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<document id="D3FB8D61141F4BD3F83F8897D6C86F3A" ID-DOI="10.11646/zootaxa.5152.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6630873" ID-ZooBank="92E9DD85-6CC6-4602-BD7C-C51F49CEEF47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654852147308" checkinUser="plazi" docAuthor="Márquez, Juan &amp; Asiain, Julieta" docDate="2022" docId="038E878977037F2E81AFA3A7FAEBF8CD" docLanguage="en" docName="zootaxa.5152.1.1.pdf" docOrigin="Zootaxa 5152 (1)" docSource="http://zoobank.org/92e9dd85-6cc6-4602-bd7c-c51f49ceef47" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Belonuchus tenuistylus Márquez &amp; Asiain 2022, sp. nov." docType="treatment" docUuid="49D6419A-A9EE-4C63-893A-C70F026C193E" docUuidSource="ZooBank" docVersion="3" lastPageNumber="46" masterDocId="FFB7FFF1772F7F038138A63CFF9FFFE4" masterDocTitle="Taxonomy of the Mexican species of Belonuchus Nordmann (Coleoptera: Staphylinidae)" masterLastPageNumber="129" masterPageNumber="1" pageNumber="45" updateTime="1699335993931" updateUser="plazi">
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<mods:title id="D4532584E3BC1F1C4C0087A021FF9A36">Taxonomy of the Mexican species of Belonuchus Nordmann (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="90A80743393636AB82CA498B9EB258FF">Márquez, Juan</mods:namePart>
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<mods:namePart id="C86F4675C2C40EEBFD1F07756C353AF3">Asiain, Julieta</mods:namePart>
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<mods:title id="BE2199B4B07D0E381AD79F3CBB8BA5EE">Zootaxa</mods:title>
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<mods:date id="5C788DC0FB0CD76D5CB0400EA7C4DA54">2022</mods:date>
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<paragraph id="8B98369F77037F2F81AFA3A7FD88FA52" blockId="44.[151,1057,1435,1535]" box="[151,535,1435,1462]" pageId="44" pageNumber="45">
<heading id="D0D081F377037F2F81AFA3A7FD88FA52" bold="true" box="[151,535,1435,1462]" fontSize="11" level="2" pageId="44" pageNumber="45" reason="4">
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<taxonomicName id="4C274D1C77037F2F81AFA3A7FE2DFA52" ID-CoL="9GK5F" authority="Márquez &amp; Asiain, 2022" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[151,434,1435,1462]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tenuistylus" status="sp. nov.">
<emphasis id="B953EA8D77037F2F81AFA3A7FE2DFA52" bold="true" box="[151,434,1435,1462]" italics="true" pageId="44" pageNumber="45">Belonuchus tenuistylus</emphasis>
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<taxonomicNameLabel id="A26057F677037F2F8081A3A0FD88FA52" box="[441,535,1436,1462]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8B98369F77037F2F81AFA3FCFBBEFA3E" blockId="44.[151,1057,1435,1535]" box="[151,1057,1472,1498]" pageId="44" pageNumber="45">http://zoobank.org/NomenclaturalActs/49d6419a-a9ee-4c63-893a-c70f026c193e</paragraph>
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<subSubSection id="C33D651477037F2E81AFA3D8FE32FAB1" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="description">
<paragraph id="8B98369F77037F2F81AFA3D8FD88FA1A" blockId="44.[151,1057,1435,1535]" box="[151,535,1508,1535]" pageId="44" pageNumber="45">
<figureCitation id="131C2A1A77037F2F81AFA3D8FF6DFA1A" box="[151,242,1508,1534]" captionStart="FIGURE 4" captionStartId="98.[151,250,1720,1744]" captionTargetBox="[170,1410,196,1684]" captionTargetId="figure-25@98.[161,1426,181,1696]" captionTargetPageId="98" captionText="FIGURE 4. Dorsal view of males of: a) Belonuchus tenuistylus (holotype), b) B. tepoztecus (paratype), c) B. trochanterinus, d) B. zunilensis (lectotype), e) B. bidens (syntype, NHM©), f) B. bidens, g) B. erichsoni, h) B. jalappensis. Scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630889" httpUri="https://zenodo.org/record/6630889/files/figure.png" pageId="44" pageNumber="45">Figs. 4a</figureCitation>
,
<figureCitation id="131C2A1A77037F2F81C6A3D8FEB5FA1A" box="[254,298,1508,1534]" captionStart="FIGURE 13" captionStartId="107.[151,250,1930,1954]" captionTargetBox="[165,1424,192,1892]" captionTargetId="figure-19@107.[151,1436,181,1907]" captionTargetPageId="107" captionText="FIGURE 13. Ventral view of the left posterior leg of males of: a) B. trochanterinus, b) B. tenuistylus, c) B. zunilensis, d) B. rufipennis, e) B. mixtecus, f) B. pollens, g) B. ephippiatus, h) B. aenigmaticus, i) B. iridescens, j) B. viridipennis. Tarsomeres of posterior leg of males of: k) B. colon, l) B. ephippiatus. Yellow ovals: median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6630917" httpUri="https://zenodo.org/record/6630917/files/figure.png" pageId="44" pageNumber="45">13b</figureCitation>
,
<figureCitation id="131C2A1A77037F2F800EA3D8FEF5FA1B" box="[310,362,1508,1535]" captionStart="FIGURE 15" captionStartId="109.[151,250,1821,1845]" captionTargetBox="[175,1417,182,1784]" captionTargetId="figure-19@109.[160,1427,181,1797]" captionTargetPageId="109" captionText="FIGURE 15. Male pregenital sternite of: a) Belonuchus alternans, b) B. oxyporinus, c) B. tepoztecus, d) B. jalappensis, e) B. iridescens, f) B. platypterus, g) B. cyanipennis, h) B. apiciventris, i) B. linuzensis, j) B. metafemoralis, k) B. rufisternus, l) B. tenuistylus, m) B. bidens, n) B. pollens, o) B. longitarsus." figureDoi="http://doi.org/10.5281/zenodo.6630921" httpUri="https://zenodo.org/record/6630921/files/figure.png" pageId="44" pageNumber="45">15l</figureCitation>
,
<figureCitation id="131C2A1A77037F2F804EA3D8FE2EFA1A" box="[374,433,1508,1534]" captionStart="FIGURE 18" captionStartId="112.[151,250,1799,1823]" captionTargetBox="[174,1411,182,1767]" captionTargetId="figure-25@112.[173,1413,181,1775]" captionTargetPageId="112" captionText="FIGURE 18. Male genital sternite of: a) Belonuchus altenans, b) B. apiciventris, c) B. cariniventris, d) B. colon, e) B. confusus, f) B. julietitae, g) B. magnistylus, h) B. linuzensis, i) B. metafemoralis, j) B. oxyporimimus, k) B. oxyporinus, l) B. pulcher, m) B. rufisternus, n) B. singularis, o) B. tenuistylus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630925" httpUri="https://zenodo.org/record/6630925/files/figure.png" pageId="44" pageNumber="45">18o</figureCitation>
,
<figureCitation id="131C2A1A77037F2F8084A3D8FE7BFA1A" box="[444,484,1508,1534]" captionStart="FIGURE 22" captionStartId="116.[151,250,1894,1918]" captionTargetBox="[162,1427,182,1859]" captionTargetId="figure-25@116.[151,1436,181,1871]" captionTargetPageId="116" captionText="FIGURE 22. Ventral (left) and lateral (right) view of the aedeagus of: a) Belonuchus oxyporimimus, b) B. oxyporinus, c) B. pulcher, d) B. rufisternus, e) B. singularis, f) B. tenuistylus, g) B. tepoztecus (at center detail of the paramere), h) B. trochanterinus, i) B. zunilensis. Scale bar = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6630937" httpUri="https://zenodo.org/record/6630937/files/figure.png" pageId="44" pageNumber="45">22f</figureCitation>
,
<figureCitation id="131C2A1A77037F2F80D7A3D8FD88FA1A" box="[495,535,1508,1534]" captionStart="FIGURE 28" captionStartId="122.[151,250,1691,1715]" captionTargetBox="[155,1424,191,1658]" captionTargetId="figure-25@122.[151,1436,181,1667]" captionTargetPageId="122" captionText="FIGURE 28. Geographical distribution in Mexico and Guatemala of: a) Belonuchus colon, B. tenuistylus and B. trochanterinus; b) B. oxyporimimus, B. pulcher and B. juanmorronei." figureDoi="http://doi.org/10.5281/zenodo.6630949" httpUri="https://zenodo.org/record/6630949/files/figure.png" pageId="44" pageNumber="45">28a</figureCitation>
</paragraph>
<paragraph id="8B98369F77037F2F81AFA010FBEDF956" blockId="44.[151,1437,1580,2039]" pageId="44" pageNumber="45">
Total body length
<quantity id="4CDF9B7A77037F2F8067A010FE5AF9A3" box="[351,453,1580,1607]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="44" pageNumber="45" unit="mm" value="10.8">10.8 mm</quantity>
. Black on head, antennomeres 13, borders of mandibles, pronotum, scutellum, posterior half of prosternum, entire sixth and anterior half of fifth visible abdominal segments, and abdominal styli. Reddish on elytra, legs, anterior half of prosternum, posterior half of mesoventrite, entire metaventrite, visible abdominal segments 14 and anterior half of fifth. Brown on antennomeres 411 and mesoventrite.
</paragraph>
<paragraph id="8B98369F77037F2F81FFA080FAF1F84B" blockId="44.[151,1437,1580,2039]" pageId="44" pageNumber="45">
<emphasis id="B953EA8D77037F2F81FFA080FE98F932" bold="true" box="[199,263,1724,1750]" pageId="44" pageNumber="45">Head</emphasis>
: transverse, slightly convex at sides and backward; ratio length/width of head 0.73. Dorsal surface with sulcate longitudinal midline visible in anterior half of head; front foveate in a small portion near to anterior margin. Eyes nearly 0.5 times the cephalic lateral length, not protruding laterally. Antennomeres 46 elongate, 710 almost as long as wide. Mandibles 1.5 times longer than head; each with two widely separated teeth (basal and middle) of moderate size; mandibular channnel with external margin expanded at basal third of its length, leaving a wide space between this and internal margin, which is carinate and ends near level of middle tooth. Apical palpomere of maxillary and labial palpi nearly 1.25 times longer than preapical palpomere. Head 1.29 times wider than pronotum.
</paragraph>
<paragraph id="8B98369F77037F2E81FFA184FF68FE35" blockId="44.[151,1437,1580,2039]" lastBlockId="45.[151,1437,151,1833]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="B953EA8D77037F2F81FFA184FEBFF836" bold="true" box="[199,288,1976,2002]" pageId="44" pageNumber="45">Thorax</emphasis>
: pronotum with five punctures in the left and seven in the right dorsal row; pronotum slightly longer than wide (ratio 1.1) and almost as wide at anterior corners as at posterior corners (ratio 1.07). Scutellum with punctures slightly denser than on elytra, its margins smooth. Elytra with punctures moderately dense, with fine setae. Prosternum moderately elevated behind anterior margin. Intercoxal process of mesoventrite shield-shaped; transverse discal ridge clearly visible, broadly U-shaped or V-shaped, slightly acute posteriorly, meeting margin of intercoxal process laterally. Profemur with external row of spines initiating a little before half of its length and not reaching apex, and not reaching level of femur where some apical spines of internal margin begin, internal margin only visible in this anterior portion; spines of moderate size, similar to some spines of protibiae. Metatrochanter modified like a small hook that does not exceed 1/4 of length of metafemur, hook curved only near apex (
<figureCitation id="131C2A1A77027F2E8415A753FA11FE6E" box="[1325,1422,367,394]" captionStart="FIGURE 13" captionStartId="107.[151,250,1930,1954]" captionTargetBox="[165,1424,192,1892]" captionTargetId="figure-19@107.[151,1436,181,1907]" captionTargetPageId="107" captionText="FIGURE 13. Ventral view of the left posterior leg of males of: a) B. trochanterinus, b) B. tenuistylus, c) B. zunilensis, d) B. rufipennis, e) B. mixtecus, f) B. pollens, g) B. ephippiatus, h) B. aenigmaticus, i) B. iridescens, j) B. viridipennis. Tarsomeres of posterior leg of males of: k) B. colon, l) B. ephippiatus. Yellow ovals: median longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.6630917" httpUri="https://zenodo.org/record/6630917/files/figure.png" pageId="45" pageNumber="46">Fig. 13b</figureCitation>
); metafemur without curvature in area of overlap with hook; metatibia straight. Tarsomeres slender, pooly flattened dorsally.
</paragraph>
<paragraph id="8B98369F77027F2E81FFA7E6FBBBFD29" blockId="45.[151,1437,151,1833]" pageId="45" pageNumber="46">
<emphasis id="B953EA8D77027F2E81FFA7E6FEA3FE10" bold="true" box="[199,316,474,500]" pageId="45" pageNumber="46">Abdomen</emphasis>
: first three visible tergites with posterior basal transverse carina well developed; depressed adjacent area with wide-superficial punctures moderately dense, which are missing or only sparse on remaining surface of these tergites, as well as following tergites, which have fine, sparse punctures. Visible sternites 14 with wide-superficial punctures in anterior half, and with fine, sparse punctures on remaining surface, as well as on remaining sternites. Male pregenital sternite weakly emarginate at posterior margin (
<figureCitation id="131C2A1A77027F2E82DDA457FBCEFD61" box="[997,1105,618,645]" captionStart="FIGURE 15" captionStartId="109.[151,250,1821,1845]" captionTargetBox="[175,1417,182,1784]" captionTargetId="figure-19@109.[160,1427,181,1797]" captionTargetPageId="109" captionText="FIGURE 15. Male pregenital sternite of: a) Belonuchus alternans, b) B. oxyporinus, c) B. tepoztecus, d) B. jalappensis, e) B. iridescens, f) B. platypterus, g) B. cyanipennis, h) B. apiciventris, i) B. linuzensis, j) B. metafemoralis, k) B. rufisternus, l) B. tenuistylus, m) B. bidens, n) B. pollens, o) B. longitarsus." figureDoi="http://doi.org/10.5281/zenodo.6630921" httpUri="https://zenodo.org/record/6630921/files/figure.png" pageId="45" pageNumber="46">Fig. 15l</figureCitation>
). Genital sternite moderately large (2.87 times longer than wide), moderately asymmetrical, anterior portion occupying 33% and posterior portion 67% of its length; apical emargination deep (
<figureCitation id="131C2A1A77027F2E83B6A48FFD60FD29" box="[654,767,691,717]" captionStart="FIGURE 18" captionStartId="112.[151,250,1799,1823]" captionTargetBox="[174,1411,182,1767]" captionTargetId="figure-25@112.[173,1413,181,1775]" captionTargetPageId="112" captionText="FIGURE 18. Male genital sternite of: a) Belonuchus altenans, b) B. apiciventris, c) B. cariniventris, d) B. colon, e) B. confusus, f) B. julietitae, g) B. magnistylus, h) B. linuzensis, i) B. metafemoralis, j) B. oxyporimimus, k) B. oxyporinus, l) B. pulcher, m) B. rufisternus, n) B. singularis, o) B. tenuistylus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630925" httpUri="https://zenodo.org/record/6630925/files/figure.png" pageId="45" pageNumber="46">Fig. 18o</figureCitation>
). Abdominal styli slender.
</paragraph>
<paragraph id="8B98369F77027F2E81FFA4EAFCECFCF2" blockId="45.[151,1437,151,1833]" pageId="45" pageNumber="46">
<emphasis id="B953EA8D77027F2E81FFA4EAFEA5FD14" bold="true" box="[199,314,726,752]" pageId="45" pageNumber="46">Aedeagus</emphasis>
: length
<quantity id="4CDF9B7A77027F2E80ACA4EBFE73FD15" box="[404,492,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="45" pageNumber="46" unit="mm" value="1.3">1.3 mm</quantity>
; cone-shaped, apex pointed; sides becoming regularly wider from apex to base; basal half notably wider than apical half; internal sac visible (
<figureCitation id="131C2A1A77027F2E8230A4C7FCF8FCF2" box="[776,871,763,790]" captionStart="FIGURE 22" captionStartId="116.[151,250,1894,1918]" captionTargetBox="[162,1427,182,1859]" captionTargetId="figure-25@116.[151,1436,181,1871]" captionTargetPageId="116" captionText="FIGURE 22. Ventral (left) and lateral (right) view of the aedeagus of: a) Belonuchus oxyporimimus, b) B. oxyporinus, c) B. pulcher, d) B. rufisternus, e) B. singularis, f) B. tenuistylus, g) B. tepoztecus (at center detail of the paramere), h) B. trochanterinus, i) B. zunilensis. Scale bar = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6630937" httpUri="https://zenodo.org/record/6630937/files/figure.png" pageId="45" pageNumber="46">Fig. 22f</figureCitation>
).
</paragraph>
<paragraph id="8B98369F77027F2E81FFA522FB67FBBE" blockId="45.[151,1437,151,1833]" pageId="45" pageNumber="46">
<emphasis id="B953EA8D77027F2E81FFA522FED8FCDC" bold="true" box="[199,327,798,824]" pageId="45" pageNumber="46">Variability</emphasis>
. In measurements and ratios, the following variation was observed: total body length in males
<quantity id="4CDF9B7A77027F2E81AFA57EFF63FCB9" box="[151,252,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="45" pageNumber="46" unit="mm" value="10.5">10.5 mm</quantity>
(range
<quantity id="4CDF9B7A77027F2E806BA57FFE73FCB9" box="[339,492,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.055" metricValueMax="1.1199999999999999" metricValueMin="0.99" pageId="45" pageNumber="46" unit="mm" value="10.55" valueMax="11.2" valueMin="9.9">9.911.2 mm</quantity>
), in females
<quantity id="4CDF9B7A77027F2E8344A57FFD4CFCB9" box="[636,723,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="45" pageNumber="46" unit="mm" value="9.8">9.8 mm</quantity>
(range
<quantity id="4CDF9B7A77027F2E8212A57EFC5CFCB9" box="[810,963,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="45" pageNumber="46" unit="mm" value="9.5" valueMax="10.5" valueMin="8.5">8.510.5 mm</quantity>
); ratio length/width of head in males 0.77 (range 0.650.85), in females 0.88 (range 0.860.90); ratio mandibular length/cephalic length in males 1.25 (range 1.001.43), in females 0.86 (range 0.810.92); ratio cephalic width/pronotal width in males 1.23 (range 1.111.40), in females 1.06 (range 1.011.08); ratio length/width of pronotum in males 1.12 (range 1.031.20), in females 1.17 (range 1.121.21); ratio anterior width/posterior width of pronotum in males 1.05 (range 1.051.08), in females 1 (without variation). Apical antennomere, some times also subapical antennomere, lighter than the preceding ones. Most specimens have five punctures in each dorsal row of pronotum, with a variation of 5:6, 6:5 or as in the
<typeStatus id="549C883D77027F2E8459A227FF54FBBD" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
. The anterior half of the mesoventrite may be reddish as on its posterior half or darker (brown).
</paragraph>
<paragraph id="8B98369F77027F2E81FFA25EFE32FAB1" blockId="45.[151,1437,151,1833]" pageId="45" pageNumber="46">
<emphasis id="B953EA8D77027F2E81FFA25EFE55FB99" bold="true" box="[199,458,1122,1149]" pageId="45" pageNumber="46">Taxonomic comments</emphasis>
. Due to its color pattern, size and modification of the male metatrochanters, this species has been confused with
<taxonomicName id="4C274D1C77027F2E8098A2BBFDFDFB45" authorityName="Sharp" authorityYear="1885" box="[416,610,1159,1185]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="trochanterinus">
<emphasis id="B953EA8D77027F2E8098A2BBFDFDFB45" box="[416,610,1159,1185]" italics="true" pageId="45" pageNumber="46">B. trochanterinus</emphasis>
</taxonomicName>
in publications reporting the presence of
<taxonomicName id="4C274D1C77027F2E8513A2BBFB72FB45" authorityName="Sharp" authorityYear="1885" box="[1067,1261,1159,1185]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="trochanterinus">
<emphasis id="B953EA8D77027F2E8513A2BBFB72FB45" box="[1067,1261,1159,1185]" italics="true" pageId="45" pageNumber="46">B. trochanterinus</emphasis>
</taxonomicName>
in
<collectingCountry id="F330760F77027F2E8437A2BBFAF7FB45" box="[1295,1384,1159,1185]" name="Mexico" pageId="45" pageNumber="46">Mexico</collectingCountry>
. The analysis of the
<typeStatus id="549C883D77027F2E807BA297FE3AFB21" box="[323,421,1195,1221]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
of this latter species has made it possible to distinguish them in a simple way because
<taxonomicName id="4C274D1C77027F2E84BCA297FEA4FB0D" authorityName="Sharp" authorityYear="1885" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="trochanterinus">
<emphasis id="B953EA8D77027F2E84BCA297FEA4FB0D" italics="true" pageId="45" pageNumber="46">B. trochanterinus</emphasis>
</taxonomicName>
has abdominal styli that are very wide apically, while they are slender and elongate in
<taxonomicName id="4C274D1C77027F2E85C7A2F3FA0AFB0D" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[1279,1429,1231,1257]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tenuistylus">
<emphasis id="B953EA8D77027F2E85C7A2F3FA0AFB0D" box="[1279,1429,1231,1257]" italics="true" pageId="45" pageNumber="46">B. tenuistylus</emphasis>
</taxonomicName>
, in addition to the differences in the shape of the genital sternite and the aedeagus of both species. Furthermore,
<taxonomicName id="4C274D1C77027F2E84BCA2CFFE90FAD5" authorityName="Márquez &amp; Asiain" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tenuistylus">
<emphasis id="B953EA8D77027F2E84BCA2CFFE90FAD5" italics="true" pageId="45" pageNumber="46">B. tenuistylus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C274D1C77027F2E80B7A32BFDD0FAD5" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[399,591,1303,1329]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="metafemoralis">
<emphasis id="B953EA8D77027F2E80B7A32BFDD0FAD5" box="[399,591,1303,1329]" italics="true" pageId="45" pageNumber="46">B. metafemoralis</emphasis>
</taxonomicName>
and in the taxonomic comments of this latter species it has been indicated how to distinguish them.
</paragraph>
</subSubSection>
<subSubSection id="C33D651477027F2E81FFA363FC70FA79" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8B98369F77027F2E81FFA363FC70FA79" blockId="45.[151,1437,151,1833]" pageId="45" pageNumber="46">
<emphasis id="B953EA8D77027F2E81FFA363FED7FA9D" bold="true" box="[199,328,1375,1401]" pageId="45" pageNumber="46">Etymology</emphasis>
. The name of this species is a combination of the words “tenuis” and “stylus”, in order to highlight the slender styli of this species that easily separate it from
<taxonomicName id="4C274D1C77027F2E821DA3BFFC77FA79" authorityName="Sharp" authorityYear="1885" box="[805,1000,1411,1437]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="trochanterinus">
<emphasis id="B953EA8D77027F2E821DA3BFFC77FA79" box="[805,1000,1411,1437]" italics="true" pageId="45" pageNumber="46">B. trochanterinus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D651477027F2E81FFA39BFAEBF8CD" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8B98369F77027F2E81FFA39BFAEBF8CD" blockId="45.[151,1437,151,1833]" pageId="45" pageNumber="46">
<emphasis id="B953EA8D77027F2E81FFA39BFDE1FA24" bold="true" box="[199,638,1446,1473]" pageId="45" pageNumber="46">
Type material (
<specimenCount id="9D21FD1677027F2E8043A39AFE7BFA25" box="[379,484,1446,1473]" pageId="45" pageNumber="46" type="male">18 males</specimenCount>
,
<specimenCount id="9D21FD1677027F2E80D7A39AFDEEFA24" box="[495,625,1446,1472]" pageId="45" pageNumber="46" type="female">17 females</specimenCount>
):
</emphasis>
<materialsCitation id="3B4F3CC277027F2E83BBA39BFC52FA25" box="[643,973,1447,1473]" collectionCode="CC-UAEH" country="Mexico" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="holotype">
<emphasis id="B953EA8D77027F2E83BBA39BFD6FFA25" bold="true" box="[643,752,1447,1473]" pageId="45" pageNumber="46">
<typeStatus id="549C883D77027F2E83BBA39BFD6FFA25" box="[643,752,1447,1473]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
</emphasis>
(male,
<collectionCode id="ED36AE5A77027F2E827BA39BFC59FA25" box="[835,966,1447,1473]" pageId="45" pageNumber="46">CC-UAEH</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="3B4F3CC277027F2E82EEA39BFD11F9ED" collectingDate="1995-08-01" collectingDateMax="1995-08-30" collectingDateMin="1995-08-01" collectorName="J. Marquez" country="Mexico" county="Tlayacapan" elevation="1874" latitude="18.968056" location="Laureles" longLatPrecision="21" longitude="-99.00139" municipality="San Jose" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Mexico, Morelos" typeStatus="holotype">
<collectingRegion id="49E3F87D77027F2E82D8A39BFB3AFA25" box="[992,1189,1447,1473]" isEnumeration="true" pageId="45" pageNumber="46">México, Morelos</collectingRegion>
,
<collectingCounty id="62F94E1377027F2E8596A39BFAADFA25" box="[1198,1330,1447,1473]" pageId="45" pageNumber="46">Tlayacapan</collectingCounty>
,
<collectingMunicipality id="6BFCACE577027F2E8404A39BFA03FA25" box="[1340,1436,1447,1473]" pageId="45" pageNumber="46">San José</collectingMunicipality>
de los
<location id="8EF8604477027F2E81DAA3F7FEDAFA01" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E81DAA3F7FEDAFA01" box="[226,325,1483,1509]" country="Mexico" county="Tlayacapan" latitude="18.968056" longLatPrecision="21" longitude="-99.00139" municipality="San Jose" name="Laureles" pageId="45" pageNumber="46" stateProvince="Mexico, Morelos">Laureles</location>
, camino a
<location id="8EF8604477027F2E8085A3F7FDBFFA01" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E8085A3F7FDBFFA01" box="[445,544,1483,1509]" country="Mexico" county="Tlayacapan" latitude="18.968056" longLatPrecision="21" longitude="-99.00139" municipality="San Jose" name="Amatlan" pageId="45" pageNumber="46" stateProvince="Mexico, Morelos">Amatlán</location>
,
<geoCoordinate id="EE13505877027F2E8314A3F6FD35FA01" box="[556,682,1482,1509]" degrees="18" direction="north" minutes="58" orientation="latitude" pageId="45" pageNumber="46" precision="15" seconds="5" value="18.968056">N18°58´5”</geoCoordinate>
,
<geoCoordinate id="EE13505877027F2E838EA3F6FCA6FA01" box="[694,825,1482,1509]" degrees="99" direction="west" minutes="00" orientation="longitude" pageId="45" pageNumber="46" precision="15" seconds="5" value="-99.00139">W99°00´5”</geoCoordinate>
, localidad 1, bosque de pino-encino,
<quantity id="4CDF9B7A77027F2E85E6A3F7FAA9FA01" box="[1246,1334,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.874" pageId="45" pageNumber="46" unit="m" value="1874.0">
<elevation id="000AD1AC77027F2E85E6A3F7FAA9FA01" box="[1246,1334,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.874" pageId="45" pageNumber="46" unit="m" value="1874.0">1874 m</elevation>
</quantity>
, NTP-80 (calamar),
<date id="FF99105F77027F2E8037A3D3FE51F9ED" box="[271,462,1518,1545]" pageId="45" pageNumber="46" value="1995-08-01" valueMax="1995-08-30" valueMin="1995-08-01">
<collectingDate id="EFDDE9B777027F2E8037A3D3FE51F9ED" box="[271,462,1518,1545]" pageId="45" pageNumber="46" value="1995-08-01" valueMax="1995-08-30" valueMin="1995-08-01">1 a 30-VIII-1995</collectingDate>
</date>
,
<collectorName id="26D2534977027F2E80E0A3D3FDCCF9ED" box="[472,595,1519,1545]" pageId="45" pageNumber="46">J. Márquez</collectorName>
col.”
</materialsCitation>
<materialsCitation id="3B4F3CC277027F2E83ACA3D3FACCF9C9" collectingDate="1995-10" collectionCode="CC-UAEH" collectorName="K. Villavicencio &amp; J. Marquez" country="Mexico" county="Tlayacapan" location="Laureles" municipality="San Jose" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<emphasis id="B953EA8D77027F2E83ACA3D3FC93F9ED" bold="true" box="[660,780,1519,1545]" pageId="45" pageNumber="46">
<typeStatus id="549C883D77027F2E83ACA3D3FC93F9ED" box="[660,780,1519,1545]" pageId="45" pageNumber="46" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="B953EA8D77027F2E8221A3D2FC75F9ED" bold="true" box="[793,1002,1518,1545]" pageId="45" pageNumber="46">
<collectingCountry id="F330760F77027F2E8221A3D2FCEBF9EC" box="[793,884,1518,1544]" name="Mexico" pageId="45" pageNumber="46">Mexico</collectingCountry>
:
<collectingRegion id="49E3F87D77027F2E8247A3D3FC7AF9ED" box="[895,997,1519,1545]" country="Mexico" name="Morelos" pageId="45" pageNumber="46">Morelos</collectingRegion>
:
</emphasis>
<collectingCounty id="62F94E1377027F2E82C1A3D3FB1FF9ED" box="[1017,1152,1519,1545]" pageId="45" pageNumber="46">Tlayacapan</collectingCounty>
,
<collectingMunicipality id="6BFCACE577027F2E85B1A3D3FB76F9ED" box="[1161,1257,1519,1545]" pageId="45" pageNumber="46">San José</collectingMunicipality>
de los
<location id="8EF8604477027F2E840DA3D3FA07F9ED" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E840DA3D3FA07F9ED" box="[1333,1432,1519,1545]" country="Mexico" county="Tlayacapan" municipality="San Jose" name="Laureles" pageId="45" pageNumber="46" stateProvince="Morelos">Laureles</location>
, NTP-80, localidad 1, bosque de pino-encino,
<date id="FF99105F77027F2E83A3A02EFD6CF9C9" box="[667,755,1554,1581]" pageId="45" pageNumber="46" value="1995-10">
<collectingDate id="EFDDE9B777027F2E83A3A02EFD6CF9C9" box="[667,755,1554,1581]" pageId="45" pageNumber="46" value="1995-10">X-1995</collectingDate>
</date>
,
<collectorName id="26D2534977027F2E83C7A02FFC28F9C9" box="[767,951,1555,1581]" pageId="45" pageNumber="46">K. Villavicencio</collectorName>
y
<collectorName id="26D2534977027F2E82EDA02FFBCCF9C9" box="[981,1107,1555,1581]" pageId="45" pageNumber="46">J. Márquez</collectorName>
cols.” (6,
<collectionCode id="ED36AE5A77027F2E85F1A02FFAD4F9C9" box="[1225,1355,1555,1581]" pageId="45" pageNumber="46">CC-UAEH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E8466A02FFDFBF9B5" collectingDate="1995-10" collectionCode="MAAS" country="Mexico" location="Same" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477027F2E8466A02FFA03F9C9" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E8466A02FFA03F9C9" box="[1374,1436,1555,1581]" country="Mexico" name="Same" pageId="45" pageNumber="46" stateProvince="Morelos">Same</location>
data, except: “
<date id="FF99105F77027F2E800AA00AFE4CF9B5" box="[306,467,1590,1617]" pageId="45" pageNumber="46" value="1995-10">
<collectingDate id="EFDDE9B777027F2E800AA00AFE4CF9B5" box="[306,467,1590,1617]" pageId="45" pageNumber="46" value="1995-10">Octubre-1995</collectingDate>
</date>
” (4,
<collectionCode id="ED36AE5A77027F2E833EA00BFDC3F9B5" box="[518,604,1591,1617]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E8355A00BFE83F991" collectionCode="MAAS" country="Mexico" location="Same" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477027F2E8355A00BFD34F9B5" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E8355A00BFD34F9B5" box="[621,683,1591,1617]" country="Mexico" name="Same" pageId="45" pageNumber="46" stateProvince="Morelos">Same</location>
data, except: “localidad 3, bosque mesófilo de montaña perturbado” (3,
<collectionCode id="ED36AE5A77027F2E8186A067FE8BF991" box="[190,276,1627,1653]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E8010A067FE84F97D" collectingDate="1996-06" collectionCode="MAAS" collectorName="J. Marquez &amp; K. Villavicencio" country="Mexico" isTailEnumeration="true" location="Tlayacapan" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477027F2E800AA067FE26F991" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E800AA067FE26F991" box="[306,441,1627,1653]" country="Mexico" name="Tlayacapan" pageId="45" pageNumber="46" stateProvince="Morelos">Tlayacapan</location>
, selva baja caducifolia, zona 4, NTP-80,
<date id="FF99105F77027F2E82B7A067FC6FF991" box="[911,1008,1627,1653]" pageId="45" pageNumber="46" value="1996-06">
<collectingDate id="EFDDE9B777027F2E82B7A067FC6FF991" box="[911,1008,1627,1653]" pageId="45" pageNumber="46" value="1996-06">VI-1996</collectingDate>
</date>
,
<collectorName id="26D2534977027F2E82C5A067FBE3F991" box="[1021,1148,1627,1653]" pageId="45" pageNumber="46">J. Márquez</collectorName>
y
<collectorName id="26D2534977027F2E85A3A067FACCF991" box="[1179,1363,1627,1653]" pageId="45" pageNumber="46">K. Villavicencio</collectorName>
cols.” (1,
<collectionCode id="ED36AE5A77027F2E8185A043FE8CF97D" box="[189,275,1663,1689]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E801EA043FBA7F959" collectingDate="1995-07" collectionCode="MAAS" collectorName="J. Marquez" country="Mexico" county="Tlayacapan" elevation="1783" latitude="18.96861" location="Laureles" longLatPrecision="21" longitude="-99.020836" municipality="San Jose" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<collectingCounty id="62F94E1377027F2E8008A043FE28F97D" box="[304,439,1663,1689]" pageId="45" pageNumber="46">Tlayacapan</collectingCounty>
,
<collectingMunicipality id="6BFCACE577027F2E80FAA043FDBBF97D" box="[450,548,1663,1689]" pageId="45" pageNumber="46">San José</collectingMunicipality>
de los
<location id="8EF8604477027F2E834EA043FD46F97D" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E834EA043FD46F97D" box="[630,729,1663,1689]" country="Mexico" county="Tlayacapan" latitude="18.96861" longLatPrecision="21" longitude="-99.020836" municipality="San Jose" name="Laureles" pageId="45" pageNumber="46" stateProvince="Morelos">Laureles</location>
, camino a
<location id="8EF8604477027F2E826AA043FC2AF97D" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E826AA043FC2AF97D" box="[850,949,1663,1689]" country="Mexico" county="Tlayacapan" latitude="18.96861" longLatPrecision="21" longitude="-99.020836" municipality="San Jose" name="Amatlan" pageId="45" pageNumber="46" stateProvince="Morelos">Amatlan</location>
,
<geoCoordinate id="EE13505877027F2E82F9A042FBA0F97D" box="[961,1087,1662,1689]" degrees="18" direction="north" minutes="58" orientation="latitude" pageId="45" pageNumber="46" precision="15" seconds="7" value="18.96861">N18°587”</geoCoordinate>
,
<geoCoordinate id="EE13505877027F2E8573A042FB43F97D" box="[1099,1244,1662,1689]" degrees="99" direction="west" minutes="01" orientation="longitude" pageId="45" pageNumber="46" precision="15" seconds="15" value="-99.020836">W99°0115”</geoCoordinate>
,
<quantity id="4CDF9B7A77027F2E85D1A043FADFF97D" box="[1257,1344,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.783" pageId="45" pageNumber="46" unit="m" value="1783.0">
<elevation id="000AD1AC77027F2E85D1A043FADFF97D" box="[1257,1344,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.783" pageId="45" pageNumber="46" unit="m" value="1783.0">1783 m</elevation>
</quantity>
, bosque mesófilo de montaña, NTP-80 (calamar),
<date id="FF99105F77027F2E8355A09EFD77F959" box="[621,744,1698,1725]" pageId="45" pageNumber="46" value="1995-07">
<collectingDate id="EFDDE9B777027F2E8355A09EFD77F959" box="[621,744,1698,1725]" pageId="45" pageNumber="46" value="1995-07">Julio-1995</collectingDate>
</date>
,
<collectorName id="26D2534977027F2E83CBA09FFCEEF959" box="[755,881,1699,1725]" pageId="45" pageNumber="46">J. Márquez</collectorName>
col.” (2,
<collectionCode id="ED36AE5A77027F2E82E3A09FFBAEF959" box="[987,1073,1699,1725]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E857BA09FFD07F905" collectingDate="1995-08" collectionCode="MAAS" country="Mexico" location="Same" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477027F2E857BA09FFB1EF959" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E857BA09FFB1EF959" box="[1091,1153,1699,1725]" country="Mexico" name="Same" pageId="45" pageNumber="46" stateProvince="Morelos">Same</location>
data, except: “bosque de pino y pino-encino,
<date id="FF99105F77027F2E804FA0FAFD9AF905" box="[375,517,1734,1761]" pageId="45" pageNumber="46" value="1995-08">
<collectingDate id="EFDDE9B777027F2E804FA0FAFD9AF905" box="[375,517,1734,1761]" pageId="45" pageNumber="46" value="1995-08">agosto-1995</collectingDate>
</date>
” (9,
<collectionCode id="ED36AE5A77027F2E8302A0FBFD0FF905" box="[570,656,1735,1761]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E8399A0FBFE85F8E1" collectingDate="1995-09" collectionCode="MAAS" country="Mexico" location="Same" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477027F2E8399A0FBFD40F905" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E8399A0FBFD40F905" box="[673,735,1735,1761]" country="Mexico" name="Same" pageId="45" pageNumber="46" stateProvince="Morelos">Same</location>
data, except: “bosque mesófilo de montaña,
<date id="FF99105F77027F2E85ECA0FAFA0CF905" box="[1236,1427,1734,1761]" pageId="45" pageNumber="46" value="1995-09">
<collectingDate id="EFDDE9B777027F2E85ECA0FAFA0CF905" box="[1236,1427,1734,1761]" pageId="45" pageNumber="46" value="1995-09">septiembre-1995</collectingDate>
</date>
” (3,
<collectionCode id="ED36AE5A77027F2E8184A0D7FE8DF8E1" box="[188,274,1771,1797]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E801CA0D7FBA2F8E1" box="[292,1085,1770,1797]" collectingDate="1995-10" collectionCode="MAAS" country="Mexico" location="Same" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477027F2E801CA0D7FEFDF8E1" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E801CA0D7FEFDF8E1" box="[292,354,1771,1797]" country="Mexico" name="Same" pageId="45" pageNumber="46" stateProvince="Morelos">Same</location>
data, except: “bosque de pino-encino,
<date id="FF99105F77027F2E822AA0D6FC34F8E1" box="[786,939,1770,1797]" pageId="45" pageNumber="46" value="1995-10">
<collectingDate id="EFDDE9B777027F2E822AA0D6FC34F8E1" box="[786,939,1770,1797]" pageId="45" pageNumber="46" value="1995-10">octubre-1995</collectingDate>
</date>
” (3,
<collectionCode id="ED36AE5A77027F2E82D8A0D7FBA9F8E1" box="[992,1078,1771,1797]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E857FA0D7FD80F8CD" collectingDate="1995-12" collectionCode="MAAS" country="Mexico" location="Same" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477027F2E857FA0D7FB1AF8E1" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E857FA0D7FB1AF8E1" box="[1095,1157,1771,1797]" country="Mexico" name="Same" pageId="45" pageNumber="46" stateProvince="Morelos">Same</location>
data, except: “bosque de pino,
<date id="FF99105F77027F2E81EFA132FE14F8CD" box="[215,395,1806,1833]" pageId="45" pageNumber="46" value="1995-12">
<collectingDate id="EFDDE9B777027F2E81EFA132FE14F8CD" box="[215,395,1806,1833]" pageId="45" pageNumber="46" value="1995-12">diciembre-1995</collectingDate>
</date>
” (1,
<collectionCode id="ED36AE5A77027F2E80F9A133FD88F8CD" box="[449,535,1807,1833]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277027F2E8311A133FAEFF8CD" box="[553,1392,1807,1833]" collectingDate="1996-06" collectionCode="MAAS" country="Mexico" location="Same" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477027F2E8311A133FDF8F8CD" LSID="urn:lsid:plazi:treatment:038E878977037F2E81AFA3A7FAEBF8CD:8EF8604477027F2E8311A133FDF8F8CD" box="[553,615,1807,1833]" country="Mexico" name="Same" pageId="45" pageNumber="46" stateProvince="Morelos">Same</location>
data, except: “bosque mesófilo de montaña,
<date id="FF99105F77027F2E8567A133FB43F8CD" box="[1119,1244,1807,1833]" pageId="45" pageNumber="46" value="1996-06">
<collectingDate id="EFDDE9B777027F2E8567A133FB43F8CD" box="[1119,1244,1807,1833]" pageId="45" pageNumber="46" value="1996-06">junio-1996</collectingDate>
</date>
” (2,
<collectionCode id="ED36AE5A77027F2E842AA133FAF7F8CD" box="[1298,1384,1807,1833]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="45" pageNumber="46" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>