405 lines
44 KiB
XML
405 lines
44 KiB
XML
<document id="3B4E1DB76E5176A4D42EF4B7D7CFE4AA" ID-CLB-Dataset="53799" ID-DOI="10.1080/00222933.2023.2171820" ID-GBIF-Dataset="e3544c70-f9c9-4daa-a16e-a827022616bc" ID-ISSN="1464-5262" ID-Zenodo-Dep="7737268" ID-ZooBank="AA69FCBE-81ED-4B41-90D6-0D32EBE887CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678694520066" checkinUser="plazi" docAuthor="Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P." docDate="2023" docId="038E87BB9B7AFFF04B36FA2EFB82642A" docLanguage="en" docName="JNATHIST.57.1-4.193-242.pdf" docOrigin="Journal of Natural History 57 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2023.2171820" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Mogannia tenebrosa Lee and Marshall 2023, sp." docType="treatment" docVersion="8" lastPageNumber="232" masterDocId="FFB7FFC39B59FFD74A52FFEEFFDF6525" masterDocTitle="Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species" masterLastPageNumber="242" masterPageNumber="193" pageNumber="228" updateTime="1699476474976" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="35791247DE824AF60E69A0167D29F791">Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species</mods:title>
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<mods:namePart id="591A11349B6AD8E75D92B5988D87AA10">Lee, Young June</mods:namePart>
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<mods:namePart id="4C29BF97713D1A84F31772303724A2E3">Marshall, David C.</mods:namePart>
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<mods:namePart id="4F808E20D47C7481A828BD95E8C9E308">Mohagan, Alma B.</mods:namePart>
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<mods:namePart id="82F0109BBD02554334D20ED9B22B3ED3">Hill, Kathy B. R.</mods:namePart>
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<mods:title id="BEA855C4B9741A3716D4CF69E5D3140D">Journal of Natural History</mods:title>
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<mods:date id="43C0EDF3C7F003E8F8DF0C3335904071">2023</mods:date>
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<treatment id="038E87BB9B7AFFF04B36FA2EFB82642A" ID-DOI="http://doi.org/10.5281/zenodo.7738589" ID-GBIF-Taxon="209450803" ID-Zenodo-Dep="7738589" LSID="urn:lsid:plazi:treatment:038E87BB9B7AFFF04B36FA2EFB82642A" httpUri="http://treatment.plazi.org/id/038E87BB9B7AFFF04B36FA2EFB82642A" lastPageId="39" lastPageNumber="232" pageId="35" pageNumber="228">
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<subSubSection id="C33D65269B7AFFF44B36FA2EFC1F60FF" box="[356,960,1472,1498]" pageId="35" pageNumber="228" type="nomenclature">
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<heading id="D0D081C19B7AFFF44B36FA2EFC1F60FF" box="[356,960,1472,1498]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="228" reason="5">
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24.
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<taxonomicName id="4C274D2E9B7AFFF44BDCFA2EFC5960FF" ID-CoL="013061d5-8011-4f53-8fa7-af44eba958d5" authority="Lee and Marshall" authorityName="Lee and Marshall" authorityYear="2023" box="[398,902,1472,1498]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="193" phylum="Arthropoda" rank="species" species="tenebrosa" status="sp.">
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<emphasis id="B953EABF9B7AFFF44BDCFA2EFD5360FF" bold="true" box="[398,652,1472,1498]" italics="true" pageId="35" pageNumber="228">Mogannia tenebrosa</emphasis>
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Lee and Marshall
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</taxonomicName>
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nov.
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<subSubSection id="C33D65269B7AFFF44865FA09FD336324" box="[567,748,1511,1537]" pageId="35" pageNumber="228" type="description">
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<paragraph id="8B9836AD9B7AFFF44865FA09FD336324" blockId="35.[567,748,1511,1537]" box="[567,748,1511,1537]" pageId="35" pageNumber="228">
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(
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<figureCitation id="131C2A289B7AFFF4486DFA09FD636324" box="[575,700,1511,1537]" captionStart="Figure 15" captionStartId="37.[160,225,1398,1421]" captionTargetBox="[162,1138,185,1353]" captionTargetId="figure-79@37.[162,1154,142,1367]" captionTargetPageId="37" captionText="Figure 15. Mogannia tenebrosa Lee and Marshall sp. nov., holotype, male, Mindanao, Philippines. (A) Dorsal habitus; (B) ventral habitus; (C) pygofer, oblique ventral view." figureDoi="http://doi.org/10.5281/zenodo.7737304" httpUri="https://zenodo.org/record/7737304/files/figure.png" pageId="35" pageNumber="228">Figures 15</figureCitation>
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,
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<figureCitation id="131C2A289B7AFFF44895FA09FD3B6324" box="[711,740,1511,1537]" captionStart="Figure 16" captionStartId="38.[160,225,708,731]" captionTargetBox="[164,1151,144,674]" captionTargetId="figure-373@38.[162,1154,142,676]" captionTargetPageId="38" captionText="Figure 16. Mogannia tenebrosa Lee and Marshall sp. nov. Waveforms (A, C) of male calling song at two time scales, the former with accompanying spectrogram (B)." figureDoi="http://doi.org/10.5281/zenodo.7737306" httpUri="https://zenodo.org/record/7737306/files/figure.png" pageId="35" pageNumber="228">16</figureCitation>
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)
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</paragraph>
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.
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, specimen code 12.PH.
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.CAB.01,
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/
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Rd., nr bdry,
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07°40.39
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N
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126°12.77
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E
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.
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. Marshall and
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<collectorName id="26D2537B9B7AFFF44B65F949FE0C63E5" box="[311,467,1703,1728]" pageId="35" pageNumber="228">A.B. Mohagan</collectorName>
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(
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).
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.
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, same location as holotype, genitalia dissection G103 (
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);
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, specimen code 12.
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.
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.HWM.01,
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,
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side
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. Hamiguitan, Ca. 1.5 h up trail from Tumalite,
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||
<geoCoordinate id="EE13506A9B7DFFF3495DFF3AFC5765C8" box="[783,904,212,237]" degrees="06" direction="north" minutes="43.95" orientation="latitude" pageId="36" pageNumber="229" precision="9" value="6.7325">
|
||
06°43.95
|
||
<emphasis id="B953EABF9B7DFFF3493DFF3AFCAA65C9" bold="true" box="[879,885,212,236]" pageId="36" pageNumber="229">ʹ</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE13506A9B7DFFF349C5FF3AFBC465C8" box="[919,1051,212,237]" degrees="126" direction="east" minutes="08.13" orientation="longitude" pageId="36" pageNumber="229" precision="9" value="126.1355">
|
||
126°08.13
|
||
<emphasis id="B953EABF9B7DFFF34E57FF3AFBD465C9" bold="true" box="[1029,1035,212,236]" pageId="36" pageNumber="229">ʹ</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
,
|
||
<date id="FF99106D9B7DFFF34E7AFF3AFF066435" pageId="36" pageNumber="229" value="2012-04-26">26 April 2012</date>
|
||
, K.B.
|
||
<collectionCode id="ED36AE689B7DFFF34B59FF19FEC36435" box="[267,284,247,272]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="36" pageNumber="229">R</collectionCode>
|
||
. Hill, D.C. Marshall and A.B.
|
||
<collectorName id="26D2537B9B7DFFF34807FF19FD606435" box="[597,703,247,272]" pageId="36" pageNumber="229">Mohagan</collectorName>
|
||
(
|
||
<collectionCode id="ED36AE689B7DFFF3489EFF19FCF66435" box="[716,809,247,272]" country="USA" httpUri="http://grbio.org/cool/jrf7-y871" name="University of Connecticut Biodiversity Research Collections" pageId="36" pageNumber="229">UCONN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D65269B7DFFF34AF2FED2FC8C64BE" pageId="36" pageNumber="229" type="etymology">
|
||
<paragraph id="8B9836AD9B7DFFF34AF2FED2FEC26473" blockId="36.[160,1155,316,411]" box="[160,285,316,342]" pageId="36" pageNumber="229">
|
||
<emphasis id="B953EABF9B7DFFF34AF2FED2FEC26473" bold="true" box="[160,285,316,342]" italics="true" pageId="36" pageNumber="229">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34AF2FEB1FC8C64BE" blockId="36.[160,1155,316,411]" pageId="36" pageNumber="229">The specific name is a Latin feminine adjective meaning ̍dark̾, in reference to the darkcoloured body and the base of the fore wings of this species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D65269B7DFFF34AF2FE28FC5863E5" pageId="36" pageNumber="229" type="description">
|
||
<paragraph id="8B9836AD9B7DFFF34AF2FE28FBD464C5" blockId="36.[160,1156,454,688]" box="[160,1035,454,480]" pageId="36" pageNumber="229">
|
||
<emphasis id="B953EABF9B7DFFF34AF2FE28FBD464C5" bold="true" box="[160,1035,454,480]" italics="true" pageId="36" pageNumber="229">
|
||
Measurements of types (
|
||
<specimenCount id="9D21FD249B7DFFF34B91FE28FDFF64C5" box="[451,544,454,480]" pageId="36" pageNumber="229" type="male">2 males</specimenCount>
|
||
,
|
||
<typeStatus id="549C880F9B7DFFF3487FFE28FD4A64C5" box="[557,661,454,480]" pageId="36" pageNumber="229" type="holotype">holotype</typeStatus>
|
||
(larger) and
|
||
<typeStatus id="549C880F9B7DFFF34960FE28FC4364C5" box="[818,924,454,480]" pageId="36" pageNumber="229" type="paratype">paratype</typeStatus>
|
||
from Mt.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34AF2FE07FEEA6726" blockId="36.[160,1156,454,688]" box="[160,309,489,515]" pageId="36" pageNumber="229">
|
||
<emphasis id="B953EABF9B7DFFF34AF2FE07FEEA6726" bold="true" box="[160,309,489,515]" italics="true" pageId="36" pageNumber="229">Hamiguitan)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34AF2FDE2FE466795" blockId="36.[160,1156,454,688]" pageId="36" pageNumber="229">Length of body: 12.5 (11.2–13.7). Length of head and thorax together: 6.3 (5.4–7.1). Length of abdomen: 6.2 (5.8–6.6). Width of head including compound eyes: 3.2 (3.1–3.2). Width of pronotum: 4.6 (4.1–5.0). Width of mesonotum: 4.0 (3.6–4.4). Width of abdominal tergite 3: 5.3 (4.6–5.9). Length of fore wing: 15.4 (14.0–16.8). Width of fore wing: 5.2 (4.8–5.5). Wing span: 33.4 (30.8–36.0).</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34AF2FD32FE5767D3" blockId="36.[160,1156,732,1520]" box="[160,392,732,758]" pageId="36" pageNumber="229">
|
||
<emphasis id="B953EABF9B7DFFF34AF2FD32FE5767D3" bold="true" box="[160,392,732,758]" italics="true" pageId="36" pageNumber="229">Description of male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34AF2FD11FCD16686" blockId="36.[160,1156,732,1520]" pageId="36" pageNumber="229">Head black to fuscous; narrow, much narrower than mesonotum. Distance between lateral ocelli and compound eyes slightly shorter than distance between lateral ocelli. Antenna mostly dull ochraceous but with black pedicel and scape. Postclypeus greatly protruding anteriad. Rostrum with small dull ochraceous patch proximally, with apex slightly passing beyond posterior margin of mid coxae.</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34A92FC43FC2766CD" blockId="36.[160,1156,732,1520]" pageId="36" pageNumber="229">Thorax including cruciform elevation black to fuscous. Anterolateral pronotal collar not dentate. Fore-, mid- and hind coxae each with light green distal membrane.</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34A92FC1CFE98619D" blockId="36.[160,1156,732,1520]" pageId="36" pageNumber="229">Wings: Fore wing partly hyaline with venation ochraceous except for basal half of costal vein pinkish brown and with broad dark brown patch on basal half of fore wing. Basal cell tinged with pinkish brown. Basal membrane pinkish brown. Hind wing hyaline. Hind wing costal cell, radial cell and medial cell with small pinkish brown patch basally. Vannus with pinkish brown patch basally on about basal one-fourth. Hind wing jugum pinkish brown.</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34A92FB2CFC7F6005" blockId="36.[160,1156,732,1520]" pageId="36" pageNumber="229">Operculum black to fuscous; hook- or meniscus-shaped, with length of 2.5–3 times the width, with apex heading towards centre of body; not reaching posterior margin of abdominal sternite II. Opercula nearly touching each other medially.</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34A92FAC4FCF960D5" blockId="36.[160,1156,732,1520]" pageId="36" pageNumber="229">Abdomen wide and short, about as long as head and thorax together; mostly black with small reddish brown patches, especially on median and sublateral parts. Tergite 3 much wider than pronotum. Timbal cover black but tinged with reddish brown, very small, triangular with anterior angle rounded, wider than long. Timbal mostly exposed. Abdominal sternites dark brown to reddish brown with black caudal margin on each of sternites II–VII. Sternite VIII and epipleurite VIII nearly red.</paragraph>
|
||
<paragraph id="8B9836AD9B7DFFF34AF2F9F2FC5863E5" blockId="36.[160,1156,1564,1728]" pageId="36" pageNumber="229">
|
||
<emphasis id="B953EABF9B7DFFF34AF2F9F2FECD6313" bold="true" box="[160,274,1564,1590]" italics="true" pageId="36" pageNumber="229">Genitalia.</emphasis>
|
||
Pygofer circular in ventral view. Dorsal beak narrow and long. Uncus short with widely concave apex. Median clasper process extending posteriad in ventral view, but curved inward in lateral view. Lateral clasper lobe short. Aedeagus with spine-like processes apically. Distal shoulder with short triangular tooth in lateral view. Upper lobe uncurved, with rounded apex in lateral view. Basal lobe indistinct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5866259B7CFFF24AF2FA98FCB5608E" ID-DOI="http://doi.org/10.5281/zenodo.7737304" ID-Zenodo-Dep="7737304" httpUri="https://zenodo.org/record/7737304/files/figure.png" pageId="37" pageNumber="230" startId="37.[160,225,1398,1421]" targetBox="[162,1138,185,1353]" targetPageId="37">
|
||
<paragraph id="8B9836AD9B7CFFF24AF2FA98FCB5608E" blockId="37.[160,1157,1398,1451]" pageId="37" pageNumber="230">
|
||
<emphasis id="B953EABF9B7CFFF24AF2FA98FED560A8" bold="true" box="[160,266,1398,1421]" pageId="37" pageNumber="230">Figure 15.</emphasis>
|
||
<taxonomicName id="4C274D2E9B7CFFF24B46FA98FD6560A8" authority="Lee and Marshall" authorityName="Lee and Marshall" authorityYear="2023" box="[276,698,1398,1421]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="193" phylum="Arthropoda" rank="species" species="tenebrosa" status="sp.">
|
||
<emphasis id="B953EABF9B7CFFF24B46FA98FE0260A8" box="[276,477,1398,1421]" italics="true" pageId="37" pageNumber="230">Mogannia tenebrosa</emphasis>
|
||
Lee and Marshall
|
||
<taxonomicNameLabel id="A26057C49B7CFFF248CCFA98FD6560A8" box="[670,698,1398,1421]" pageId="37" pageNumber="230" rank="species">sp.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov., holotype, male, Mindanao, Philippines. (A) Dorsal habitus; (B) ventral habitus; (C) pygofer, oblique ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33D65269B7CFFF14AF2FA17FB8B6006" lastPageId="38" lastPageNumber="231" pageId="37" pageNumber="230" type="discussion">
|
||
<paragraph id="8B9836AD9B7CFFF24AF2FA17FED96336" blockId="37.[159,1155,1529,1728]" box="[160,262,1529,1555]" pageId="37" pageNumber="230">
|
||
<emphasis id="B953EABF9B7CFFF24AF2FA17FED96336" bold="true" box="[160,262,1529,1555]" italics="true" pageId="37" pageNumber="230">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7CFFF24AF2F9F3FC2A63E5" blockId="37.[159,1155,1529,1728]" pageId="37" pageNumber="230">
|
||
This species resembles
|
||
<taxonomicName id="4C274D2E9B7CFFF24BC9F9F3FD266313" authority="(Germar, 1830)" baseAuthorityName="Germar" baseAuthorityYear="1830" box="[411,761,1565,1590]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="230" phylum="Arthropoda" rank="species" species="conica">
|
||
<emphasis id="B953EABF9B7CFFF24BC9F9F3FD8D6313" box="[411,594,1565,1590]" italics="true" pageId="37" pageNumber="230">Mogannia conica</emphasis>
|
||
(Germar, 1830)
|
||
</taxonomicName>
|
||
(TL:
|
||
<collectingRegion id="49E3F84F9B7CFFF2497EF9F3FC826313" box="[812,861,1565,1590]" country="Indonesia" name="Jawa Timur" pageId="37" pageNumber="230">Java</collectingRegion>
|
||
) because its fore wing also has a broad dark-coloured patch halfway from the base, but
|
||
<taxonomicName id="4C274D2E9B7CFFF2490CF9D1FC17637D" baseAuthorityName="Germar" baseAuthorityYear="1830" box="[862,968,1599,1624]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="230" phylum="Arthropoda" rank="species" species="conica">
|
||
<emphasis id="B953EABF9B7CFFF2490CF9D1FC17637D" box="[862,968,1599,1624]" italics="true" pageId="37" pageNumber="230">M. conica</emphasis>
|
||
</taxonomicName>
|
||
has the reddish postclypeus (vs black in the new species),narrower patch on the fore wing, smoothly rounded distal shoulder of the male pygofer (vs with a short triangular tooth in the new species), and a lobe-like process under the lateral clasper lobe (vs absent in the new species).
|
||
</paragraph>
|
||
<caption id="DF5866259B7FFFF14AF2FD2AFD2867DD" ID-DOI="http://doi.org/10.5281/zenodo.7737306" ID-Zenodo-Dep="7737306" httpUri="https://zenodo.org/record/7737306/files/figure.png" pageId="38" pageNumber="231" startId="38.[160,225,708,731]" targetBox="[164,1151,144,674]" targetPageId="38">
|
||
<paragraph id="8B9836AD9B7FFFF14AF2FD2AFD2867DD" blockId="38.[160,1155,708,760]" pageId="38" pageNumber="231">
|
||
<emphasis id="B953EABF9B7FFFF14AF2FD2AFED967FE" bold="true" box="[160,262,708,731]" pageId="38" pageNumber="231">Figure 16.</emphasis>
|
||
<taxonomicName id="4C274D2E9B7FFFF14B59FD2AFD4367FE" authority="Lee and Marshall" authorityName="Lee and Marshall" authorityYear="2023" box="[267,668,708,731]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="193" phylum="Arthropoda" rank="species" species="tenebrosa" status="sp.">
|
||
<emphasis id="B953EABF9B7FFFF14B59FD2AFE0F67FE" box="[267,464,708,731]" italics="true" pageId="38" pageNumber="231">Mogannia tenebrosa</emphasis>
|
||
Lee and Marshall
|
||
<taxonomicNameLabel id="A26057C49B7FFFF1482DFD2AFD4367FE" box="[639,668,708,731]" pageId="38" pageNumber="231" rank="species">sp.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov. Waveforms (A, C) of male calling song at two time scales, the former with accompanying spectrogram (B).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9836AD9B7FFFF14A92FCCAFD6866ED" blockId="38.[159,1156,804,1315]" pageId="38" pageNumber="231">
|
||
Also similar is
|
||
<taxonomicName id="4C274D2E9B7FFFF14B22FCCAFD286618" authority="Boulard, 2008" authorityName="Boulard" authorityYear="2008" box="[368,759,804,829]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="funettae">
|
||
<emphasis id="B953EABF9B7FFFF14B22FCCAFD966618" box="[368,585,804,829]" italics="true" pageId="38" pageNumber="231">Mogannia funettae</emphasis>
|
||
Boulard, 2008
|
||
</taxonomicName>
|
||
from northern
|
||
<collectingCountry id="F330763D9B7FFFF149E4FCCAFBC46618" box="[950,1051,804,829]" name="Thailand" pageId="38" pageNumber="231">Thailand</collectingCountry>
|
||
, but like
|
||
<emphasis id="B953EABF9B7FFFF14AF2FCA9FE506645" box="[160,399,839,864]" italics="true" pageId="38" pageNumber="231">
|
||
<taxonomicName id="4C274D2E9B7FFFF14AF2FCA9FED66645" baseAuthorityName="Germar" baseAuthorityYear="1830" box="[160,265,839,864]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="conica">M. conica</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C274D2E9B7FFFF14B41FCA9FE506645" authorityName="Boulard" authorityYear="2008" box="[275,399,839,864]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="funettae">M. funettae</taxonomicName>
|
||
</emphasis>
|
||
has the fore wing basal cell clear or tinged with bright colour rather than infuscated with dark brown to match the remainder of the basal fore wing (see
|
||
<bibRefCitation id="EFB64B5C9B7FFFF14AF2FC62FEEA6680" author="Boulard M" box="[160,309,908,933]" pageId="38" pageNumber="231" pagination="436" refId="ref19423" refString="Boulard M. 2013. The cicadas of Thailand, volume 2: taxonomy and sonic ethology. Manchester (United Kingdom): Siri Scientific Press; p. 436." type="book chapter" year="2013">Boulard 2013</bibRefCitation>
|
||
). As discussed below, the song of
|
||
<taxonomicName id="4C274D2E9B7FFFF148E5FC62FB5B6680" authority="Lee and Marshall" authorityName="Lee and Marshall" authorityYear="2023" box="[695,1156,908,933]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="193" phylum="Arthropoda" rank="species" species="tenebrosa" status="sp.">
|
||
<emphasis id="B953EABF9B7FFFF148E5FC62FC476680" box="[695,920,908,933]" italics="true" pageId="38" pageNumber="231">Mogannia tenebrosa</emphasis>
|
||
Lee and Marshall
|
||
<taxonomicNameLabel id="A26057C49B7FFFF14E36FC62FB5B6680" box="[1124,1156,908,933]" pageId="38" pageNumber="231" rank="species">sp.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov. also suggests a relationship to
|
||
<taxonomicName id="4C274D2E9B7FFFF14866FC41FD6E66ED" authorityName="Boulard" authorityYear="2008" box="[564,689,943,968]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="funettae">
|
||
<emphasis id="B953EABF9B7FFFF14866FC41FD6E66ED" box="[564,689,943,968]" italics="true" pageId="38" pageNumber="231">M. funettae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7FFFF14A92FC3CFB5B6115" blockId="38.[159,1156,804,1315]" pageId="38" pageNumber="231">
|
||
This species is similar also to
|
||
<taxonomicName id="4C274D2E9B7FFFF14BB8FC3CFC5766CE" authority="Matsumura, 1907" authorityName="Matsumura" authorityYear="1907" box="[490,904,978,1003]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="formosana">
|
||
<emphasis id="B953EABF9B7FFFF14BB8FC3CFD1466CE" box="[490,715,978,1003]" italics="true" pageId="38" pageNumber="231">Mogannia formosana</emphasis>
|
||
Matsumura,1907
|
||
</taxonomicName>
|
||
because of the blackish body colour and the brownish patch on the fore wing, but
|
||
<taxonomicName id="4C274D2E9B7FFFF1497BFC1AFC646128" authorityName="Matsumura" authorityYear="1907" box="[809,955,1012,1037]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="formosana">
|
||
<emphasis id="B953EABF9B7FFFF1497BFC1AFC646128" box="[809,955,1012,1037]" italics="true" pageId="38" pageNumber="231">M. formosana</emphasis>
|
||
</taxonomicName>
|
||
has a much more brightly coloured patch on the fore wing and light colouration on the ventral side of the body.
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7FFFF14A92FBD4FB8B6006" blockId="38.[159,1156,804,1315]" pageId="38" pageNumber="231">
|
||
Two other
|
||
<taxonomicName id="4C274D2E9B7FFFF14B6BFBD4FE776176" authorityName="Amyot and Audinet-Serville" authorityYear="1843" box="[313,424,1082,1107]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953EABF9B7FFFF14B6BFBD4FE776176" box="[313,424,1082,1107]" italics="true" pageId="38" pageNumber="231">Mogannia</emphasis>
|
||
</taxonomicName>
|
||
previously recorded from the
|
||
<collectingCountry id="F330763D9B7FFFF148A9FBD4FCA96176" box="[763,886,1082,1107]" name="Philippines" pageId="38" pageNumber="231">Philippines</collectingCountry>
|
||
are
|
||
<taxonomicName id="4C274D2E9B7FFFF149F5FBD4FEE96150" authority="Distant, 1917" authorityName="Distant" authorityYear="1917" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="indigotea">
|
||
<emphasis id="B953EABF9B7FFFF149F5FBD4FB5B6176" box="[935,1156,1082,1107]" italics="true" pageId="38" pageNumber="231">Mogannia indigotea</emphasis>
|
||
Distant, 1917
|
||
</taxonomicName>
|
||
(TL:
|
||
<collectingCountry id="F330763D9B7FFFF14B22FBB2FE616150" box="[368,446,1116,1141]" name="Vietnam" pageId="38" pageNumber="231">Tonkin</collectingCountry>
|
||
) and
|
||
<taxonomicName id="4C274D2E9B7FFFF14850FBB2FCA86150" authority="(Signoret, 1847)" baseAuthorityName="Signoret" baseAuthorityYear="1847" box="[514,887,1116,1141]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="viridis">
|
||
<emphasis id="B953EABF9B7FFFF14850FBB2FD656150" box="[514,698,1116,1141]" italics="true" pageId="38" pageNumber="231">Mogannia viridis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B5C9B7FFFF1489BFBB2FCAF6150" author="Signoret MV" box="[713,880,1116,1141]" pageId="38" pageNumber="231" pagination="293 - 296" refId="ref20857" refString="Signoret MV. 1847. Description de deux Hemipteres Homopteres, tribu des Octicelles groupe des cicadides. Annales de la Societe entomologique de France. 1847: 293 - 296." type="journal article" year="1847">Signoret, 1847</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(TL: Java) although we suspect that they are not distributed in the
|
||
<collectingCountry id="F330763D9B7FFFF148C1FB91FCCE61BD" box="[659,785,1151,1176]" name="Philippines" pageId="38" pageNumber="231">Philippines</collectingCountry>
|
||
. Both differ from
|
||
<taxonomicName id="4C274D2E9B7FFFF1498AFB91FBB961BD" box="[984,1126,1151,1176]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="B953EABF9B7FFFF1498AFB91FBB961BD" box="[984,1126,1151,1176]" italics="true" pageId="38" pageNumber="231">M. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
in the main body colour (
|
||
<taxonomicName id="4C274D2E9B7FFFF14BC9FB4CFDFD619E" authorityName="Distant" authorityYear="1917" box="[411,546,1186,1211]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="indigotea">
|
||
<emphasis id="B953EABF9B7FFFF14BC9FB4CFDFD619E" box="[411,546,1186,1211]" italics="true" pageId="38" pageNumber="231">M. indigotea</emphasis>
|
||
</taxonomicName>
|
||
is dark indigo blue, and
|
||
<taxonomicName id="4C274D2E9B7FFFF1497CFB4CFC52619E" baseAuthorityName="Signoret" baseAuthorityYear="1847" box="[814,909,1186,1211]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="viridis">
|
||
<emphasis id="B953EABF9B7FFFF1497CFB4CFC52619E" box="[814,909,1186,1211]" italics="true" pageId="38" pageNumber="231">M. viridis</emphasis>
|
||
</taxonomicName>
|
||
green to ochraceous). While the dark body colour of
|
||
<taxonomicName id="4C274D2E9B7FFFF14BAAFB2AFD5E61F8" authorityName="Distant" authorityYear="1917" box="[504,641,1220,1245]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="indigotea">
|
||
<emphasis id="B953EABF9B7FFFF14BAAFB2AFD5E61F8" box="[504,641,1220,1245]" italics="true" pageId="38" pageNumber="231">M. indigotea</emphasis>
|
||
</taxonomicName>
|
||
could be confused in poorly preserved specimens, the fore wing colour pattern is different, with some sections of the fore wing hyaline or tinted in
|
||
<taxonomicName id="4C274D2E9B7FFFF14B2CFAE4FE3A6006" authorityName="Distant" authorityYear="1917" box="[382,485,1290,1315]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="indigotea">
|
||
<emphasis id="B953EABF9B7FFFF14B2CFAE4FE3A6006" box="[382,485,1290,1315]" italics="true" pageId="38" pageNumber="231">indigotea</emphasis>
|
||
</taxonomicName>
|
||
but fully infuscated in
|
||
<taxonomicName id="4C274D2E9B7FFFF148B8FAE4FC8A6006" box="[746,853,1290,1315]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="B953EABF9B7FFFF148B8FAE4FC8A6006" box="[746,853,1290,1315]" italics="true" pageId="38" pageNumber="231">tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFB64B5C9B7FFFF149C3FAE4FB986006" author="Chen X & Yang M & Wei C" box="[913,1095,1290,1315]" pageId="38" pageNumber="231" pagination="1 - 35" refId="ref19490" refString="Chen X, Yang M, Wei C 2012. Review of the cicada genus Mogannia Amyot & Serville from China, with descriptions of three new species (Hemiptera: Cicadidae). Zootaxa. 3568 (1): 1 - 35." type="journal article" year="2012">Chen et al. 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D65269B7FFFF04AF2FAA2FB82642A" lastPageId="39" lastPageNumber="232" pageId="38" pageNumber="231" type="description">
|
||
<paragraph id="8B9836AD9B7FFFF14AF2FAA2FEB86043" blockId="38.[159,1155,1356,1728]" box="[160,359,1356,1382]" pageId="38" pageNumber="231">
|
||
<emphasis id="B953EABF9B7FFFF14AF2FAA2FEB86043" bold="true" box="[160,359,1356,1382]" pageId="38" pageNumber="231">
|
||
<emphasis id="B953EABF9B7FFFF14AF2FAA2FF036043" bold="true" box="[160,220,1356,1382]" italics="true" pageId="38" pageNumber="231">Song</emphasis>
|
||
(
|
||
<figureCitation id="131C2A289B7FFFF14ABEFAA2FE806043" box="[236,351,1356,1382]" captionStart="Figure 16" captionStartId="38.[160,225,708,731]" captionTargetBox="[164,1151,144,674]" captionTargetId="figure-373@38.[162,1154,142,676]" captionTargetPageId="38" captionText="Figure 16. Mogannia tenebrosa Lee and Marshall sp. nov. Waveforms (A, C) of male calling song at two time scales, the former with accompanying spectrogram (B)." figureDoi="http://doi.org/10.5281/zenodo.7737306" httpUri="https://zenodo.org/record/7737306/files/figure.png" pageId="38" pageNumber="231">Figure 16</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7FFFF14AF2FA81FC1F63E5" blockId="38.[159,1155,1356,1728]" pageId="38" pageNumber="231">
|
||
The song of the
|
||
<typeStatus id="549C880F9B7FFFF14B09FA81FE6360AD" box="[347,444,1391,1416]" pageId="38" pageNumber="231" type="holotype">holotype</typeStatus>
|
||
male was recorded and is illustrated here. The sound is a highpitched staccato buzz that continues indefinitely (at least 62s in our recording). Syllables containing approximately six pulses each are closely grouped or coalesced into echemes comprising four syllables, with the beginning and ending pulses of each syllable sometimes overlapping. The syllables are produced at about 200/s, and the echemes at about 20/s. There is minor amplitude modulation that helps to define the syllables,but otherwise sound intensity and frequency are not regularly modulated.The frequency spectrum is broad,reaching from at least 8 kHz to above 20 kHz, but most energy is concentrated within the range 8–12 kHz. The dominant frequency is about 10 kHz.Males were observed to sing while perching on the upper sides of leaves, as noted by
|
||
<bibRefCitation id="EFB64B5C9B7FFFF14B98F949FDB763E5" author="Boulard M" box="[458,616,1703,1728]" pageId="38" pageNumber="231" pagination="436" refId="ref19423" refString="Boulard M. 2013. The cicadas of Thailand, volume 2: taxonomy and sonic ethology. Manchester (United Kingdom): Siri Scientific Press; p. 436." type="book chapter" year="2013">Boulard (2013)</bibRefCitation>
|
||
for other
|
||
<taxonomicName id="4C274D2E9B7FFFF14880F949FCE263E5" authorityName="Amyot and Audinet-Serville" authorityYear="1843" box="[722,829,1703,1728]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="231" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953EABF9B7FFFF14880F949FCE263E5" box="[722,829,1703,1728]" italics="true" pageId="38" pageNumber="231">Mogannia</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F330763D9B7FFFF1490CF949FC6263E5" box="[862,957,1703,1728]" name="Thailand" pageId="38" pageNumber="231">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9836AD9B7EFFF04A92FF60FB82642A" blockId="39.[160,1156,142,271]" pageId="39" pageNumber="232">
|
||
No songs have been published previously for the genus
|
||
<taxonomicName id="4C274D2E9B7EFFF04910FF60FC6E6582" authorityName="Amyot and Audinet-Serville" authorityYear="1843" box="[834,945,142,167]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="232" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953EABF9B7EFFF04910FF60FC6E6582" box="[834,945,142,167]" italics="true" pageId="39" pageNumber="232">Mogannia</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<collectingCountry id="F330763D9B7EFFF04E51FF60FB5E6582" box="[1027,1153,142,167]" name="Philippines" pageId="39" pageNumber="232">Philippines</collectingCountry>
|
||
. The song of
|
||
<taxonomicName id="4C274D2E9B7EFFF04B7BFF5FFE7A65EF" authorityName="Boulard" authorityYear="2008" box="[297,421,177,202]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="232" phylum="Arthropoda" rank="species" species="funettae">
|
||
<emphasis id="B953EABF9B7EFFF04B7BFF5FFE7A65EF" box="[297,421,177,202]" italics="true" pageId="39" pageNumber="232">M. funettae</emphasis>
|
||
</taxonomicName>
|
||
, as shown in
|
||
<bibRefCitation id="EFB64B5C9B7EFFF04868FF5FFD0165EF" author="Boulard M" box="[570,734,177,202]" pageId="39" pageNumber="232" pagination="436" refId="ref19423" refString="Boulard M. 2013. The cicadas of Thailand, volume 2: taxonomy and sonic ethology. Manchester (United Kingdom): Siri Scientific Press; p. 436." type="book chapter" year="2013">Boulard (2013)</bibRefCitation>
|
||
, is very similar to that of
|
||
<taxonomicName id="4C274D2E9B7EFFF049A0FF5FFBA165EF" box="[1010,1150,177,202]" class="Insecta" family="Cicadidae" genus="Mogannia" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="232" phylum="Arthropoda" rank="species" species="tenebrosa">
|
||
<emphasis id="B953EABF9B7EFFF049A0FF5FFBA165EF" box="[1010,1150,177,202]" italics="true" pageId="39" pageNumber="232">M. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
, differing mainly in having a somewhat faster echeme repetition rate of about 28/s and inter-echeme intervals that are about the same duration as the echemes themselves.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |