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<mods:title id="E9BF345A4EE80D32BD3D0B6D329FA624">Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul &amp; Jocqué, 2004 (Araneae: Zodariidae)</mods:title>
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<treatment id="038E87ACFFB50423FF26FC7F06FEA85B" ID-DOI="http://doi.org/10.5281/zenodo.7984001" ID-GBIF-Taxon="209663658" ID-Zenodo-Dep="7984001" LSID="urn:lsid:plazi:treatment:038E87ACFFB50423FF26FC7F06FEA85B" httpUri="http://treatment.plazi.org/id/038E87ACFFB50423FF26FC7F06FEA85B" lastPageNumber="396" pageId="15" pageNumber="396">
<subSubSection id="C33D6531FFB50423FF26FC7F017FAA21" box="[151,893,1014,1041]" pageId="15" pageNumber="396" type="nomenclature">
<paragraph id="8B9836BAFFB50423FF26FC7F017FAA21" blockId="15.[151,893,1014,1077]" box="[151,893,1014,1041]" pageId="15" pageNumber="396">
<heading id="D0D081D6FFB50423FF26FC7F017FAA21" bold="true" box="[151,893,1014,1041]" fontSize="11" level="1" pageId="15" pageNumber="396" reason="1">
<emphasis id="B953EAA8FFB50423FF26FC7F017FAA21" bold="true" box="[151,893,1014,1041]" pageId="15" pageNumber="396">
<taxonomicName id="4C274D39FFB50423FF26FC7F0022AA21" authority="Simon, 1905" authorityName="Simon" authorityYear="1905" box="[151,544,1014,1041]" class="Arachnida" family="Zodariidae" genus="Capheris" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="species" species="nitidiceps">
<emphasis id="B953EAA8FFB50423FF26FC7F0382AA20" bold="true" box="[151,384,1014,1041]" italics="true" pageId="15" pageNumber="396">Capheris nitidiceps</emphasis>
<bibRefCitation id="EFB64B4BFFB50423FE36FC7F0022AA21" author="Simon, E." box="[391,544,1014,1040]" pageId="15" pageNumber="396" pagination="160 - 180" refId="ref11073" refString="Simon, E. (1905) Voyage de M. Maurice Maindron dans l'Inde meridionale (mai a novembre 1901). 7 me Memoire. Arachnides (1 re partie). Annales de la Societe Entomologique de France, 74 (1 - 2), 160 - 180." type="journal article" year="1905">Simon, 1905</bibRefCitation>
</taxonomicName>
comb, rest., nomen dubium
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D6531FFB50423FF26FB920316AA04" box="[151,276,1051,1077]" pageId="15" pageNumber="396" type="description">
<paragraph id="8B9836BAFFB50423FF26FB920316AA04" blockId="15.[151,893,1014,1077]" box="[151,276,1051,1077]" pageId="15" pageNumber="396">
<figureCitation id="131C2A3FFFB50423FF26FB9202EEAA04" box="[151,236,1051,1077]" captionStart="FIGURE 12" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[156,1431,494,1881]" captionTargetId="figure-180@16.[151,1436,489,1886]" captionTargetPageId="16" captionText="FIGURE 12. Capheris nitidiceps Simon, 1905 comb. rest., holotype female (subadult) (AR-16434).A habitus, dorsal. B same, ventral. C prosoma showing sternum, ventral. D opisthosoma showing pre-epigyne, ventral. E original label found in the type vial. Scale bars: AB, 0.5 mm. Figures CD without scale provided. © MNHN, Paris." figureDoi="http://doi.org/10.5281/zenodo.7984081" httpUri="https://zenodo.org/record/7984081/files/figure.png" pageId="15" pageNumber="396">Figs 12</figureCitation>
,
<figureCitation id="131C2A3FFFB50423FF49FB920316AA04" box="[248,276,1051,1077]" captionStart="FIGURE 18" captionStartId="22.[151,250,1974,1998]" captionTargetBox="[225,1367,1107,1944]" captionTargetId="figure-58@22.[219,1371,1102,1952]" captionTargetPageId="22" captionText="FIGURE 18. Distribution of the genus Capheris Simon, 1893 in India. ● Capheris escheri Reimoser, 1934 comb. rest., ■ Capheris nitidiceps Simon, 1905 comb. rest., ▲ Capheris stillata Simon, 1905 comb. rest." figureDoi="http://doi.org/10.5281/zenodo.7984101" httpUri="https://zenodo.org/record/7984101/files/figure.png" pageId="15" pageNumber="396">18</figureCitation>
</paragraph>
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<paragraph id="8B9836BAFFB50423FF26FBD600A7AA46" blockId="15.[151,808,1119,1174]" box="[151,677,1119,1143]" pageId="15" pageNumber="396">
<treatmentCitationGroup id="AB371194FFB50423FF26FBD600A7AA46" box="[151,677,1119,1143]" pageId="15" pageNumber="396">
<treatmentCitation id="0A8610ABFFB50423FF26FBD6001AAA46" author="Simon, E." box="[151,536,1119,1143]" page="171" pageId="15" pageNumber="396" year="1905">
<taxonomicName id="4C274D39FFB50423FF26FBD6001AAA46" authority="Simon, 1905: 171" authorityName="Simon" authorityPageNumber="171" authorityYear="1905" box="[151,536,1119,1143]" class="Arachnida" family="Zodariidae" genus="Capheris" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="species" species="nitidiceps">
<emphasis id="B953EAA8FFB50423FF26FBD6035FAA46" box="[151,349,1119,1143]" italics="true" pageId="15" pageNumber="396">Capheris nitidiceps</emphasis>
<bibRefCitation id="EFB64B4BFFB50423FED2FBD6001AAA46" author="Simon, E." box="[355,536,1119,1143]" pageId="15" pageNumber="396" pagination="160 - 180" refId="ref11073" refString="Simon, E. (1905) Voyage de M. Maurice Maindron dans l'Inde meridionale (mai a novembre 1901). 7 me Memoire. Arachnides (1 re partie). Annales de la Societe Entomologique de France, 74 (1 - 2), 160 - 180." type="journal article" year="1905">Simon, 1905: 171</bibRefCitation>
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(subadult
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).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9836BAFFB50423FF26FBF7012AAAA7" blockId="15.[151,808,1119,1174]" box="[151,808,1150,1174]" pageId="15" pageNumber="396">
<treatmentCitationGroup id="AB371194FFB50423FF26FBF7012AAAA7" box="[151,808,1150,1174]" pageId="15" pageNumber="396">
<taxonomicName id="4C274D39FFB50423FF26FBF70353AAA7" ID-CoL="32ZNQ" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[151,337,1150,1174]" class="Arachnida" family="Zodariidae" genus="Cydrela" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="species" species="nitidiceps">
<emphasis id="B953EAA8FFB50423FF26FBF70353AAA7" box="[151,337,1150,1174]" italics="true" pageId="15" pageNumber="396">Cydrela nitidiceps</emphasis>
</taxonomicName>
<treatmentCitation id="0A8610ABFFB50423FEDAFBF70021AAA7" author="Jocque, R." box="[363,547,1150,1174]" page="123" pageId="15" pageNumber="396" year="2009">
<bibRefCitation id="EFB64B4BFFB50423FEDAFBF70021AAA7" author="Jocque, R." box="[363,547,1150,1174]" pageId="15" pageNumber="396" pagination="77 - 148" refId="ref10916" refString="Jocque, R. (2009) Some keep it short: on the radiation in the Afrotropical spider genera Capheris and Systenoplacis (Araneae, Zodariidae) without male pedipalp complexity increase. Journal of Afrotropical Zoology, 5, 77 - 148." type="journal article" year="2009">Jocqué, 2009: 123</bibRefCitation>
</treatmentCitation>
(transfer from
<taxonomicName id="4C274D39FFB50423FD0CFBF70118AAA7" ID-CoL="3GZ6" baseAuthorityName="Reimoser" baseAuthorityYear="1934" box="[701,794,1150,1174]" class="Arachnida" family="Zodariidae" genus="Capheris" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="B953EAA8FFB50423FD0CFBF70118AAA7" box="[701,794,1150,1174]" italics="true" pageId="15" pageNumber="396">Capheris</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33D6531FFB50423FF26FB4D075DAB33" pageId="15" pageNumber="396" type="materials_examined">
<paragraph id="8B9836BAFFB50423FF26FB4D075DAB33" blockId="15.[151,1437,1220,1643]" pageId="15" pageNumber="396">
<emphasis id="B953EAA8FFB50423FF26FB4D0346AAEF" bold="true" box="[151,324,1220,1246]" pageId="15" pageNumber="396">Type material.</emphasis>
<materialsCitation id="3B4F3CE7FFB50423FEFBFB4D075DAB33" ID-GBIF-Occurrence="4120748309" collectionCode="MNHN" collectorName="M. M. Maindron &amp; Paris" country="India" elevation="1591" latitude="11.333333" location="Coonoor" longLatPrecision="1295" longitude="76.8" municipality="Nilgiris" pageId="15" pageNumber="396" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<typeStatus id="549C8818FFB50423FEFBFB4D03B0AAEF" box="[330,434,1220,1246]" pageId="15" pageNumber="396" type="holotype">Holotype</typeStatus>
subadult
<specimenCount id="9D21FD33FFB50423FDACFB4C0030AAEE" box="[541,562,1221,1247]" pageId="15" pageNumber="396" type="female"></specimenCount>
from
<emphasis id="B953EAA8FFB50423FDC4FB4D00D3AAEF" bold="true" box="[629,721,1220,1246]" pageId="15" pageNumber="396">
<collectingCountry id="F330762AFFB50423FDC4FB4D00CEAAEF" box="[629,716,1220,1246]" name="India" pageId="15" pageNumber="396">INDIA</collectingCountry>
:
</emphasis>
<emphasis id="B953EAA8FFB50423FD66FB4D015BAAEF" box="[727,857,1220,1246]" italics="true" pageId="15" pageNumber="396">
<collectingRegion id="49E3F858FFB50423FD66FB4D015BAAEF" box="[727,857,1220,1246]" country="India" name="Tamil Nadu" pageId="15" pageNumber="396">Tamil Nadu</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6BFCACC0FFB50423FCD7FB4D01BCAAEF" box="[870,958,1220,1246]" pageId="15" pageNumber="396">Nilgiris</collectingMunicipality>
,
<location id="8EF86061FFB50423FC78FB4D062FAAEF" LSID="urn:lsid:plazi:treatment:038E87ACFFB50423FF26FC7F06FEA85B:8EF86061FFB50423FC78FB4D062FAAEF" box="[969,1069,1220,1246]" country="India" latitude="11.333333" longLatPrecision="1295" longitude="76.8" municipality="Nilgiris" name="Coonoor" pageId="15" pageNumber="396" stateProvince="Tamil Nadu">Coonoor</location>
,
<geoCoordinate id="EE13507DFFB50423FB86FB4D0694AAEF" box="[1079,1174,1220,1246]" degrees="11" direction="north" minutes="20" orientation="latitude" pageId="15" pageNumber="396" precision="925" value="11.333333">11°20N</geoCoordinate>
<geoCoordinate id="EE13507DFFB50423FB2DFB4D06FEAAEF" box="[1180,1276,1220,1246]" degrees="76" direction="east" minutes="48" orientation="longitude" pageId="15" pageNumber="396" precision="925" value="76.8">76°48E</geoCoordinate>
,
<elevation id="000AD189FFB50423FAB6FB4D0795AAEF" box="[1287,1431,1220,1247]" metricMagnitude="3" metricUnit="m" metricValue="1.591" pageId="15" pageNumber="396" unit="m" value="1591.0">
<quantity id="4CDF9B5FFFB50423FAB6FB4D0759AAEE" box="[1287,1371,1220,1247]" metricMagnitude="3" metricUnit="m" metricValue="1.591" pageId="15" pageNumber="396" unit="m" value="1591.0">1591 m</quantity>
a.s.l.
</elevation>
, date unknown, leg.
<collectorName id="26D2536CFFB50423FEC2FB600030AB33" box="[371,562,1256,1283]" pageId="15" pageNumber="396">M. M. Maindron</collectorName>
, repository
<collectionCode id="ED36AE7FFFB50423FD07FB60010CAB32" box="[694,782,1257,1283]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="396">MNHN</collectionCode>
,
<collectorName id="26D2536CFFB50423FCABFB610150AB33" box="[794,850,1256,1282]" pageId="15" pageNumber="396">Paris</collectorName>
(AR-16434), examined based on photographs.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6531FFB50423FF76FA850708ABA2" pageId="15" pageNumber="396" type="discussion">
<paragraph id="8B9836BAFFB50423FF76FA850708ABA2" blockId="15.[151,1437,1220,1643]" pageId="15" pageNumber="396">
<emphasis id="B953EAA8FFB50423FF76FA850339AB17" bold="true" box="[199,315,1292,1318]" pageId="15" pageNumber="396">Remarks.</emphasis>
This species has to be considered a
<emphasis id="B953EAA8FFB50423FD7AFA840172AB17" box="[715,880,1292,1318]" italics="true" pageId="15" pageNumber="396">nomen dubium</emphasis>
due to the immature stage of the female
<typeStatus id="549C8818FFB50423FA82FA85079BAB17" box="[1331,1433,1292,1318]" pageId="15" pageNumber="396" type="holotype">holotype</typeStatus>
. The type locality of this species lies hardly 267 kms away from the type locality of
<taxonomicName id="4C274D39FFB50423FBE7FAB906CBAB7B" authorityName="Reimoser" authorityYear="1934" box="[1110,1225,1328,1354]" class="Arachnida" family="Zodariidae" genus="Capheris" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="species" species="escheri">
<emphasis id="B953EAA8FFB50423FBE7FAB906CBAB7B" box="[1110,1225,1328,1354]" italics="true" pageId="15" pageNumber="396">C. escheri</emphasis>
</taxonomicName>
that let suggests a possible synonymy of the latter species with it; however, confirmation requires the examination of adult specimens of
<taxonomicName id="4C274D39FFB50423FF07FAF10341ABA3" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[182,323,1400,1426]" class="Arachnida" family="Zodariidae" genus="Cydrela" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="species" species="nitidiceps">
<emphasis id="B953EAA8FFB50423FF07FAF10341ABA3" box="[182,323,1400,1426]" italics="true" pageId="15" pageNumber="396">C. nitidiceps</emphasis>
</taxonomicName>
from the type locality. Until adult specimens are found, it is treated as
<emphasis id="B953EAA8FFB50423FBECFAF00701ABA3" box="[1117,1283,1400,1426]" italics="true" pageId="15" pageNumber="396">nomen dubium</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6531FFB50423FF76FA1506FEA85B" pageId="15" pageNumber="396" type="distribution">
<paragraph id="8B9836BAFFB50423FF76FA150153AB87" blockId="15.[151,1437,1220,1643]" box="[199,849,1436,1463]" pageId="15" pageNumber="396">
<emphasis id="B953EAA8FFB50423FF76FA150363AB87" bold="true" box="[199,353,1436,1462]" pageId="15" pageNumber="396">Distribution.</emphasis>
<collectingCountry id="F330762AFFB50423FED6FA1503A3AB87" box="[359,417,1436,1462]" name="India" pageId="15" pageNumber="396">India</collectingCountry>
(
<collectingRegion id="49E3F858FFB50423FE1EFA150035AB87" box="[431,567,1436,1462]" country="India" name="Tamil Nadu" pageId="15" pageNumber="396">Tamil Nadu</collectingRegion>
) (
<bibRefCitation id="EFB64B4BFFB50423FDFFFA1500DBAB86" author="Simon, E." box="[590,729,1436,1463]" pageId="15" pageNumber="396" pagination="160 - 180" refId="ref11073" refString="Simon, E. (1905) Voyage de M. Maurice Maindron dans l'Inde meridionale (mai a novembre 1901). 7 me Memoire. Arachnides (1 re partie). Annales de la Societe Entomologique de France, 74 (1 - 2), 160 - 180." type="journal article" year="1905">Simon 1905</bibRefCitation>
) (
<figureCitation id="131C2A3FFFB50423FD41FA150146AB87" box="[752,836,1436,1462]" captionStart="FIGURE 18" captionStartId="22.[151,250,1974,1998]" captionTargetBox="[225,1367,1107,1944]" captionTargetId="figure-58@22.[219,1371,1102,1952]" captionTargetPageId="22" captionText="FIGURE 18. Distribution of the genus Capheris Simon, 1893 in India. ● Capheris escheri Reimoser, 1934 comb. rest., ■ Capheris nitidiceps Simon, 1905 comb. rest., ▲ Capheris stillata Simon, 1905 comb. rest." figureDoi="http://doi.org/10.5281/zenodo.7984101" httpUri="https://zenodo.org/record/7984101/files/figure.png" pageId="15" pageNumber="396">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8B9836BAFFB50423FF76FA4906FEA85B" blockId="15.[151,1437,1220,1643]" pageId="15" pageNumber="396">
<emphasis id="B953EAA8FFB50423FF76FA490010ABEB" bold="true" box="[199,530,1472,1498]" pageId="15" pageNumber="396">Justification of the transfer.</emphasis>
This species was originally placed under
<taxonomicName id="4C274D39FFB50423FC57FA4906F0ABEA" authority="(Simon 1905)" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[998,1266,1472,1499]" class="Arachnida" family="Zodariidae" genus="Capheris" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="B953EAA8FFB50423FC57FA49064FABEB" box="[998,1101,1472,1498]" italics="true" pageId="15" pageNumber="396">Capheris</emphasis>
(
<bibRefCitation id="EFB64B4BFFB50423FBEDFA4906EBABEA" author="Simon, E." box="[1116,1257,1472,1499]" pageId="15" pageNumber="396" pagination="160 - 180" refId="ref11073" refString="Simon, E. (1905) Voyage de M. Maurice Maindron dans l'Inde meridionale (mai a novembre 1901). 7 me Memoire. Arachnides (1 re partie). Annales de la Societe Entomologique de France, 74 (1 - 2), 160 - 180." type="journal article" year="1905">Simon 1905</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="EFB64B4BFFB50423FB4CFA49079EABEB" author="Jocque, R." box="[1277,1436,1472,1498]" pageId="15" pageNumber="396" pagination="77 - 148" refId="ref10916" refString="Jocque, R. (2009) Some keep it short: on the radiation in the Afrotropical spider genera Capheris and Systenoplacis (Araneae, Zodariidae) without male pedipalp complexity increase. Journal of Afrotropical Zoology, 5, 77 - 148." type="journal article" year="2009">Jocqué (2009)</bibRefCitation>
erroneously transferred it to
<taxonomicName id="4C274D39FFB50423FE6DFA6D0037ABCF" baseAuthorityName="Jocque" baseAuthorityYear="2009" box="[476,565,1508,1534]" class="Arachnida" family="Zodariidae" genus="Cydrela" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="B953EAA8FFB50423FE6DFA6D0037ABCF" box="[476,565,1508,1534]" italics="true" pageId="15" pageNumber="396">Cydrela</emphasis>
</taxonomicName>
, without proper justification. An examination of the
<typeStatus id="549C8818FFB50423FB21FA6D06F0ABCF" box="[1168,1266,1508,1534]" pageId="15" pageNumber="396" type="holotype">holotype</typeStatus>
of this species showed that it has diagnostic features of
<taxonomicName id="4C274D39FFB50423FDC5F9810137A813" box="[628,821,1544,1570]" pageId="15" pageNumber="396">
<emphasis id="B953EAA8FFB50423FDC5F98100D9A813" box="[628,731,1544,1570]" italics="true" pageId="15" pageNumber="396">Capheris</emphasis>
species
</taxonomicName>
including ALEs almost touching, situated in front of AMEs, straight PER, sternum shied-shaped, anterolaterally with indentations accommodating endites, and very stout palps (
<figureCitation id="131C2A3FFFB50423FEAEF9D903AEA85B" box="[287,428,1616,1642]" captionStart="FIGURE 12" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[156,1431,494,1881]" captionTargetId="figure-180@16.[151,1436,489,1886]" captionTargetPageId="16" captionText="FIGURE 12. Capheris nitidiceps Simon, 1905 comb. rest., holotype female (subadult) (AR-16434).A habitus, dorsal. B same, ventral. C prosoma showing sternum, ventral. D opisthosoma showing pre-epigyne, ventral. E original label found in the type vial. Scale bars: AB, 0.5 mm. Figures CD without scale provided. © MNHN, Paris." figureDoi="http://doi.org/10.5281/zenodo.7984081" httpUri="https://zenodo.org/record/7984081/files/figure.png" pageId="15" pageNumber="396">Figs 12AC</figureCitation>
). Based on these observations, the transfer of
<taxonomicName id="4C274D39FFB50423FC1FF9D9063EA85B" baseAuthorityName="Simon" baseAuthorityYear="1905" box="[942,1084,1616,1643]" class="Arachnida" family="Zodariidae" genus="Cydrela" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="396" phylum="Arthropoda" rank="species" species="nitidiceps">
<emphasis id="B953EAA8FFB50423FC1FF9D901C3A85B" box="[942,961,1616,1642]" italics="true" pageId="15" pageNumber="396">C</emphasis>
.
<emphasis id="B953EAA8FFB50423FC7EF9D9063EA85B" box="[975,1084,1616,1642]" italics="true" pageId="15" pageNumber="396">nitidiceps</emphasis>
</taxonomicName>
is fully justified.
</paragraph>
</subSubSection>
</treatment>
</document>