0FE533A90" box="[412,456,1146,1199]" gridcol="1" gridrow="10" pageId="9" pageNumber="436">13.7</td>
<td id="76905D8DFFED5F92C92FFBC0FDFA3A90" box="[565,609,1146,1199]" gridcol="2" gridrow="10" pageId="9" pageNumber="436">15.7</td>
<td id="76905D8DFFED5F92C9D4FBC0FC503A90" box="[718,971,1146,1199]" gridcol="3" gridrow="10" pageId="9" pageNumber="436">14.4 ± 0.6 11.6 13.7–15.7</td>
<td id="76905D8DFFED5F92CF23FBC0FBFE3A90" box="[1081,1125,1146,1199]" gridcol="4" gridrow="10" pageId="9" pageNumber="436">10.9</td>
<td id="76905D8DFFED5F92CFC8FBC0FA183A90" box="[1234,1411,1146,1199]" gridcol="5" gridrow="10" pageId="9" pageNumber="436">11.3 ± 0.4 10.9–11.6</td>
</tr>
<tr id="354134F1FFED5F92CB8DFB70FA183AC0" box="[151,1411,1226,1279]" gridrow="11" pageId="9" pageNumber="436">
<th id="76905D8DFFED5F92CB8DFB70FEBC3AC0" box="[151,295,1226,1279]" gridcol="0" gridrow="11" pageId="9" pageNumber="436">PL 10.5</th>
<td id="76905D8DFFED5F92CA86FB70FE533AC0" box="[412,456,1226,1279]" gridcol="1" gridrow="11" pageId="9" pageNumber="436">10.5</td>
<td id="76905D8DFFED5F92C92FFB70FDFA3AC0" box="[565,609,1226,1279]" gridcol="2" gridrow="11" pageId="9" pageNumber="436">11.3</td>
<td id="76905D8DFFED5F92C9D4FB70FC503AC0" box="[718,971,1226,1279]" gridcol="3" gridrow="11" pageId="9" pageNumber="436">10.8 ± 0.3 8.4 10.5–11.3</td>
<td id="76905D8DFFED5F92CF23FB70FBFE3AC0" box="[1081,1125,1226,1279]" gridcol="4" gridrow="11" pageId="9" pageNumber="436">7.7</td>
<td id="76905D8DFFED5F92CFC8FB70FA183AC0" box="[1234,1411,1226,1279]" gridcol="5" gridrow="11" pageId="9" pageNumber="436">8.1 ± 0.4 7.7–8.4</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCE36B3FFEDA064CBDDFAF6FC6C3827" blockId="9.[151,1437,1356,2029]" pageId="9" pageNumber="436">
<emphasis id="B905EAA1FFEDA064CBDDFAF6FE323B5A" bold="true" box="[199,425,1356,1381]" pageId="9" pageNumber="436">Coloration in life.</emphasis>
Dorsal base color dark brown to black. Top of head, lores and back with yellow reticulations bearing greenish flecks. Large, yellow tubercle at rictus, followed anteriorly by smaller yellow tubercle at posterior end of upper lip. An even smaller, yellow tubercle present between rictal and posterior upper labial tubercles on left side of head but absent on right side. Three small, yellow patches along upper lip, below eye, lore, and rostrum. Yellow patches on ventral side of mandible. Flanks with small yellow spots. Yellow crossbars on front and hind limbs. Venter light gray with fine, white spotting.
</paragraph>
<paragraph id="8BCE36B3FFEDA064CBDDF99EFDF438BB" blockId="9.[151,1437,1356,2029]" pageId="9" pageNumber="436">
<emphasis id="B905EAA1FFEDA064CBDDF99EFD973802" bold="true" box="[199,524,1572,1597]" pageId="9" pageNumber="436">Coloration in preservative.</emphasis>
Yellow coloration creamy white and dorsal reticulations light gray. Keratinized tips on tubercles brown. Venter uniform creamy white with very fine dark brown stippling that is most dense on gular region, tibia, tarsus, manus, and pes.
</paragraph>
<paragraph id="8BCE36B3FFEDA064CBDDF935FDEA3963" blockId="9.[151,1437,1356,2029]" pageId="9" pageNumber="436">
<emphasis id="B905EAA1FFEDA064CBDDF935FEA43897" bold="true" box="[199,319,1679,1704]" pageId="9" pageNumber="436">Variation.</emphasis>
All
<typeStatus id="54CA8811FFEDA064CA60F92AFE7E3897" box="[378,485,1680,1704]" pageId="9" pageNumber="436" type="paratype">paratypes</typeStatus>
closely resemble
<typeStatus id="54CA8811FFEDA064C9A4F935FCBB3897" box="[702,800,1679,1704]" pageId="9" pageNumber="436" type="holotype">holotype</typeStatus>
in coloration and pattern. Female
<typeStatus id="54CA8811FFEDA064CFA5F92AFAB13897" box="[1215,1322,1680,1704]" pageId="9" pageNumber="436" type="paratype">paratypes</typeStatus>
(LSUHC 11211, 11214) have additional yellow tubercle below tympanum between rictal and posterior upper labial tubercle, tubercle on right side larger than one on the left. Male
<typeStatus id="54CA8811FFEDA064C9E3F962FCFF38CF" box="[761,868,1752,1776]" pageId="9" pageNumber="436" type="paratype">paratypes</typeStatus>
(LSUHC 10899, 11172) smaller in size, SVL 21.0 mm and
<quantity id="4C899B56FFEDA064CBE2F946FEC7392B" box="[248,348,1788,1813]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="9" pageNumber="436" unit="mm" value="23.6">23.6 mm</quantity>
respectively. Vocal slit present on left side of mouth leads into median subgular vocal sac. Small, indistinct nuptial pad of brown asperities on medial surface of first finger between base and distal phalangeal joint. Nuptial pad not visible on LSUHC 10899.
</paragraph>
</subSubSection>
<subSubSection id="C36B6538FFEDA067CBDDF8DDFBAF3A9F" lastPageId="10" lastPageNumber="437" pageId="9" pageNumber="436" type="materials_examined">
<paragraph id="8BCE36B3FFEDA067CBDDF8DDFBAF3A9F" blockId="9.[151,1437,1356,2029]" lastBlockId="10.[151,1436,151,1544]" lastPageId="10" lastPageNumber="437" pageId="9" pageNumber="436">
<emphasis id="B905EAA1FFEDA064CBDDF8DDFEF639BF" bold="true" box="[199,365,1895,1920]" pageId="9" pageNumber="436">Comparisons.</emphasis>
Morphological data for species comparisons were obtained from
<bibRefCitation id="EFE04B42FFEDA064CF53F8DDFA8639BF" box="[1097,1309,1895,1920]" pageId="9" pageNumber="436" refString="Wood, P. L. Jr., Grismer, L. L., Norhayati, A. &amp; Senawi, J. (2008) Two new species of torrent-dwelling toads Ansonia Stoliczka, 1870 (Anura: Bufonidae) from Peninsular Malaysia. Herpetologica, 64 (3), 321 - 340. http: // dx. doi. org / 10.1655 / 07 - 065.1" type="journal article">
Wood
<emphasis id="B905EAA1FFEDA064CF88F8D3FB5139BF" box="[1170,1226,1896,1920]" italics="true" pageId="9" pageNumber="436">et al.</emphasis>
(2008)
</bibRefCitation>
,
<bibRefCitation id="EFE04B42FFEDA064CE32F8DDFEBF399B" pageId="9" pageNumber="436" refString="Wilkinson, J. A., Sellas, A. B. &amp; Vindum, J. (2012) A new species of Ansonia (Anura: Bufonidae) from northern Tanintharyi Division, Myanmar. Zootaxa, 3163, 54 - 68." type="journal article">
Wilkinson
<emphasis id="B905EAA1FFEDA064CB8DF837FF4B399B" box="[151,208,1932,1956]" italics="true" pageId="9" pageNumber="436">et al.</emphasis>
(2012)
</bibRefCitation>
, and material examined (see Appendix).
<taxonomicName id="4C714D30FFEDA064C9EEF836FC03399B" box="[756,920,1932,1956]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="436" phylum="Chordata" rank="species" species="lumut" status="sp. nov.">
<emphasis id="B905EAA1FFEDA064C9EEF836FC03399B" box="[756,920,1932,1956]" italics="true" pageId="9" pageNumber="436">Ansonia lumut</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657DAFFEDA064C885F836FC61399A" box="[927,1018,1932,1957]" pageId="9" pageNumber="436" rank="species">
<emphasis id="B905EAA1FFEDA064C885F836FC61399A" bold="true" box="[927,1018,1932,1957]" pageId="9" pageNumber="436">sp. nov.</emphasis>
</taxonomicNameLabel>
is most closely related to three other Peninsular Malaysian species,
<taxonomicName id="4C714D30FFEDA064CAEDF80AFD1C39F7" box="[503,647,1968,1992]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="436" phylum="Chordata" rank="species" species="malayana">
<emphasis id="B905EAA1FFEDA064CAEDF80AFD1C39F7" box="[503,647,1968,1992]" italics="true" pageId="9" pageNumber="436">A. malayana</emphasis>
</taxonomicName>
,
<emphasis id="B905EAA1FFEDA064C982F80AFCD039F7" box="[664,843,1968,1992]" italics="true" pageId="9" pageNumber="436">
<taxonomicName id="4C714D30FFEDA064C982F80AFCDC39F7" box="[664,839,1968,1992]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="436" phylum="Chordata" rank="species" species="penangensis">A. penangensis</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C714D30FFEDA064C892F80AFBCC39F7" box="[904,1111,1968,1992]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="436" phylum="Chordata" rank="species" species="jeetsukumarani">
<emphasis id="B905EAA1FFEDA064C892F80AFBCC39F7" box="[904,1111,1968,1992]" italics="true" pageId="9" pageNumber="436">A. jeetsukumarani</emphasis>
</taxonomicName>
but can be morphologically distinguished from them by the absence of a light interscapular spot (vs. present in
<taxonomicName id="4C714D30FFEDA064CF56F86EFB4139D3" box="[1100,1242,2004,2028]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="436" phylum="Chordata" rank="species" species="malayana">
<emphasis id="B905EAA1FFEDA064CF56F86EFB4139D3" box="[1100,1242,2004,2028]" italics="true" pageId="9" pageNumber="436">A. malayana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C714D30FFEDA064CFF0F86EFA0E39D3" box="[1258,1429,2004,2028]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="436" phylum="Chordata" rank="species" species="penangensis">
<emphasis id="B905EAA1FFEDA064CFF0F86EFA0E39D3" box="[1258,1429,2004,2028]" italics="true" pageId="9" pageNumber="436">A. penangensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C714D30FFEEA067CBD7FF22FE3B3E8F" box="[205,416,152,176]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="jeetsukumarani">
<emphasis id="B905EAA1FFEEA067CBD7FF22FE3B3E8F" box="[205,416,152,176]" italics="true" pageId="10" pageNumber="437">A. jeetsukumarani</emphasis>
</taxonomicName>
), dorsum with greenish-yellow reticulations (vs. orange hourglass-shaped lines in
<taxonomicName id="4C714D30FFEEA067CE9EFF22FE9E3EEB" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="malayana">
<emphasis id="B905EAA1FFEEA067CE9EFF22FE9E3EEB" italics="true" pageId="10" pageNumber="437">A. malayana</emphasis>
</taxonomicName>
, almost uniform black in
<taxonomicName id="4C714D30FFEEA067C959FF06FD693EEB" box="[579,754,188,212]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="penangensis">
<emphasis id="B905EAA1FFEEA067C959FF06FD693EEB" box="[579,754,188,212]" italics="true" pageId="10" pageNumber="437">A. penangensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C714D30FFEEA067C82EFF06FB9F3EEB" box="[820,1028,188,212]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="jeetsukumarani">
<emphasis id="B905EAA1FFEEA067C82EFF06FB9F3EEB" box="[820,1028,188,212]" italics="true" pageId="10" pageNumber="437">A. jeetsukumarani</emphasis>
</taxonomicName>
), large, yellow rictal tubercle (vs. whitish in
<emphasis id="B905EAA1FFEEA067CA15FF5AFDCC3EC8" box="[271,599,224,248]" italics="true" pageId="10" pageNumber="437">
<taxonomicName id="4C714D30FFEEA067CA15FF5AFE043EC7" box="[271,415,224,248]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="malayana">A. malayana</taxonomicName>
,
<taxonomicName id="4C714D30FFEEA067CAB6FF5AFDCC3EC8" box="[428,599,224,247]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="penangensis">A. penangensis</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C714D30FFEEA067C98FFF5AFCFA3EC7" box="[661,865,224,248]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="jeetsukumarani">
<emphasis id="B905EAA1FFEEA067C98FFF5AFCFA3EC7" box="[661,865,224,248]" italics="true" pageId="10" pageNumber="437">A. jeetsukumarani</emphasis>
</taxonomicName>
), and yellow cross-bars on limbs (vs. orange in
<emphasis id="B905EAA1FFEEA067CE9EFF5AFE263F23" italics="true" pageId="10" pageNumber="437">
<taxonomicName id="4C714D30FFEEA067CE9EFF5AFE933F23" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="malayana">A. malayana</taxonomicName>
,
<taxonomicName id="4C714D30FFEEA067CA0EFEBEFE263F23" box="[276,445,260,284]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="penangensis">A. penangensis</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C714D30FFEEA067CAE1FEBEFD533F23" box="[507,712,260,284]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="jeetsukumarani">
<emphasis id="B905EAA1FFEEA067CAE1FEBEFD533F23" box="[507,712,260,284]" italics="true" pageId="10" pageNumber="437">A. jeetsukumarani</emphasis>
</taxonomicName>
). On the Thai-Malay Peninsula and Indochina, it differs from
<taxonomicName id="4C714D30FFEEA067CE9EFEBEFF4F3F7F" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="latiffi">
<emphasis id="B905EAA1FFEEA067CE9EFEBEFF4F3F7F" italics="true" pageId="10" pageNumber="437">A. latiffi</emphasis>
</taxonomicName>
by smaller body size in males (vs. SVL
<quantity id="4C899B56FFEEA067C9B6FE92FCC93F7F" box="[684,850,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.615" metricValueMax="3.82" metricValueMin="3.41" pageId="10" pageNumber="437" unit="mm" value="36.15" valueMax="38.2" valueMin="34.1">34.1–38.2 mm</quantity>
) and females (vs. SVL
<quantity id="4C899B56FFEEA067CF7FFE9DFA953F7F" box="[1125,1294,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="5.06" metricValueMax="5.07" metricValueMin="5.05" pageId="10" pageNumber="437" unit="mm" value="50.6" valueMax="50.7" valueMin="50.5">50.5–50.7 mm</quantity>
), first finger shorter than second (vs. first finger reaching tip of second), absence of tarsal ridge (vs. presence), dorsum blackish with greenish-yellow reticulations (vs. almost uniform brown), presence of large, yellow rictal tubercle (vs. absence), and yellow crossbars on limbs (vs. pale orange); from
<taxonomicName id="4C714D30FFEEA067C869FE2EFC663F93" box="[883,1021,404,428]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="latirostra">
<emphasis id="B905EAA1FFEEA067C869FE2EFC663F93" box="[883,1021,404,428]" italics="true" pageId="10" pageNumber="437">A. latirostra</emphasis>
</taxonomicName>
by the absence of interorbital ridges (vs. present), no rictal gland (vs. present), and greenish-yellow reticulations on dorsum (vs. almost uniform black); from
<taxonomicName id="4C714D30FFEEA067CBCEFE66FEF93FCB" box="[212,354,476,500]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="tiomanica">
<emphasis id="B905EAA1FFEEA067CBCEFE66FEF93FCB" box="[212,354,476,500]" italics="true" pageId="10" pageNumber="437">A. tiomanica</emphasis>
</taxonomicName>
by smaller body size in males (vs. SVL
<quantity id="4C899B56FFEEA067C838FE66FC1D3FCB" box="[802,902,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="10" pageNumber="437" unit="mm" value="31.2">31.2 mm</quantity>
) and females (vs. SVL
<quantity id="4C899B56FFEEA067CF96FE66FB683FCB" box="[1164,1267,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" pageId="10" pageNumber="437" unit="mm" value="38.4">38.4 mm</quantity>
), and greenishyellow reticulations on dorsum (vs. yellow spots); from
<taxonomicName id="4C714D30FFEEA067C802FDBAFC2E3C27" box="[792,949,512,536]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="endauensis">
<emphasis id="B905EAA1FFEEA067C802FDBAFC2E3C27" box="[792,949,512,536]" italics="true" pageId="10" pageNumber="437">A. endauensis</emphasis>
</taxonomicName>
by having blackish dorsum with greenishyellow reticulations (vs. almost uniform black), and yellow crossbars on limbs (vs. orange); from
<taxonomicName id="4C714D30FFEEA067CFF1FD9EFAE33C03" box="[1259,1400,548,572]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="B905EAA1FFEEA067CFF1FD9EFAE33C03" box="[1259,1400,548,572]" italics="true" pageId="10" pageNumber="437">A. siamensis</emphasis>
</taxonomicName>
by smaller body size in males (vs. SVL 28.0 mm) and females (vs. SVL 35.0 mm), finger tips not expanded into discs (vs. expanded into distinct discs), distinct dorsal tubercles (vs. reduced or absent), presence of large, yellow rictal tubercle (vs. absence), and slightly less than two phalanges free of web on fifth toe (vs. one phalanx free); from
<taxonomicName id="4C714D30FFEEA067CE9EFD2AFF6F3CF3" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="kraensis">
<emphasis id="B905EAA1FFEEA067CE9EFD2AFF6F3CF3" italics="true" pageId="10" pageNumber="437">A. kraensis</emphasis>
</taxonomicName>
by the absence of a light interscapular spot (vs. present), presence of large, yellow rictal tubercle (vs. absence); slightly less than two phalanges free of web on fifth toe (vs. half phalanx free), dorsum with greenishyellow reticulations (vs. dorsum brown with darker hourglass-shaped pattern), and venter light gray with fine, white spots (vs. dark brown with distinct, white reticulations); from
<taxonomicName id="4C714D30FFEEA067C88EFC9AFAE33D07" authority="Matsui, Nabhitabhata &amp; Panha" authorityName="Matsui, Nabhitabhata &amp; Panha" box="[916,1400,799,824]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="inthanon">
<emphasis id="B905EAA1FFEEA067C88EFC9AFB833D07" box="[916,1048,800,824]" italics="true" pageId="10" pageNumber="437">A. inthanon</emphasis>
Matsui, Nabhitabhata &amp; Panha
</taxonomicName>
by larger body size in females (vs. SVL
<quantity id="4C899B56FFEEA067C928FCFEFD423D63" box="[562,729,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.425" metricValueMax="2.52" metricValueMin="2.33" pageId="10" pageNumber="437" unit="mm" value="24.25" valueMax="25.2" valueMin="23.3">23.3–25.2 mm</quantity>
), absence of light, interscapular spot (vs. presence), presence of large, yellow tubercle (vs. absence), dorsum blackish with greenish-yellow reticulations (vs. brown with lighter, indistinct markings), presence of dorsolateral row of enlarged tubercles (vs. absence), slightly less than two phalanges free of web on fifth toe (vs. one phalanx free), and venter light gray with fine, white spots (vs. brown with bright yellow blotches); from
<taxonomicName id="4C714D30FFEEA067C939FC6EFD2E3DD3" box="[547,693,980,1004]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="thinthinae">
<emphasis id="B905EAA1FFEEA067C939FC6EFD2E3DD3" box="[547,693,980,1004]" italics="true" pageId="10" pageNumber="437">A. thinthinae</emphasis>
</taxonomicName>
by the absence of a light patch below eye (vs. presence), absence of light interscapular spot (vs. presence), dorsum black with greenish-yellow reticulations (vs. dark brown, lacking distinct markings), slightly less than two phalanges free of web on fifth toe (vs. half phalanx free), venter light gray with fine, white spots (vs. dark brown with yellow reticulations), and males with nuptial pads on first finger (vs. nuptial pad covering first and second fingers). Differences between the new species and other geographically and phylogenetically distant congeners are summarized in
<bibRefCitation id="EFE04B42FFEEA067C9E2FB3DFC5D3A9F" author="Wood" box="[760,966,1159,1184]" pageId="10" pageNumber="437" refString="Wood, P. L. Jr., Grismer, L. L., Norhayati, A. &amp; Senawi, J. (2008) Two new species of torrent-dwelling toads Ansonia Stoliczka, 1870 (Anura: Bufonidae) from Peninsular Malaysia. Herpetologica, 64 (3), 321 - 340. http: // dx. doi. org / 10.1655 / 07 - 065.1" type="journal article" year="2008">
Wood
<emphasis id="B905EAA1FFEEA067C858FB33FCE83A9F" box="[834,883,1160,1184]" italics="true" pageId="10" pageNumber="437">et al</emphasis>
. (2008
</bibRefCitation>
:
<tableCitation id="C6F30308FFEEA067C8CBFB3DFBB93A9F" box="[977,1058,1159,1184]" captionStart="TABLE 1" captionStartId="1.[151,239,1317,1340]" captionTargetBox="[159,1361,1368,1721]" captionTargetPageId="1" captionText="TABLE 1. Primers used in this study for PCR amplification and sequencing reactions." httpUri="http://table.plazi.org/id/DF0E663BFFE5A06CCB8DFA9FFBAA3B02" pageId="10" pageNumber="437" tableUuid="DF0E663BFFE5A06CCB8DFA9FFBAA3B02">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B6538FFEEA067CBDDFB16FBB83BA2" pageId="10" pageNumber="437" type="distribution">
<paragraph id="8BCE36B3FFEEA067CBDDFB16FBB83BA2" blockId="10.[151,1436,151,1544]" pageId="10" pageNumber="437">
<emphasis id="B905EAA1FFEEA067CBDDFB16FDD73AFA" bold="true" box="[199,588,1196,1221]" pageId="10" pageNumber="437">Distribution and natural history.</emphasis>
<taxonomicName id="4C714D30FFEEA067C948FB16FD6F3AFB" box="[594,756,1196,1220]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="lumut" status="sp. nov.">
<emphasis id="B905EAA1FFEEA067C948FB16FD6F3AFB" box="[594,756,1196,1220]" italics="true" pageId="10" pageNumber="437">Ansonia lumut</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657DAFFEEA067C9E0FB16FCC83AFA" box="[762,851,1196,1221]" pageId="10" pageNumber="437" rank="species">
<emphasis id="B905EAA1FFEEA067C9E0FB16FCC83AFA" bold="true" box="[762,851,1196,1221]" pageId="10" pageNumber="437">sp. nov.</emphasis>
</taxonomicNameLabel>
occurs in hill dipterocarp forest on Gunung Tebu (at
<quantity id="4C899B56FFEEA067CB8DFB75FF463AD8" box="[151,221,1231,1256]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="10" pageNumber="437" unit="m" value="610.0">610 m</quantity>
elevation) and Gunung Lawit (at
<quantity id="4C899B56FFEEA067C94EFB6AFD7A3AD8" box="[596,737,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.035" metricValueMax="1.28" metricValueMin="0.79" pageId="10" pageNumber="437" unit="m" value="1035.0" valueMax="1280.0" valueMin="790.0">790–1280 m</quantity>
elevation;
<bibRefCitation id="EFE04B42FFEEA067C846FB75FC463AD7" author="Dring" box="[860,989,1231,1256]" pageId="10" pageNumber="437" refString="Dring, J. C. M. (1979) Amphibians and reptiles from northern Terengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum Natural History (Zoology), 34, 181 - 241." type="journal article" year="1979">Dring 1979</bibRefCitation>
) in the northeastern state of Terengganu (
<figureCitation id="134A2A36FFEEA067CB85FB4EFF7E3B33" box="[159,229,1268,1293]" captionStart="FIGURE 4" captionStartId="11.[151,250,1722,1744]" captionTargetBox="[166,1425,617,1647]" captionTargetId="figure@11.[155,1431,617,1650]" captionTargetPageId="11" captionText="FIGURE 4. Distribution of Ansonia in Peninsular Malaysia. Localities are color coded to corresponding species. Green star = type locality of A. lumut sp. nov. at Gunung Tebu, green triangle = Gunung Lawit; triangles = members in the A. lumut clade; circles = members outside of the A. lumut clade." httpUri="https://zenodo.org/record/230627/files/figure.png" pageId="10" pageNumber="437">Fig. 4</figureCitation>
). During the day, specimens were found in small rock fissures along a slow moving stream. The stream was approximately
<quantity id="4C899B56FFEEA067CA25FAADFEF23B10" box="[319,361,1303,1328]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="437" unit="m" value="5.0">5 m</quantity>
wide under a semi-closed canopy forest and consisted of dark-colored granite rock covered with algae and moss (
<figureCitation id="134A2A36FFEEA067CA49FA86FE033B6A" box="[339,408,1340,1365]" captionStart="FIGURE 5" captionStartId="12.[151,250,1068,1090]" captionTargetBox="[151,1436,193,1046]" captionTargetId="figure@12.[151,1436,193,1047]" captionTargetPageId="12" captionText="FIGURE 5. Habitat of Ansonia lumut sp. nov. at Gunung Tebu, Terengganu." httpUri="https://zenodo.org/record/230628/files/figure.png" pageId="10" pageNumber="437">Fig. 5</figureCitation>
). At night, toads were found on rocks and small, overhanging vegetation within the stream and along its banks away from strong torrent zones. Female
<typeStatus id="54CA8811FFEEA067C85EFADBFC3F3B46" box="[836,932,1377,1401]" pageId="10" pageNumber="437" type="paratype">paratype</typeStatus>
LSUHC 11214 was found to be gravid with unfertilized, unpigmented eggs that were approximately
<quantity id="4C899B56FFEEA067C80AFA3EFC0C3BA3" box="[784,919,1412,1436]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="10" pageNumber="437" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
in diameter.
</paragraph>
</subSubSection>
<subSubSection id="C36B6538FFEEA067CBDDFA1DFEE53837" pageId="10" pageNumber="437" type="etymology">
<paragraph id="8BCE36B3FFEEA067CBDDFA1DFEE53837" blockId="10.[151,1436,151,1544]" pageId="10" pageNumber="437">
<emphasis id="B905EAA1FFEEA067CBDDFA1DFED53BFF" bold="true" box="[199,334,1447,1472]" pageId="10" pageNumber="437">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="4C714D30FFEEA067C956FA1DFD093BFF" box="[588,658,1447,1472]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="lumut">lumut</taxonomicName>
” is derived from the Malay word for moss, in reference to the new species’ color pattern, which gives it a mossy appearance. Suggested English name: Mossy Stream Toad; Malay name: Kodok
<taxonomicName id="4C714D30FFEEA067CA2FFA55FEE13837" box="[309,378,1519,1544]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="437" phylum="Chordata" rank="species" species="lumut">lumut</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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