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<document id="18B9F9C3416F529ACC7AC8ECABE8F0BF" ID-CLB-Dataset="21707" ID-DOI="10.11646/zootaxa.4790.2.10" ID-GBIF-Dataset="3f492afe-37c4-4bc9-b7d1-5f8d804cf5bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889408" ID-ZooBank="7F4A9B99-C12E-473B-B258-7BE5697F4986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859385935" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="038E87D4FF80FFA1FF1579E0FB387125" docLanguage="en" docName="zootaxa.4790.2.10.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus armatus Perkins 2020, new species" docType="treatment" docVersion="7" lastPageNumber="362" masterDocId="FFB7FFACFF81FFA5FF827F44FFB87066" masterDocTitle="Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="374" masterPageNumber="358" pageNumber="359" updateTime="1698842324292" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="155D577BEFE9A15BDFAC1DB41D3F9BE5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2E119EB02B4340B8B169C0C97F0F9C04">
<mods:title id="2441EAD9DC4BCD41CFD498779CB69530">Taxonomy of Ecuadorian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="75E685AAE78A524F39A92386CD2D8AD4" type="personal">
<mods:role id="E8EF4A07B66EA4CBC9DA417B665AB496">
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</mods:role>
<mods:namePart id="B91DD89FD089A849B81CE0D8D8E1F11A">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="87A9486ED7ADA0AAFBEADE0E1EA92494">text</mods:typeOfResource>
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<mods:title id="577B732C82036FF4B90E13DA3DF0C7EF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0AD8C6F6E90E5D191D736DD9BDA8F7DD">
<mods:date id="4EEEF87B50B777B43414AD2DB5927FFC">2020</mods:date>
<mods:detail id="59C968568D5E0FE884F916C88A97786A" type="pubDate">
<mods:number id="7A2DCC6329F921111D6E5012B0E84E55">2020-06-11</mods:number>
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<mods:detail id="BE752CD2EF3910CEEFB9D6FE2C7CE194" type="volume">
<mods:number id="18122C0E00976CDC796213AA623524D9">4790</mods:number>
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<mods:number id="829E0D4C0ABE0305032CD167E8C184A9">2</mods:number>
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<mods:classification id="521C3690330DB126FF4EF7B6CD64436E">journal article</mods:classification>
<mods:identifier id="4FFCC996685FE8124AACE7CEDC414A42" type="CLB-Dataset">21707</mods:identifier>
<mods:identifier id="565FDF001EA08313C7B00783DA4B9CF1" type="DOI">10.11646/zootaxa.4790.2.10</mods:identifier>
<mods:identifier id="738FADF146490BC0BC0C55506D82E903" type="GBIF-Dataset">3f492afe-37c4-4bc9-b7d1-5f8d804cf5bb</mods:identifier>
<mods:identifier id="502049FEF6663502FED94674DBA168C3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7F8CF9B6558018FE1717BD65ACE4DC74" type="Zenodo-Dep">3889408</mods:identifier>
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</mods:mods>
<treatment id="038E87D4FF80FFA1FF1579E0FB387125" ID-GBIF-Taxon="164501161" LSID="urn:lsid:plazi:treatment:038E87D4FF80FFA1FF1579E0FB387125" httpUri="http://treatment.plazi.org/id/038E87D4FF80FFA1FF1579E0FB387125" lastPageId="4" lastPageNumber="362" pageId="1" pageNumber="359">
<subSubSection id="C33D6549FF80FFA4FF1579E0FD9E76D8" box="[151,550,1700,1727]" pageId="1" pageNumber="359" type="nomenclature">
<paragraph id="8B9836C2FF80FFA4FF1579E0FD9E76D8" blockId="1.[151,550,1700,1763]" box="[151,550,1700,1727]" pageId="1" pageNumber="359">
<heading id="D0D081AEFF80FFA4FF1579E0FD9E76D8" bold="true" box="[151,550,1700,1727]" fontSize="11" level="1" pageId="1" pageNumber="359" reason="1">
<emphasis id="B953EAD0FF80FFA4FF1579E0FD9E76D8" bold="true" box="[151,550,1700,1727]" pageId="1" pageNumber="359">
<taxonomicName id="4C274D41FF80FFA4FF1579E0FE3F76D9" ID-CoL="8Y4P6" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,391,1700,1727]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="359" phylum="Arthropoda" rank="species" species="armatus" status="sp. nov.">
<emphasis id="B953EAD0FF80FFA4FF1579E0FE3F76D9" bold="true" box="[151,391,1700,1727]" italics="true" pageId="1" pageNumber="359">Hydrochus armatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26057ABFF80FFA4FE1479E1FD9E76D8" box="[406,550,1700,1727]" pageId="1" pageNumber="359" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF80FFA4FF15798CFF477685" box="[151,255,1736,1763]" pageId="1" pageNumber="359" type="description">
<paragraph id="8B9836C2FF80FFA4FF15798CFF477685" blockId="1.[151,550,1700,1763]" box="[151,255,1736,1763]" pageId="1" pageNumber="359">
<figureCitation id="131C2A47FF80FFA4FF15798CFF5B7685" box="[151,227,1736,1763]" captionStart="FIGURE 1" captionStartId="2.[151,250,2019,2044]" captionTargetBox="[192,1405,448,1963]" captionTargetId="figure@2.[151,1435,417,1993]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3889410" httpUri="https://zenodo.org/record/3889410/files/figure.png" pageId="1" pageNumber="359">Figs. 1</figureCitation>
,
<figureCitation id="131C2A47FF80FFA4FF73798DFF477685" box="[241,255,1737,1763]" captionStart="FIGURE 2" captionStartId="3.[151,250,1586,1611]" captionTargetBox="[167,1423,193,1555]" captionTargetId="figure@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURE 2. Hydrochus armatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889412" httpUri="https://zenodo.org/record/3889412/files/figure.png" pageId="1" pageNumber="359">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF80FFA4FF157855FCE87715" pageId="1" pageNumber="359" type="materials_examined">
<paragraph id="8B9836C2FF80FFA4FF157855FCE87715" blockId="1.[151,1437,1808,2015]" pageId="1" pageNumber="359">
<emphasis id="B953EAD0FF80FFA4FF157855FEFE774D" bold="true" box="[151,326,1808,1835]" pageId="1" pageNumber="359">Type Material.</emphasis>
<materialsCitation id="3B4F3C9FFF80FFA4FEC97854FD9D7729" ID-GBIF-Occurrence="2632886312" collectingDate="1974-04-10" collectionCode="NMNH" collectorName="Boyce A. Drummond" country="Ecuador" location="Rio" municipality="Limoncocha" pageId="1" pageNumber="359" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
<typeStatus id="549C8860FF80FFA4FEC97854FE0B774C" box="[331,435,1808,1834]" pageId="1" pageNumber="359" type="holotype">Holotype</typeStatus>
(male): “
<collectingCountry id="F3307652FF80FFA4FD9A7854FDC4774C" box="[536,636,1808,1834]" name="Ecuador" pageId="1" pageNumber="359">Ecuador</collectingCountry>
:
<collectingRegion id="49E3F820FF80FFA4FD077855FC95774C" box="[645,813,1808,1835]" country="Ecuador" name="Napo" pageId="1" pageNumber="359">Napo Province</collectingRegion>
,
<collectingMunicipality id="6BFCACB8FF80FFA4FCB57854FC7E774C" box="[823,966,1808,1834]" pageId="1" pageNumber="359">Limoncocha</collectingMunicipality>
, on
<location id="8EF86019FF80FFA4FC727854FBA0774C" LSID="urn:lsid:plazi:treatment:038E87D4FF80FFA1FF1579E0FB387125:8EF86019FF80FFA4FC727854FBA0774C" box="[1008,1048,1808,1834]" country="Ecuador" municipality="Limoncocha" name="Rio" pageId="1" pageNumber="359" stateProvince="Napo">Rio</location>
<collectingRegion id="49E3F820FF80FFA4FB9C7855FBE3774D" box="[1054,1115,1809,1835]" country="Ecuador" name="Napo" pageId="1" pageNumber="359">Napo</collectingRegion>
<date id="FF991002FF80FFA4FBDD7855FB58774D" box="[1119,1248,1809,1835]" pageId="1" pageNumber="359" value="1974-04-10">
<collectingDate id="EFDDE9EAFF80FFA4FBDD7855FB58774D" box="[1119,1248,1809,1835]" pageId="1" pageNumber="359" value="1974-04-10">10-IV-1974</collectingDate>
</date>
<collectorName id="26D25314FF80FFA4FB647855FF627728" pageId="1" pageNumber="359">Boyce A. Drummond</collectorName>
,
<collectingMethod id="52664ED5FF80FFA4FF667871FE117729" box="[228,425,1844,1871]" pageId="1" pageNumber="359">III blacklight trap</collectingMethod>
” (
<collectionCode id="ED36AE07FF80FFA4FE437871FDA57729" box="[449,541,1845,1871]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="359" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3C9FFF80FFA4FDAC7871FBF57729" ID-GBIF-Occurrence="2632886329" box="[558,1101,1844,1871]" collectingDate="1974-04-10" collectionCode="NMNH" collectorName="Boyce A. Drummond" country="Ecuador" location="Rio" municipality="Limoncocha" pageId="1" pageNumber="359" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<typeStatus id="549C8860FF80FFA4FDAC7871FD237729" box="[558,667,1845,1871]" pageId="1" pageNumber="359" type="paratype">Paratypes</typeStatus>
(2): Same data as holotype (1
<collectionCode id="ED36AE07FF80FFA4FC687871FBFD7729" box="[1002,1093,1845,1871]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="359" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C9FFF80FFA4FBD47871FCF57715" ID-GBIF-Occurrence="2632886315" collectingDate="1977-06-12" collectionCode="NMNH" collectorName="W. E. Steiner" country="Ecuador" latitude="-1.55" location="Limoncocha" longLatPrecision="1308" longitude="-75.583336" pageId="1" pageNumber="359" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<collectingRegion id="49E3F820FF80FFA4FBD47871FB477728" box="[1110,1279,1844,1871]" country="Ecuador" name="Napo" pageId="1" pageNumber="359">Napo Province</collectingRegion>
,
<location id="8EF86019FF80FFA4FA8B7870FA207728" LSID="urn:lsid:plazi:treatment:038E87D4FF80FFA1FF1579E0FB387125:8EF86019FF80FFA4FA8B7870FA207728" box="[1289,1432,1844,1870]" country="Ecuador" latitude="-1.55" longLatPrecision="1308" longitude="-75.583336" name="Limoncocha" pageId="1" pageNumber="359" stateProvince="Napo">Limoncocha</location>
, (
<geoCoordinate id="EE135005FF80FFA4FF23781DFF487715" box="[161,240,1881,1907]" degrees="1" direction="south" minutes="33" orientation="latitude" pageId="1" pageNumber="359" precision="925" value="-1.55">1°33’S</geoCoordinate>
,
<geoCoordinate id="EE135005FF80FFA4FF7F781CFEDF7715" box="[253,359,1880,1907]" degrees="75" direction="west" minutes="35" orientation="longitude" pageId="1" pageNumber="359" precision="925" value="-75.583336">75°35’W</geoCoordinate>
),
<date id="FF991002FF80FFA4FEF9781CFE4D7714" box="[379,501,1880,1906]" pageId="1" pageNumber="359" value="1977-06-12">
<collectingDate id="EFDDE9EAFF80FFA4FEF9781CFE4D7714" box="[379,501,1880,1906]" pageId="1" pageNumber="359" value="1977-06-12">12.vi.1977</collectingDate>
</date>
, leg.
<collectorName id="26D25314FF80FFA4FDB2781DFD7C7714" box="[560,708,1880,1907]" pageId="1" pageNumber="359">W. E. Steiner</collectorName>
(1
<collectionCode id="ED36AE07FF80FFA4FD68781DFCFD7715" box="[746,837,1881,1907]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="359" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF80FFA7FF457838FE7470B1" lastPageId="2" lastPageNumber="360" pageId="1" pageNumber="359" type="diagnosis">
<paragraph id="8B9836C2FF80FFA7FF457838FE7470B1" blockId="1.[151,1437,1808,2015]" lastBlockId="2.[151,1437,153,395]" lastPageId="2" lastPageNumber="360" pageId="1" pageNumber="359">
<emphasis id="B953EAD0FF80FFA4FF457838FE6F77F1" bold="true" box="[199,471,1916,1943]" pageId="1" pageNumber="359">Differential Diagnosis.</emphasis>
Among Ecuadorian species, recognized by the combination of moderate size (ca.
<quantity id="4CDF9B27FF80FFA4FF1578E5FEBA77DD" box="[151,258,1953,1979]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="359" unit="mm" value="4.1">4.10 mm</quantity>
), the moderately deep pronotal impressions, and the distinctive male genitalia, which have the parameres strongly scleotized and, in dorsal/ventral views widened at midlength, in addition to the strongly sclerotized aedeagus (
<figureCitation id="131C2A47FF83FFA7FE8F7FDDFEEB70D5" box="[269,339,153,179]" captionStart="FIGURE 2" captionStartId="3.[151,250,1586,1611]" captionTargetBox="[167,1423,193,1555]" captionTargetId="figure@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURE 2. Hydrochus armatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889412" httpUri="https://zenodo.org/record/3889412/files/figure.png" pageId="2" pageNumber="360">Fig. 2</figureCitation>
; described below). This species has a generalized
<taxonomicName id="4C274D41FF83FFA7FC057FDDFBB970D5" authorityName="Leach" authorityYear="1817" box="[903,1025,153,179]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B953EAD0FF83FFA7FC057FDDFBB970D5" box="[903,1025,153,179]" italics="true" pageId="2" pageNumber="360">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF83FFA1FF457FA4FC8C7179" lastPageId="4" lastPageNumber="362" pageId="2" pageNumber="360" type="description">
<paragraph id="8B9836C2FF83FFA7FF457FA4FA2F71ED" blockId="2.[151,1437,153,395]" pageId="2" pageNumber="360">
<emphasis id="B953EAD0FF83FFA7FF457FA4FEE9709D" bold="true" box="[199,337,224,251]" pageId="2" pageNumber="360">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="131C2A47FF83FFA7FDDF7FA5FD1B709D" box="[605,675,225,251]" captionStart="FIGURE 2" captionStartId="3.[151,250,1586,1611]" captionTargetBox="[167,1423,193,1555]" captionTargetId="figure@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURE 2. Hydrochus armatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889412" httpUri="https://zenodo.org/record/3889412/files/figure.png" pageId="2" pageNumber="360">Fig. 2</figureCitation>
). Size:
<typeStatus id="549C8860FF83FFA7FD757FA5FCE1709D" box="[759,857,225,251]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 4.10/1.42; head width 0.97; pronotum l/w 0.94/1.03; PA 0.90; PB 0.84; elytra 2.55/1.42. Body size moderate (ca.
<quantity id="4CDF9B27FF83FFA7FF157E6DFF467125" box="[151,254,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="360" unit="mm" value="4.1">4.10 mm</quantity>
). Dorsum grey to greenish grey, with slight to moderately strong iridescence, elytra with black spots. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 1–1.5x strial puncture diameter. Interstria 9
<superScript id="7C529B8AFF83FFA7FACC7E08FAE3713C" attach="left" box="[1358,1371,332,346]" fontSize="6" pageId="2" pageNumber="360">th</superScript>
more convex than others, overhanging 10
<superScript id="7C529B8AFF83FFA7FDAA7E34FD8D7118" attach="left" box="[552,565,368,382]" fontSize="6" pageId="2" pageNumber="360">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C529B8AFF83FFA7FC407E34FC777118" attach="left" box="[962,975,368,382]" fontSize="6" pageId="2" pageNumber="360">th</superScript>
moderately raised, moderately elongate.
</paragraph>
<caption id="DF58664AFF83FFA7FF1578A7FD2B779A" ID-DOI="http://doi.org/10.5281/zenodo.3889410" ID-Zenodo-Dep="3889410" box="[151,659,2019,2044]" httpUri="https://zenodo.org/record/3889410/files/figure.png" pageId="2" pageNumber="360" startId="2.[151,250,2019,2044]" targetBox="[192,1405,448,1963]" targetPageId="2">
<paragraph id="8B9836C2FF83FFA7FF1578A7FD2B779A" blockId="2.[151,659,2019,2044]" box="[151,659,2019,2044]" pageId="2" pageNumber="360">
<emphasis id="B953EAD0FF83FFA7FF1578A7FEAB779A" bold="true" box="[151,275,2019,2044]" pageId="2" pageNumber="360">FIGURE 1.</emphasis>
Dorsal habitus of
<taxonomicName id="4C274D41FF83FFA7FE4C78A7FD84779D" authorityName="Leach" authorityYear="1817" box="[462,572,2019,2043]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="B953EAD0FF83FFA7FE4C78A7FD84779D" box="[462,572,2019,2043]" italics="true" pageId="2" pageNumber="360">Hydrochus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<caption id="DF58664AFF82FFA6FF157976FC00762D" ID-DOI="http://doi.org/10.5281/zenodo.3889412" ID-Zenodo-Dep="3889412" box="[151,952,1586,1611]" httpUri="https://zenodo.org/record/3889412/files/figure.png" pageId="3" pageNumber="361" startId="3.[151,250,1586,1611]" targetBox="[167,1423,193,1555]" targetPageId="3">
<paragraph id="8B9836C2FF82FFA6FF157976FC00762D" blockId="3.[151,952,1586,1611]" box="[151,952,1586,1611]" pageId="3" pageNumber="361">
<emphasis id="B953EAD0FF82FFA6FF157976FEAB762D" bold="true" box="[151,275,1586,1611]" pageId="3" pageNumber="361">FIGURE 2.</emphasis>
<taxonomicName id="4C274D41FF82FFA6FE9B7977FE59762D" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[281,481,1587,1611]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="361" phylum="Arthropoda" rank="species" species="armatus" status="sp. nov.">
<emphasis id="B953EAD0FF82FFA6FE9B7977FE59762D" box="[281,481,1587,1611]" italics="true" pageId="3" pageNumber="361">Hydrochus armatus</emphasis>
</taxonomicName>
<emphasis id="B953EAD0FF82FFA6FE657977FD99762D" bold="true" box="[487,545,1587,1611]" pageId="3" pageNumber="361">
<taxonomicNameLabel id="A26057ABFF82FFA6FE657977FD99762D" box="[487,545,1587,1611]" pageId="3" pageNumber="361" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<paragraph id="8B9836C2FF82FFA6FF45793EFDF476BD" blockId="3.[151,1437,1657,2044]" pageId="3" pageNumber="361">Pronotum slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; anterior margin slightly bisinuate; anterior depressions deeper than posterior, reliefs lateral to anterior depressions rounded.</paragraph>
<paragraph id="8B9836C2FF82FFA6FF4579A1FC2F7745" blockId="3.[151,1437,1657,2044]" pageId="3" pageNumber="361">Elytra with spaces between strial punctures narrow walls to ca. 1x their diameter; apices conjointly rounded in dorsal view, in lateral view outer margin with very slight angulation.</paragraph>
<paragraph id="8B9836C2FF82FFA6FF457869FDBF770D" blockId="3.[151,1437,1657,2044]" pageId="3" pageNumber="361">Ventral characters: Mentum coarsely punctate, with median fovea. Submentum with two foveae. Legs testaceous, femora darker than tibiae.</paragraph>
<paragraph id="8B9836C2FF82FFA1FF457831FF6D70B1" blockId="3.[151,1437,1657,2044]" lastBlockId="4.[151,1436,153,323]" lastPageId="4" lastPageNumber="362" pageId="3" pageNumber="361">
Male genitalia (
<figureCitation id="131C2A47FF82FFA6FEFC7831FE7D77E9" box="[382,453,1909,1935]" captionStart="FIGURE 2" captionStartId="3.[151,250,1586,1611]" captionTargetBox="[167,1423,193,1555]" captionTargetId="figure@3.[151,1435,181,1561]" captionTargetPageId="3" captionText="FIGURE 2. Hydrochus armatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3889412" httpUri="https://zenodo.org/record/3889412/files/figure.png" pageId="3" pageNumber="361">Fig. 2</figureCitation>
) general characters: Genitalia very large, strongly sclerotized, with margins between strongly sclerotized areas and weakly or non-sclerotized areas very distinct; parameres longer than aedeagus, plsa small, parameres in dorsal/ventral views with middle 1/3 widened and strongly sclerotized, in lateral view parameres much wider, pointed apically, with two moderately large pwsa; aedeagus lanceolate, with alsm, apex sharply pointed; basal piece slightly shorter than parameres, dorsal margin arcuate in lateral view, orifice strongly sclerotized.
</paragraph>
<paragraph id="8B9836C2FF85FFA1FF457FA5FCC2709D" blockId="4.[151,1436,153,323]" box="[199,890,225,251]" pageId="4" pageNumber="362">Dorsal surface: adbl moderately large; adtl at about distal 1/3.</paragraph>
<paragraph id="8B9836C2FF85FFA1FF457E41FC8C7179" blockId="4.[151,1436,153,323]" box="[199,820,260,287]" pageId="4" pageNumber="362">Ventral surface: avtl at about proximal 1/5 of aedeagus.</paragraph>
</subSubSection>
<subSubSection id="C33D6549FF85FFA1FF457E6DFB387125" box="[199,1152,297,323]" pageId="4" pageNumber="362" type="etymology">
<paragraph id="8B9836C2FF85FFA1FF457E6DFB387125" blockId="4.[151,1436,153,323]" box="[199,1152,297,323]" pageId="4" pageNumber="362">
<emphasis id="B953EAD0FF85FFA1FF457E6DFEF67125" bold="true" box="[199,334,297,323]" pageId="4" pageNumber="362">Etymology.</emphasis>
Named in reference to the shape and sclerotization of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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