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<document id="3F6D7923B2BEB0A97D406E173A48AC10" ID-DOI="10.11646/zootaxa.5152.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6630873" ID-ZooBank="92E9DD85-6CC6-4602-BD7C-C51F49CEEF47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654852147308" checkinUser="plazi" docAuthor="Márquez, Juan &amp; Asiain, Julieta" docDate="2022" docId="038E878977027F2C81AFA153FB4AFD61" docLanguage="en" docName="zootaxa.5152.1.1.pdf" docOrigin="Zootaxa 5152 (1)" docSource="http://zoobank.org/92e9dd85-6cc6-4602-bd7c-c51f49ceef47" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Belonuchus tepoztecus Márquez &amp; Asiain 2022, sp. nov." docType="treatment" docUuid="4790A935-2AD7-4EF0-BD6F-BAD5E420780D" docUuidSource="ZooBank" docVersion="3" lastPageNumber="48" masterDocId="FFB7FFF1772F7F038138A63CFF9FFFE4" masterDocTitle="Taxonomy of the Mexican species of Belonuchus Nordmann (Coleoptera: Staphylinidae)" masterLastPageNumber="129" masterPageNumber="1" pageNumber="46" updateTime="1699335993931" updateUser="plazi">
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<mods:title id="FA66BAEE58A9084E88529296A6A74DD3">Taxonomy of the Mexican species of Belonuchus Nordmann (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="A2DB42C15492E74BC8C7395E3F2769DF">Márquez, Juan</mods:namePart>
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<mods:namePart id="632998BD8AFDE90ED50F491D139338BC">Asiain, Julieta</mods:namePart>
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<treatment id="038E878977027F2C81AFA153FB4AFD61" ID-DOI="http://doi.org/10.5281/zenodo.6630859" ID-Zenodo-Dep="6630859" LSID="urn:lsid:zoobank.org:act:4790A935-2AD7-4EF0-BD6F-BAD5E420780D" httpUri="http://treatment.plazi.org/id/038E878977027F2C81AFA153FB4AFD61" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46">
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<paragraph id="8B98369F77027F2E81AFA153FD8FF86E" blockId="45.[151,1059,1903,2003]" box="[151,528,1903,1930]" pageId="45" pageNumber="46">
<heading id="D0D081F377027F2E81AFA153FD8FF86E" bold="true" box="[151,528,1903,1930]" fontSize="11" level="2" pageId="45" pageNumber="46" reason="4">
<emphasis id="B953EA8D77027F2E81AFA153FD8FF86E" bold="true" box="[151,528,1903,1930]" pageId="45" pageNumber="46">
<taxonomicName id="4C274D1C77027F2E81AFA153FE33F86E" ID-CoL="9GKKD" authority="Márquez &amp; Asiain, 2022" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[151,428,1903,1930]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tepoztecus" status="sp. nov.">
<emphasis id="B953EA8D77027F2E81AFA153FE33F86E" bold="true" box="[151,428,1903,1930]" italics="true" pageId="45" pageNumber="46">Belonuchus tepoztecus</emphasis>
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<taxonomicNameLabel id="A26057F677027F2E808BA14CFD8FF86E" box="[435,528,1904,1930]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B98369F77027F2E81AFA1A8FBBCF84B" blockId="45.[151,1059,1903,2003]" box="[151,1059,1940,1967]" pageId="45" pageNumber="46">http://zoobank.org/NomenclaturalActs/4790a935-2ad7-4ef0-bd6f-bad5e420780d</paragraph>
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<subSubSection id="C33D651477027F2D81AFA184FA1AF871" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="8B98369F77027F2E81AFA184FDEAF836" blockId="45.[151,1059,1903,2003]" box="[151,629,1976,2003]" pageId="45" pageNumber="46">
<figureCitation id="131C2A1A77027F2E81AFA184FF6BF836" box="[151,244,1976,2002]" captionStart="FIGURE 4" captionStartId="98.[151,250,1720,1744]" captionTargetBox="[170,1410,196,1684]" captionTargetId="figure-25@98.[161,1426,181,1696]" captionTargetPageId="98" captionText="FIGURE 4. Dorsal view of males of: a) Belonuchus tenuistylus (holotype), b) B. tepoztecus (paratype), c) B. trochanterinus, d) B. zunilensis (lectotype), e) B. bidens (syntype, NHM©), f) B. bidens, g) B. erichsoni, h) B. jalappensis. Scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630889" httpUri="https://zenodo.org/record/6630889/files/figure.png" pageId="45" pageNumber="46">Figs. 4b</figureCitation>
,
<figureCitation id="131C2A1A77027F2E81C7A184FEB4F836" box="[255,299,1976,2002]" captionStart="FIGURE 11" captionStartId="105.[151,250,1833,1857]" captionTargetBox="[175,1420,203,1797]" captionTargetId="figure-19@105.[151,1436,181,1809]" captionTargetPageId="105" captionText="FIGURE 11. Mesoventrite of males of: a) Belonuchus cariniventris, b) B. oxyporinus, c) B. rufisternus, d) B. tepoztecus, e) B. bidens, f) B. jalappensis, g) B. longitarsus, h) B. ephippiatus, i) B. dichrous, j) B. flavipennis, k) B. platypterus, l) B. juanmorronei, m) B. iridescens. Red arrow: intercoxal process of the mesoventrite, blue arrow: transverse discal ridge, green arrow: longitudinal carina in anterior half." figureDoi="http://doi.org/10.5281/zenodo.6630913" httpUri="https://zenodo.org/record/6630913/files/figure.png" pageId="45" pageNumber="46">11d</figureCitation>
,
<figureCitation id="131C2A1A77027F2E800EA184FEFCF836" box="[310,355,1976,2002]" captionStart="FIGURE 12" captionStartId="106.[151,250,1870,1894]" captionTargetBox="[191,1398,191,1835]" captionTargetId="figure-25@106.[165,1422,181,1846]" captionTargetPageId="106" captionText="FIGURE 12. Mesoventrite of males of: a) Belonuchus viridipennis, b) Belonuchus sp. Q, c) B. cyanipennis. Ventral view of the left femur of male anterior leg of: d) B. apiciventris, e) B. oxyporimimus, f) B. oxyporinus, g) B. tepoztecus, h) B. bidens, i) B. viridipennis. Ventral view of the left posterior leg of males of: j) B. julietitae, k) B. linuzensis (small specimen), l) B. linuzensis (large specimen), m) B. metafemoralis, n) B. oxyporimimus, o) B. oxyporinus. Red arrow: intercoxal process of the mesoventrite, blue arrow: transverse discal ridge." figureDoi="http://doi.org/10.5281/zenodo.6630915" httpUri="https://zenodo.org/record/6630915/files/figure.png" pageId="45" pageNumber="46">
<quantity id="4CDF9B7A77027F2E800EA184FEFCF836" box="[310,355,1976,2002]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="45" pageNumber="46" unit="g" value="12.0">12g</quantity>
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,
<figureCitation id="131C2A1A77027F2E8056A184FE04F836" box="[366,411,1976,2002]" captionStart="FIGURE 14" captionStartId="108.[151,250,1944,1968]" captionTargetBox="[185,1402,182,1894]" captionTargetId="figure-25@108.[167,1419,181,1920]" captionTargetPageId="108" captionText="FIGURE 14. Dorsal (left) and ventral (right) view of the first four visible abdominal segments of males of: a) Belonuchus singularis, b) B. tepoztecus. Abdominal styli of male of: c) B. alternans, d) B. confusus, e) B. singularis, f) B. magnistylus (right: detail of left stylus in ventral view), g) B. trochanterinus (right: detail of left stylus in ventral view)." figureDoi="http://doi.org/10.5281/zenodo.6630919" httpUri="https://zenodo.org/record/6630919/files/figure.png" pageId="45" pageNumber="46">14b</figureCitation>
,
<figureCitation id="131C2A1A77027F2E809EA184FE4EF837" box="[422,465,1976,2003]" captionStart="FIGURE 15" captionStartId="109.[151,250,1821,1845]" captionTargetBox="[175,1417,182,1784]" captionTargetId="figure-19@109.[160,1427,181,1797]" captionTargetPageId="109" captionText="FIGURE 15. Male pregenital sternite of: a) Belonuchus alternans, b) B. oxyporinus, c) B. tepoztecus, d) B. jalappensis, e) B. iridescens, f) B. platypterus, g) B. cyanipennis, h) B. apiciventris, i) B. linuzensis, j) B. metafemoralis, k) B. rufisternus, l) B. tenuistylus, m) B. bidens, n) B. pollens, o) B. longitarsus." figureDoi="http://doi.org/10.5281/zenodo.6630921" httpUri="https://zenodo.org/record/6630921/files/figure.png" pageId="45" pageNumber="46">15c</figureCitation>
,
<figureCitation id="131C2A1A77027F2E80E5A184FD97F836" box="[477,520,1976,2002]" captionStart="FIGURE 19" captionStartId="113.[151,250,1823,1847]" captionTargetBox="[165,1426,181,1793]" captionTargetId="figure-19@113.[161,1426,181,1800]" captionTargetPageId="113" captionText="FIGURE 19. Male genital sternite of: a) Belonuchus tepoztecus, b) B. trochanterinus, c) B. zunilensis, d) B. bidens, e) B. erichsoni, f) B. jalappensis, g) B. mixtecus, h) B. pollens, i) B. rufipennis, j) B. basiventris, k) B. connexus, l) B. longitarsus, m) B. rufiventris, n) B. xanthomelas, o) B. ephippiatus, p) B. moroni. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630927" httpUri="https://zenodo.org/record/6630927/files/figure.png" pageId="45" pageNumber="46">19a</figureCitation>
,
<figureCitation id="131C2A1A77027F2E832BA184FDDFF836" box="[531,576,1976,2002]" captionStart="FIGURE 22" captionStartId="116.[151,250,1894,1918]" captionTargetBox="[162,1427,182,1859]" captionTargetId="figure-25@116.[151,1436,181,1871]" captionTargetPageId="116" captionText="FIGURE 22. Ventral (left) and lateral (right) view of the aedeagus of: a) Belonuchus oxyporimimus, b) B. oxyporinus, c) B. pulcher, d) B. rufisternus, e) B. singularis, f) B. tenuistylus, g) B. tepoztecus (at center detail of the paramere), h) B. trochanterinus, i) B. zunilensis. Scale bar = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6630937" httpUri="https://zenodo.org/record/6630937/files/figure.png" pageId="45" pageNumber="46">
<quantity id="4CDF9B7A77027F2E832BA184FDDFF836" box="[531,576,1976,2002]" metricMagnitude="-2" metricUnit="kg" metricValue="2.2" pageId="45" pageNumber="46" unit="g" value="22.0">22g</quantity>
</figureCitation>
,
<figureCitation id="131C2A1A77027F2E8373A184FDEAF836" box="[587,629,1976,2002]" captionStart="FIGURE 29" captionStartId="123.[151,250,1780,1804]" captionTargetBox="[164,1422,195,1747]" captionTargetId="figure-19@123.[151,1436,181,1756]" captionTargetPageId="123" captionText="FIGURE 29. Geographical distribution in Mexico of: a) B. linuzensis and Belonuchus sp. Q; b) B. cyanipennis, B. flavipennis and B. tepoztecus." figureDoi="http://doi.org/10.5281/zenodo.6630951" httpUri="https://zenodo.org/record/6630951/files/figure.png" pageId="45" pageNumber="46">29b</figureCitation>
</paragraph>
<paragraph id="8B98369F77017F2D81AFA6ABFC1FFEA5" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
Total body length
<quantity id="4CDF9B7A77017F2D8053A6ABFE4BFF55" box="[363,468,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="46" pageNumber="47" unit="mm" value="11.6">11.6 mm</quantity>
. Black on head, antennae, mandibles (except apex and internal margins), pronotum, scutellum, posterior half of prosternum, sixth visible abdominal segment (except anterior border) and abdominal styli. Reddish on elytra, legs, sterna (except posterior half of prosternum and anterior half of mesoventrite), visible abdominal segments 15 and anterior border of sixth. Brown at apex and internal margins of mandibles, maxillary and labial palpi, anterior half of mesoventrite and genital segment.
</paragraph>
<paragraph id="8B98369F77017F2D81FFA777FAB2FDD9" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B953EA8D77017F2D81FFA777FE98FE81" bold="true" box="[199,263,331,357]" pageId="46" pageNumber="47">Head</emphasis>
: transverse, ratio length/width 0.69. Dorsal surface with sulcate longitudinal midline visible from anterior margin to almost posterior margin of head, but less notable in posterior half; front clearly foveate between antennal insertions. Eyes slightly longer than 0.5 times the cephalic lateral length, not protruding laterally.Antennomeres 45 elongate, 67 as long as wide, 810 transverse. Mandibles 1.58 times longer than head; each with two well separated teeth (basal and middle); mandibular channel well developed, with external and internal margins moderately separated at base, internal margin carinate, extending forward to level of middle tooth. Apical palpomere of maxillary and labial palpi nearly 1.25 times longer than preapical palpomere. Head 1.3 times wider than pronotum.
</paragraph>
<paragraph id="8B98369F77017F2D81FFA47BFBD7FC2D" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B953EA8D77017F2D81FFA47BFEBFFD85" bold="true" box="[199,288,583,609]" pageId="46" pageNumber="47">Thorax</emphasis>
: each dorsal row of pronotum with five punctures; pronotum almost as long as wide (ratio 1.06) and almost as wide at anterior corners as at posterior corners (ratio 1.06). Scutellum with punctures denser than on elytra, latter with sparse punctation, with fine, pale setae. Prosternum with an elevated area next to anterior margin and a transverse line near the place where black color initiates. Intercoxal process of mesoventrite shield-shaped; transverse discal ridge well developed, broadly U-shaped, reaching close to margin of intercoxal process, but not touching it (
<figureCitation id="131C2A1A77017F2D8026A4C7FEE2FCF2" box="[286,381,763,790]" captionStart="FIGURE 11" captionStartId="105.[151,250,1833,1857]" captionTargetBox="[175,1420,203,1797]" captionTargetId="figure-19@105.[151,1436,181,1809]" captionTargetPageId="105" captionText="FIGURE 11. Mesoventrite of males of: a) Belonuchus cariniventris, b) B. oxyporinus, c) B. rufisternus, d) B. tepoztecus, e) B. bidens, f) B. jalappensis, g) B. longitarsus, h) B. ephippiatus, i) B. dichrous, j) B. flavipennis, k) B. platypterus, l) B. juanmorronei, m) B. iridescens. Red arrow: intercoxal process of the mesoventrite, blue arrow: transverse discal ridge, green arrow: longitudinal carina in anterior half." figureDoi="http://doi.org/10.5281/zenodo.6630913" httpUri="https://zenodo.org/record/6630913/files/figure.png" pageId="46" pageNumber="47">Fig. 11d</figureCitation>
). Profemur with external row of spines initiating near basal third and reaching the apex or very close to it, the most apical spines are placed at same level as spines of internal apex that are less numerous and shorter; external spines increasing in size from base to apex, where being conspicuous (
<figureCitation id="131C2A1A77017F2D8541A57FFB43FCB9" box="[1145,1244,835,861]" captionStart="FIGURE 12" captionStartId="106.[151,250,1870,1894]" captionTargetBox="[191,1398,191,1835]" captionTargetId="figure-25@106.[165,1422,181,1846]" captionTargetPageId="106" captionText="FIGURE 12. Mesoventrite of males of: a) Belonuchus viridipennis, b) Belonuchus sp. Q, c) B. cyanipennis. Ventral view of the left femur of male anterior leg of: d) B. apiciventris, e) B. oxyporimimus, f) B. oxyporinus, g) B. tepoztecus, h) B. bidens, i) B. viridipennis. Ventral view of the left posterior leg of males of: j) B. julietitae, k) B. linuzensis (small specimen), l) B. linuzensis (large specimen), m) B. metafemoralis, n) B. oxyporimimus, o) B. oxyporinus. Red arrow: intercoxal process of the mesoventrite, blue arrow: transverse discal ridge." figureDoi="http://doi.org/10.5281/zenodo.6630915" httpUri="https://zenodo.org/record/6630915/files/figure.png" pageId="46" pageNumber="47">
Fig.
<quantity id="4CDF9B7A77017F2D8597A57FFB43FCB9" box="[1199,1244,835,861]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="46" pageNumber="47" unit="g" value="12.0">12g</quantity>
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). Metatrochanter modified like a hook that is as long as or longer than 1/2 length of metafemur, hook elongate, slender, with apex strongly curved and acute; metafemur with internal margin curved at level of overlap with hook; metatibia clearly curved at internal margin near basal third. Tarsomeres moderately flattened dorsally.
</paragraph>
<paragraph id="8B98369F77017F2D81FFA5EEFEB8FAE9" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B953EA8D77017F2D81FFA5EEFEA3FC08" bold="true" box="[199,316,978,1004]" pageId="46" pageNumber="47">Abdomen</emphasis>
: first three visible tergites with posterior basal transverse carina well developed, adjacent zone slightly depressed and narrow (
<figureCitation id="131C2A1A77017F2D80CDA5CBFDC8FBF6" box="[501,599,1015,1042]" captionStart="FIGURE 14" captionStartId="108.[151,250,1944,1968]" captionTargetBox="[185,1402,182,1894]" captionTargetId="figure-25@108.[167,1419,181,1920]" captionTargetPageId="108" captionText="FIGURE 14. Dorsal (left) and ventral (right) view of the first four visible abdominal segments of males of: a) Belonuchus singularis, b) B. tepoztecus. Abdominal styli of male of: c) B. alternans, d) B. confusus, e) B. singularis, f) B. magnistylus (right: detail of left stylus in ventral view), g) B. trochanterinus (right: detail of left stylus in ventral view)." figureDoi="http://doi.org/10.5281/zenodo.6630919" httpUri="https://zenodo.org/record/6630919/files/figure.png" pageId="46" pageNumber="47">Fig. 14b</figureCitation>
); tergite one with sparse, wide-superficial punctures in depression, becoming denser toward remaining surface; depression on visible tergites 23 with wide-superficial punctures denser than on remaining surface; all tergites also with fine punctures that are denser on last three visible tergites. All sternites (
<figureCitation id="131C2A1A77017F2D8198A25FFE9FFB9A" box="[160,256,1123,1150]" captionStart="FIGURE 14" captionStartId="108.[151,250,1944,1968]" captionTargetBox="[185,1402,182,1894]" captionTargetId="figure-25@108.[167,1419,181,1920]" captionTargetPageId="108" captionText="FIGURE 14. Dorsal (left) and ventral (right) view of the first four visible abdominal segments of males of: a) Belonuchus singularis, b) B. tepoztecus. Abdominal styli of male of: c) B. alternans, d) B. confusus, e) B. singularis, f) B. magnistylus (right: detail of left stylus in ventral view), g) B. trochanterinus (right: detail of left stylus in ventral view)." figureDoi="http://doi.org/10.5281/zenodo.6630919" httpUri="https://zenodo.org/record/6630919/files/figure.png" pageId="46" pageNumber="47">Fig. 14b</figureCitation>
) with wide-superficial punctures denser than on tergites, but on sternites 56 less dense than on remaining sternites, also all sternites with fine punctures as on tergites. Male pregenital sternite without emargination at posterior margin (
<figureCitation id="131C2A1A77017F2D8067A297FE20FB21" box="[351,447,1194,1221]" captionStart="FIGURE 15" captionStartId="109.[151,250,1821,1845]" captionTargetBox="[175,1417,182,1784]" captionTargetId="figure-19@109.[160,1427,181,1797]" captionTargetPageId="109" captionText="FIGURE 15. Male pregenital sternite of: a) Belonuchus alternans, b) B. oxyporinus, c) B. tepoztecus, d) B. jalappensis, e) B. iridescens, f) B. platypterus, g) B. cyanipennis, h) B. apiciventris, i) B. linuzensis, j) B. metafemoralis, k) B. rufisternus, l) B. tenuistylus, m) B. bidens, n) B. pollens, o) B. longitarsus." figureDoi="http://doi.org/10.5281/zenodo.6630921" httpUri="https://zenodo.org/record/6630921/files/figure.png" pageId="46" pageNumber="47">Fig. 15c</figureCitation>
). Genital sternite short (2.31 times longer than wide), moderately asymmetrical, anterior portion occupying 40% and posterior portion 60% of its length; apical sinuation deep (
<figureCitation id="131C2A1A77017F2D8549A2F3FB4DFB0D" box="[1137,1234,1231,1257]" captionStart="FIGURE 19" captionStartId="113.[151,250,1823,1847]" captionTargetBox="[165,1426,181,1793]" captionTargetId="figure-19@113.[161,1426,181,1800]" captionTargetPageId="113" captionText="FIGURE 19. Male genital sternite of: a) Belonuchus tepoztecus, b) B. trochanterinus, c) B. zunilensis, d) B. bidens, e) B. erichsoni, f) B. jalappensis, g) B. mixtecus, h) B. pollens, i) B. rufipennis, j) B. basiventris, k) B. connexus, l) B. longitarsus, m) B. rufiventris, n) B. xanthomelas, o) B. ephippiatus, p) B. moroni. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630927" httpUri="https://zenodo.org/record/6630927/files/figure.png" pageId="46" pageNumber="47">Fig. 19a</figureCitation>
). Abdominal styli very slender.
</paragraph>
<paragraph id="8B98369F77017F2D81FFA32AFBCCFA9D" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B953EA8D77017F2D81FFA32AFEA5FAD4" bold="true" box="[199,314,1302,1328]" pageId="46" pageNumber="47">Aedeagus</emphasis>
: length
<quantity id="4CDF9B7A77017F2D80A0A32BFE6CFAD5" box="[408,499,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="46" pageNumber="47" unit="mm" value="1.8">1.8 mm</quantity>
; oval-elongate shape, apex wide and slightly convex; sides with slight constriction before apex, widened toward base; basal half moderately wider than apical half, most notable in lateral view; paramere with irregular anterior margin (or a little jagged); internal sac visible (
<figureCitation id="131C2A1A77017F2D82DCA363FBD9FA9D" box="[996,1094,1375,1401]" captionStart="FIGURE 22" captionStartId="116.[151,250,1894,1918]" captionTargetBox="[162,1427,182,1859]" captionTargetId="figure-25@116.[151,1436,181,1871]" captionTargetPageId="116" captionText="FIGURE 22. Ventral (left) and lateral (right) view of the aedeagus of: a) Belonuchus oxyporimimus, b) B. oxyporinus, c) B. pulcher, d) B. rufisternus, e) B. singularis, f) B. tenuistylus, g) B. tepoztecus (at center detail of the paramere), h) B. trochanterinus, i) B. zunilensis. Scale bar = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6630937" httpUri="https://zenodo.org/record/6630937/files/figure.png" pageId="46" pageNumber="47">
Fig.
<quantity id="4CDF9B7A77017F2D8520A363FBD9FA9D" box="[1048,1094,1375,1401]" metricMagnitude="-2" metricUnit="kg" metricValue="2.2" pageId="46" pageNumber="47" unit="g" value="22.0">22g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="8B98369F77017F2D81FFA3BEFABAF97D" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B953EA8D77017F2D81FFA3BEFED8FA78" bold="true" box="[199,327,1410,1436]" pageId="46" pageNumber="47">Variability</emphasis>
. Total body length in males
<quantity id="4CDF9B7A77017F2D83ADA3BFFD63FA79" box="[661,764,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="46" pageNumber="47" unit="mm" value="10.8">10.8 mm</quantity>
(range 9.912.0 mm), in females
<quantity id="4CDF9B7A77017F2D85BFA3BFFB7FFA79" box="[1159,1248,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="46" pageNumber="47" unit="mm" value="9.8">9.8 mm</quantity>
(range
<quantity id="4CDF9B7A77017F2D8402A3BFFF58FA25" metricMagnitude="-3" metricUnit="m" metricValue="9.7" metricValueMax="10.6" metricValueMin="8.8" pageId="46" pageNumber="47" unit="mm" value="9.7" valueMax="10.6" valueMin="8.8">8.810.6 mm</quantity>
); ratio length/width of head in males 0.69 (range 0.670.74), in females 0.84 (range 0.820.87); ratio mandibular length/cephalic length in males 1.56 (range 1.331.70), in females 0.90 (range 0.820.96); ratio cephalic width/ pronotal width in males 1.30 (range 1.211.36), in females 1.08 (range 1.001.15); ratio length/width of pronotum in males 1.06 (range 1.021.10), in females 1.14 (range 1.101.18); ratio anterior width/posterior width of pronotum in males 1.1 (range 1.071.18), in females 1.03 (range 1.001.04). Sulcate longitudinal midline of head may be less visible than in the
<typeStatus id="549C883D77017F2D805EA067FE53F991" box="[358,460,1627,1653]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
. In some specimens the entire prosternum is black. Posterior margin of male pregenital sternite can be weakly emarginate or without emargination, with the latter state being the most frequent.
</paragraph>
<paragraph id="8B98369F77017F2D81FFA09EFA1AF871" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B953EA8D77017F2D81FFA09EFE54F959" bold="true" box="[199,459,1698,1725]" pageId="46" pageNumber="47">Taxonomic comments</emphasis>
. This species is very similar to
<taxonomicName id="4C274D1C77017F2D8217A09FFC4DF958" authorityName="Sharp" authorityYear="1885" box="[815,978,1699,1724]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="apiciventris">
<emphasis id="B953EA8D77017F2D8217A09FFC4DF958" box="[815,978,1699,1724]" italics="true" pageId="46" pageNumber="47">B. apiciventris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C274D1C77017F2D8531A09FFB3DF959" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[1033,1186,1699,1725]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="rufisternus">
<emphasis id="B953EA8D77017F2D8531A09FFB3DF959" box="[1033,1186,1699,1725]" italics="true" pageId="46" pageNumber="47">B. rufisternus</emphasis>
</taxonomicName>
, both in the color pattern as well as in the total length and other characters. In the description of
<taxonomicName id="4C274D1C77017F2D82E0A0FBFBE4F904" authorityName="Sharp" authorityYear="1885" box="[984,1147,1735,1760]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="apiciventris">
<emphasis id="B953EA8D77017F2D82E0A0FBFBE4F904" box="[984,1147,1735,1760]" italics="true" pageId="46" pageNumber="47">B. apiciventris</emphasis>
</taxonomicName>
the main differences with this species are pointed out.
<taxonomicName id="4C274D1C77017F2D80EFA0D7FD49F8E0" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[471,726,1771,1797]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="tepoztecus">
<emphasis id="B953EA8D77017F2D80EFA0D7FD49F8E0" box="[471,726,1771,1797]" italics="true" pageId="46" pageNumber="47">Belonuchus tepoztecus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C274D1C77017F2D82E9A0D7FBF5F8E1" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[977,1130,1771,1797]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="rufisternus">
<emphasis id="B953EA8D77017F2D82E9A0D7FBF5F8E1" box="[977,1130,1771,1797]" italics="true" pageId="46" pageNumber="47">B. rufisternus</emphasis>
</taxonomicName>
by having the anterior half of the mesoventrite black (or at least dark) and the posterior half reddish, five punctures in each dorsal row of pronotum, male pregenital sternite with emarginate posterior margin and different male genital sternite and aedeagus.
<taxonomicName id="4C274D1C77017F2D81AFA16BFE03F895" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[151,412,1879,1905]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="rufisternus">
<emphasis id="B953EA8D77017F2D81AFA16BFE03F895" box="[151,412,1879,1905]" italics="true" pageId="46" pageNumber="47">Belonuchus rufisternus</emphasis>
</taxonomicName>
has the entire mesoventrite reddish, six punctures in each dorsal row of pronotum, male pregenital sternite moderately emarginate in posterior margin, and a different male genital sternite and aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C33D651477017F2D81FFA1A3FCB4F839" pageId="46" pageNumber="47" type="etymology">
<paragraph id="8B98369F77017F2D81FFA1A3FCB4F839" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
<emphasis id="B953EA8D77017F2D81FFA1A3FED7F85D" bold="true" box="[199,328,1951,1977]" pageId="46" pageNumber="47">Etymology</emphasis>
. The name of this species is derived from the geographical place of origin of the specimens, which is known as “Sierra del Tepozteco” in the state of
<collectingRegion id="49E3F87D77017F2D83FEA1FFFCB8F839" box="[710,807,1987,2013]" country="Mexico" name="Morelos" pageId="46" pageNumber="47">Morelos</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D651477007F2C81FFA6ABFB4AFD61" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8B98369F77007F2C81FFA6ABFB4AFD61" blockId="47.[151,1437,150,645]" pageId="47" pageNumber="48">
<emphasis id="B953EA8D77007F2C81FFA6ABFD9BFF54" bold="true" box="[199,516,150,177]" pageId="47" pageNumber="48">
Type material (
<specimenCount id="9D21FD1677007F2C80BDA6AAFE68FF54" box="[389,503,150,176]" pageId="47" pageNumber="48" type="male">63 males</specimenCount>
):
</emphasis>
<materialsCitation id="3B4F3CC277007F2C8337A6ABFCFDFF55" box="[527,866,151,177]" collectionCode="CC-UAEH" pageId="47" pageNumber="48" specimenCount="1" typeStatus="holotype">
<emphasis id="B953EA8D77007F2C8337A6ABFDE3FF55" bold="true" box="[527,636,151,177]" pageId="47" pageNumber="48">
<typeStatus id="549C883D77007F2C8337A6ABFDE3FF55" box="[527,636,151,177]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
(male,
<collectionCode id="ED36AE5A77007F2C83E0A6ABFCC4FF55" box="[728,859,151,177]" pageId="47" pageNumber="48">CC-UAEH</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="3B4F3CC277007F2C8248A6ABFD97FF1D" collectingDate="1995-07-01" collectingDateMax="1995-07-30" collectingDateMin="1995-07-01" collectorName="J. Marquez" county="Tlayacapan" elevation="1783" latitude="18.978333" location="Laureles" longLatPrecision="880" longitude="-99.01667" municipality="San Jose" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
<collectingRegion id="49E3F87D77007F2C8242A6ABFC49FF55" box="[890,982,151,177]" country="Mexico" name="Mexico" pageId="47" pageNumber="48">México</collectingRegion>
:
<collectingRegion id="49E3F87D77007F2C82DDA6ABFBDAFF55" box="[997,1093,151,177]" country="Mexico" name="Morelos" pageId="47" pageNumber="48">Morelos</collectingRegion>
,
<collectingCounty id="62F94E1377007F2C856BA6ABFB48FF55" box="[1107,1239,151,177]" pageId="47" pageNumber="48">Tlayacapan</collectingCounty>
,
<collectingMunicipality id="6BFCACE577007F2C85DEA6ABFAD4FF55" box="[1254,1355,151,177]" pageId="47" pageNumber="48">San José</collectingMunicipality>
de los
<location id="8EF8604477007F2C81AFA687FF65FF31" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C81AFA687FF65FF31" box="[151,250,187,213]" county="Tlayacapan" latitude="18.978333" longLatPrecision="880" longitude="-99.01667" municipality="San Jose" name="Laureles" pageId="47" pageNumber="48" stateProvince="Mexico">Laureles</location>
, camino a
<location id="8EF8604477007F2C8042A687FE42FF31" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8042A687FE42FF31" box="[378,477,187,213]" county="Tlayacapan" latitude="18.978333" longLatPrecision="880" longitude="-99.01667" municipality="San Jose" name="Amatlan" pageId="47" pageNumber="48" stateProvince="Mexico">Amatlán</location>
, bosque mesófilo de montaña,
<quantity id="4CDF9B7A77007F2C827FA687FC3EFF31" box="[839,929,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.783" pageId="47" pageNumber="48" unit="m" value="1783.0">
<elevation id="000AD1AC77007F2C827FA687FC3EFF31" box="[839,929,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.783" pageId="47" pageNumber="48" unit="m" value="1783.0">1783 m</elevation>
</quantity>
,
<geoCoordinate id="EE13505877007F2C8297A686FBB7FF31" box="[943,1064,186,213]" degrees="18" direction="north" minutes="58.7" orientation="latitude" pageId="47" pageNumber="48" precision="92" value="18.978333">N18°58.7</geoCoordinate>
,
<geoCoordinate id="EE13505877007F2C850EA687FB00FF31" box="[1078,1183,187,213]" degrees="99" direction="west" minutes="01" orientation="longitude" pageId="47" pageNumber="48" precision="925" value="-99.01667">W99°01</geoCoordinate>
, NTP-80 (calamar),
<date id="FF99105F77007F2C84B6A687FEA3FF1D" pageId="47" pageNumber="48" value="1995-07-01" valueMax="1995-07-30" valueMin="1995-07-01">
<collectingDate id="EFDDE9B777007F2C84B6A687FEA3FF1D" pageId="47" pageNumber="48" value="1995-07-01" valueMax="1995-07-30" valueMin="1995-07-01">1 a 30-VII-1995</collectingDate>
</date>
,
<collectorName id="26D2534977007F2C8071A6E3FE57FF1D" box="[329,456,223,249]" pageId="47" pageNumber="48">J. Márquez</collectorName>
col.”
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C832DA6E3FACCFF1D" box="[533,1363,223,250]" collectingDate="1995-07-01" collectingDateMax="1995-07-30" collectingDateMin="1995-07-01" collectionCode="CC-UAEH" collectorName="J. Marquez" county="Tlayacapan" elevation="1783" latitude="18.978333" location="Laureles" longLatPrecision="880" longitude="-99.01667" municipality="San Jose" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<emphasis id="B953EA8D77007F2C832DA6E3FD12FF1D" bold="true" box="[533,653,223,249]" pageId="47" pageNumber="48">
<typeStatus id="549C883D77007F2C832DA6E3FD12FF1D" box="[533,653,223,249]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
</emphasis>
(all specimen males): same data as holotype (16,
<collectionCode id="ED36AE5A77007F2C85F1A6E3FAD4FF1D" box="[1225,1355,223,249]" pageId="47" pageNumber="48">CC-UAEH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C8466A6E3FD12FEF9" collectingDate="1996-08-01" collectingDateMax="1996-08-30" collectingDateMin="1996-08-01" collectionCode="MAAS" location="Same" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<location id="8EF8604477007F2C8466A6E3FA03FF1D" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8466A6E3FA03FF1D" box="[1374,1436,223,249]" name="Same" pageId="47" pageNumber="48" stateProvince="Mexico">Same</location>
data, except: “
<date id="FF99105F77007F2C8001A73FFE65FEF9" box="[313,506,259,285]" pageId="47" pageNumber="48" value="1996-08-01" valueMax="1996-08-30" valueMin="1996-08-01">
<collectingDate id="EFDDE9B777007F2C8001A73FFE65FEF9" box="[313,506,259,285]" pageId="47" pageNumber="48" value="1996-08-01" valueMax="1996-08-30" valueMin="1996-08-01">1 a 30-VIII-1996</collectingDate>
</date>
” (6,
<collectionCode id="ED36AE5A77007F2C8317A73FFD1AFEF9" box="[559,645,259,285]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C83AFA73FFB5FFEF9" box="[663,1216,258,285]" collectingDate="1995-09-01" collectingDateMax="1995-09-30" collectingDateMin="1995-09-01" collectionCode="MAAS" location="Same" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
<location id="8EF8604477007F2C83AFA73FFD4AFEF9" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C83AFA73FFD4AFEF9" box="[663,725,259,285]" name="Same" pageId="47" pageNumber="48" stateProvince="Mexico">Same</location>
data, except: “
<date id="FF99105F77007F2C8247A73FFBB3FEF9" box="[895,1068,258,285]" pageId="47" pageNumber="48" value="1995-09-01" valueMax="1995-09-30" valueMin="1995-09-01">
<collectingDate id="EFDDE9B777007F2C8247A73FFBB3FEF9" box="[895,1068,258,285]" pageId="47" pageNumber="48" value="1995-09-01" valueMax="1995-09-30" valueMin="1995-09-01">1 a 30-IX-1995</collectingDate>
</date>
” (5,
<collectionCode id="ED36AE5A77007F2C855AA73FFB27FEF9" box="[1122,1208,259,285]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C85F2A73EFCDFFE81" collectingDate="1995-07-01" collectingDateMax="1995-07-30" collectingDateMin="1995-07-01" collectionCode="MAAS" collectorName="J. Marquez" country="Mexico" elevation="1550" latitude="18.953611" location="Santa Catarina" longLatPrecision="21" longitude="-99.004166" municipality="Tlayacapan" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<emphasis id="B953EA8D77007F2C85F2A73EFA03FEF9" bold="true" box="[1226,1436,258,285]" pageId="47" pageNumber="48">
<collectingCountry id="F330760F77007F2C85F2A73EFABAFEF8" box="[1226,1317,258,284]" name="Mexico" pageId="47" pageNumber="48">Mexico</collectingCountry>
:
<collectingRegion id="49E3F87D77007F2C8409A73FFA08FEF9" box="[1329,1431,259,285]" country="Mexico" name="Morelos" pageId="47" pageNumber="48">Morelos</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BFCACE577007F2C8199A71BFEB7FEA5" box="[161,296,295,321]" pageId="47" pageNumber="48">Tlayacapan</collectingMunicipality>
, camino a
<location id="8EF8604477007F2C80A7A71BFDD9FEA5" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C80A7A71BFDD9FEA5" box="[415,582,295,321]" country="Mexico" latitude="18.953611" longLatPrecision="21" longitude="-99.004166" municipality="Tlayacapan" name="Santa Catarina" pageId="47" pageNumber="48" stateProvince="Morelos">Santa Catarina</location>
,
<geoCoordinate id="EE13505877007F2C8369A71AFD42FEA5" box="[593,733,294,321]" degrees="18" direction="north" minutes="57" orientation="latitude" pageId="47" pageNumber="48" precision="15" seconds="13" value="18.953611">N18°57´13”</geoCoordinate>
,
<geoCoordinate id="EE13505877007F2C83D0A71AFCE6FEA5" box="[744,889,294,321]" degrees="99" direction="west" minutes="00" orientation="longitude" pageId="47" pageNumber="48" precision="15" seconds="15" value="-99.004166">W99°00´15”</geoCoordinate>
,
<quantity id="4CDF9B7A77007F2C82BDA71AFC43FEA5" box="[901,988,294,321]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="47" pageNumber="48" unit="m" value="1550.0">
<elevation id="000AD1AC77007F2C82BDA71AFC43FEA5" box="[901,988,294,321]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="47" pageNumber="48" unit="m" value="1550.0">1550 m</elevation>
</quantity>
, Loc-4, selva tropical caducifolia,
<date id="FF99105F77007F2C8458A71BFE9DFE81" pageId="47" pageNumber="48" value="1995-07-01" valueMax="1995-07-30" valueMin="1995-07-01">
<collectingDate id="EFDDE9B777007F2C8458A71BFE9DFE81" pageId="47" pageNumber="48" value="1995-07-01" valueMax="1995-07-30" valueMin="1995-07-01">1-30- VII-1995</collectingDate>
</date>
, K. Villavicencio y
<collectorName id="26D2534977007F2C80D1A777FDF7FE81" box="[489,616,331,357]" pageId="47" pageNumber="48">J. Márquez</collectorName>
cols.” (3,
<collectionCode id="ED36AE5A77007F2C83DBA777FCA6FE81" box="[739,825,331,357]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C8275A777FA06FE81" box="[845,1433,330,357]" collectingDate="1995-08-01" collectingDateMax="1995-08-30" collectingDateMin="1995-08-01" collectionCode="MAAS" country="Mexico" location="Same" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477007F2C8275A777FC14FE81" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8275A777FC14FE81" box="[845,907,331,357]" country="Mexico" name="Same" pageId="47" pageNumber="48" stateProvince="Morelos">Same</location>
data, except: “
<date id="FF99105F77007F2C8504A777FA9FFE81" box="[1084,1280,330,357]" pageId="47" pageNumber="48" value="1995-08-01" valueMax="1995-08-30" valueMin="1995-08-01">
<collectingDate id="EFDDE9B777007F2C8504A777FA9FFE81" box="[1084,1280,330,357]" pageId="47" pageNumber="48" value="1995-08-01" valueMax="1995-08-30" valueMin="1995-08-01">1 a 30-VIII-1995</collectingDate>
</date>
” (1,
<collectionCode id="ED36AE5A77007F2C8403A777FA0EFE81" box="[1339,1425,331,357]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C81AFA753FD4AFE6D" box="[151,725,366,393]" collectingDate="1995-09-01" collectingDateMax="1995-09-30" collectingDateMin="1995-09-01" collectionCode="MAAS" country="Mexico" location="Same" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477007F2C81AFA753FF4AFE6D" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C81AFA753FF4AFE6D" box="[151,213,367,393]" country="Mexico" name="Same" pageId="47" pageNumber="48" stateProvince="Morelos">Same</location>
data, except: “
<date id="FF99105F77007F2C80BAA753FDAEFE6D" box="[386,561,366,393]" pageId="47" pageNumber="48" value="1995-09-01" valueMax="1995-09-30" valueMin="1995-09-01">
<collectingDate id="EFDDE9B777007F2C80BAA753FDAEFE6D" box="[386,561,366,393]" pageId="47" pageNumber="48" value="1995-09-01" valueMax="1995-09-30" valueMin="1995-09-01">1 a 30-IX-1995</collectingDate>
</date>
” (10,
<collectionCode id="ED36AE5A77007F2C834FA753FD52FE6D" box="[631,717,367,393]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C83D8A753FA8AFE6D" box="[736,1301,366,393]" collectingDate="1995-10-01" collectingDateMax="1995-10-30" collectingDateMin="1995-10-01" collectionCode="MAAS" country="Mexico" location="Same" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477007F2C83D8A753FC81FE6D" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C83D8A753FC81FE6D" box="[736,798,367,393]" country="Mexico" name="Same" pageId="47" pageNumber="48" stateProvince="Morelos">Same</location>
data, except: “
<date id="FF99105F77007F2C82F3A753FBEEFE6D" box="[971,1137,366,393]" pageId="47" pageNumber="48" value="1995-10-01" valueMax="1995-10-30" valueMin="1995-10-01">
<collectingDate id="EFDDE9B777007F2C82F3A753FBEEFE6D" box="[971,1137,366,393]" pageId="47" pageNumber="48" value="1995-10-01" valueMax="1995-10-30" valueMin="1995-10-01">1 a 30-X-1995</collectingDate>
</date>
” (10,
<collectionCode id="ED36AE5A77007F2C858FA753FA92FE6D" box="[1207,1293,367,393]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C8418A753FDD4FE49" collectingDate="1995-11-01" collectingDateMax="1995-11-30" collectingDateMin="1995-11-01" collectionCode="MAAS" country="Mexico" location="Same" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477007F2C8418A753FAC1FE6D" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8418A753FAC1FE6D" box="[1312,1374,367,393]" country="Mexico" name="Same" pageId="47" pageNumber="48" stateProvince="Morelos">Same</location>
data, except: “
<date id="FF99105F77007F2C8038A7AFFE2DFE49" box="[256,434,402,429]" pageId="47" pageNumber="48" value="1995-11-01" valueMax="1995-11-30" valueMin="1995-11-01">
<collectingDate id="EFDDE9B777007F2C8038A7AFFE2DFE49" box="[256,434,402,429]" pageId="47" pageNumber="48" value="1995-11-01" valueMax="1995-11-30" valueMin="1995-11-01">1 a 30-XI-1995</collectingDate>
</date>
” (2,
<collectionCode id="ED36AE5A77007F2C80D5A7AFFDDCFE49" box="[493,579,403,429]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C8360A7AFFB41FE49" box="[600,1246,402,429]" collectingDate="1995-12-01" collectingDateMax="1995-12-30" collectingDateMin="1995-12-01" collectionCode="CC-UAEH" country="Mexico" location="Same" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477007F2C8360A7AFFD09FE49" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8360A7AFFD09FE49" box="[600,662,403,429]" country="Mexico" name="Same" pageId="47" pageNumber="48" stateProvince="Morelos">Same</location>
data, except: “
<date id="FF99105F77007F2C827FA7AFFB9CFE49" box="[839,1027,402,429]" pageId="47" pageNumber="48" value="1995-12-01" valueMax="1995-12-30" valueMin="1995-12-01">
<collectingDate id="EFDDE9B777007F2C827FA7AFFB9CFE49" box="[839,1027,402,429]" pageId="47" pageNumber="48" value="1995-12-01" valueMax="1995-12-30" valueMin="1995-12-01">1 a 30-XII-1995</collectingDate>
</date>
” (2,
<collectionCode id="ED36AE5A77007F2C8506A7AFFB5FFE49" box="[1086,1216,403,429]" pageId="47" pageNumber="48">CC-UAEH</collectionCode>
). “
</materialsCitation>
<materialsCitation id="3B4F3CC277007F2C85E6A7AFFB16FE11" collectingDate="1995-09-01" collectingDateMax="1995-11-30" collectingDateMin="1995-09-01" collectionCode="MAAS" collectorName="J. Marquez" country="Mexico" county="Tlayacapan" elevation="2000" latitude="18.975834" location="Laureles" longLatPrecision="21" longitude="-99.013054" municipality="San Jose" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<collectingCounty id="62F94E1377007F2C85E6A7AFFAFAFE49" box="[1246,1381,403,429]" pageId="47" pageNumber="48">Tlayacapan</collectingCounty>
,
<collectingMunicipality id="6BFCACE577007F2C844AA7AFFF58FE35" pageId="47" pageNumber="48">San José</collectingMunicipality>
de los
<location id="8EF8604477007F2C8024A78BFEE0FE35" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8024A78BFEE0FE35" box="[284,383,439,465]" country="Mexico" county="Tlayacapan" latitude="18.975834" longLatPrecision="21" longitude="-99.013054" municipality="San Jose" name="Laureles" pageId="47" pageNumber="48" stateProvince="Morelos">Laureles</location>
, Loc-1,
<geoCoordinate id="EE13505877007F2C80E7A78AFDF5FE35" box="[479,618,438,465]" degrees="18" direction="north" minutes="58" orientation="latitude" pageId="47" pageNumber="48" precision="15" seconds="33" value="18.975834">N18°5833”</geoCoordinate>
,
<geoCoordinate id="EE13505877007F2C834FA78BFC97FE35" box="[631,776,439,465]" degrees="99" direction="west" minutes="00" orientation="longitude" pageId="47" pageNumber="48" precision="15" seconds="47" value="-99.013054">W99°0047”</geoCoordinate>
,
<quantity id="4CDF9B7A77007F2C822EA78BFCF0FE35" box="[790,879,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="47" pageNumber="48" unit="m" value="2000.0">
<elevation id="000AD1AC77007F2C822EA78BFCF0FE35" box="[790,879,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="47" pageNumber="48" unit="m" value="2000.0">2000 m</elevation>
</quantity>
, bosque de pino-encino, NTP-80 (calamar,
<date id="FF99105F77007F2C8458A78BFF66FE11" pageId="47" pageNumber="48" value="1995-09-01" valueMax="1995-09-30" valueMin="1995-09-01">
<collectingDate id="EFDDE9B777007F2C8458A78BFF66FE11" pageId="47" pageNumber="48" value="1995-09-01" valueMax="1995-09-30" valueMin="1995-09-01">1-30- IX-1995</collectingDate>
</date>
,
<collectorName id="26D2534977007F2C803CA7E7FE1FFE11" box="[260,384,475,501]" pageId="47" pageNumber="48">J. Márquez</collectorName>
col.” (1,
<collectionCode id="ED36AE5A77007F2C80DFA7E7FDA2FE11" box="[487,573,475,501]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
). Same data, except: “
<date id="FF99105F77007F2C820EA7E7FC7DFE11" box="[822,994,474,501]" pageId="47" pageNumber="48" value="1995-11-01" valueMax="1995-11-30" valueMin="1995-11-01">
<collectingDate id="EFDDE9B777007F2C820EA7E7FC7DFE11" box="[822,994,474,501]" pageId="47" pageNumber="48" value="1995-11-01" valueMax="1995-11-30" valueMin="1995-11-01">1 a 30-XI-1995</collectingDate>
</date>
” (1,
<collectionCode id="ED36AE5A77007F2C8520A7E7FBF1FE11" box="[1048,1134,475,501]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
). “
</materialsCitation>
<materialsCitation id="3B4F3CC277007F2C85B1A7E7FE45FDD9" collectingDate="1989-10-28" collectingDateMax="1989-11-19" collectingDateMin="1989-10-28" collectionCode="MAAS" collectorName="G. Quiroz &amp; J. L. Navarrete" country="Mexico" county="Tlayacapan" elevation="1803" location="Laureles" municipality="San Jose" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<collectingCounty id="62F94E1377007F2C85B1A7E7FA8FFE11" box="[1161,1296,475,501]" pageId="47" pageNumber="48">Tlayacapan</collectingCounty>
,
<collectingMunicipality id="6BFCACE577007F2C8422A7E7FAE4FE11" box="[1306,1403,475,501]" pageId="47" pageNumber="48">San José</collectingMunicipality>
de los
<location id="8EF8604477007F2C8187A7C3FEBDFDFD" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8187A7C3FEBDFDFD" box="[191,290,511,537]" country="Mexico" county="Tlayacapan" municipality="San Jose" name="Laureles" pageId="47" pageNumber="48" stateProvince="Morelos">Laureles</location>
, bosque mesófilo de montaña,
<location id="8EF8604477007F2C8345A7C3FD2FFDFD" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8345A7C3FD2FFDFD" box="[637,688,511,537]" country="Mexico" county="Tlayacapan" municipality="San Jose" name="Loc." pageId="47" pageNumber="48" stateProvince="Morelos">Loc.</location>
3,
<quantity id="4CDF9B7A77007F2C83EBA7C3FCB5FDFD" box="[723,810,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.803" pageId="47" pageNumber="48" unit="m" value="1803.0">
<elevation id="000AD1AC77007F2C83EBA7C3FCB5FDFD" box="[723,810,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.803" pageId="47" pageNumber="48" unit="m" value="1803.0">1803 m</elevation>
</quantity>
, NTP-80 (pulpo),
<date id="FF99105F77007F2C82C0A7C3FB4CFDFD" box="[1016,1235,511,537]" pageId="47" pageNumber="48" value="1989-10-28" valueMax="1989-11-19" valueMin="1989-10-28">
<collectingDate id="EFDDE9B777007F2C82C0A7C3FB4CFDFD" box="[1016,1235,511,537]" pageId="47" pageNumber="48" value="1989-10-28" valueMax="1989-11-19" valueMin="1989-10-28">28-X a 19-XI-1989</collectingDate>
</date>
,
<collectorName id="26D2534977007F2C85E7A7C3FAD0FDFD" box="[1247,1359,511,537]" pageId="47" pageNumber="48">G. Quiroz</collectorName>
y
<collectorName id="26D2534977007F2C8453A7C3FE9AFDD9" pageId="47" pageNumber="48">J. L. Navarrete</collectorName>
cols.” (1,
<collectionCode id="ED36AE5A77007F2C8044A41FFE4DFDD9" box="[380,466,547,573]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C80DDA41FFC67FD85" collectingDate="1990-09-08" collectingDateMax="1991-08-24" collectingDateMin="1990-09-08" collectionCode="MAAS" country="Mexico" county="Same" elevation="1751" location="Loc." municipality="Loc." pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<collectingCounty id="62F94E1377007F2C80DDA41FFDBCFDD9" box="[485,547,547,573]" pageId="47" pageNumber="48">Same</collectingCounty>
data, except:
<collectingMunicipality id="6BFCACE577007F2C83FAA41FFD6AFDD9" box="[706,757,547,573]" pageId="47" pageNumber="48">Loc.</collectingMunicipality>
5,
<quantity id="4CDF9B7A77007F2C8221A41EFCEEFDD9" box="[793,881,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.751" pageId="47" pageNumber="48" unit="m" value="1751.0">
<elevation id="000AD1AC77007F2C8221A41EFCEEFDD9" box="[793,881,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.751" pageId="47" pageNumber="48" unit="m" value="1751.0">1751 m</elevation>
</quantity>
,
<date id="FF99105F77007F2C8246A41FFBE0FDD9" box="[894,1151,547,573]" pageId="47" pageNumber="48" value="1991-07-21" valueMax="1991-08-24" valueMin="1991-07-21">
<collectingDate id="EFDDE9B777007F2C8246A41FFBE0FDD9" box="[894,1151,547,573]" pageId="47" pageNumber="48" value="1991-07-21" valueMax="1991-08-24" valueMin="1991-07-21">21-VII a 24-VIII-1991</collectingDate>
</date>
” (1,
<collectionCode id="ED36AE5A77007F2C858FA41FFA92FDD9" box="[1207,1293,547,573]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
). Same data, except: “bosque de pino-encino,
<location id="8EF8604477007F2C833BA47BFDA9FD85" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C833BA47BFDA9FD85" box="[515,566,583,609]" country="Mexico" county="Same" municipality="Loc." name="Loc." pageId="47" pageNumber="48" stateProvince="Morelos">Loc.</location>
1,
<quantity id="4CDF9B7A77007F2C836FA47BFD32FD85" box="[599,685,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="47" pageNumber="48" unit="m" value="1830.0">
<elevation id="000AD1AC77007F2C836FA47BFD32FD85" box="[599,685,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="47" pageNumber="48" unit="m" value="1830.0">1830 m</elevation>
</quantity>
,
<date id="FF99105F77007F2C8380A47BFCF9FD85" box="[696,870,583,609]" pageId="47" pageNumber="48" value="1990-09-08" valueMax="1990-09-29" valueMin="1990-09-08">
<collectingDate id="EFDDE9B777007F2C8380A47BFCF9FD85" box="[696,870,583,609]" pageId="47" pageNumber="48" value="1990-09-08" valueMax="1990-09-29" valueMin="1990-09-08">8 a 29-IX-1990</collectingDate>
</date>
” (1,
<collectionCode id="ED36AE5A77007F2C82A2A47BFC6FFD85" box="[922,1008,583,609]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C8539A47BFE85FD61" collectingDate="1990-06-03" collectingDateMax="1990-06-30" collectingDateMin="1990-06-03" collectionCode="MAAS" country="Mexico" location="Same" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<location id="8EF8604477007F2C8539A47BFBA0FD85" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C8539A47BFBA0FD85" box="[1025,1087,583,609]" country="Mexico" name="Same" pageId="47" pageNumber="48" stateProvince="Morelos">Same</location>
data, except: “
<date id="FF99105F77007F2C85DCA47BFA0DFD85" box="[1252,1426,583,609]" pageId="47" pageNumber="48" value="1990-06-03" valueMax="1990-06-30" valueMin="1990-06-03">
<collectingDate id="EFDDE9B777007F2C85DCA47BFA0DFD85" box="[1252,1426,583,609]" pageId="47" pageNumber="48" value="1990-06-03" valueMax="1990-06-30" valueMin="1990-06-03">3 a 30-VI-1990</collectingDate>
</date>
” (1,
<collectionCode id="ED36AE5A77007F2C8185A457FE8CFD61" box="[189,275,619,645]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3CC277007F2C801DA457FB4EFD61" box="[293,1233,619,645]" collectingDate="1998-06-22" collectingDateMax="1990-06-30" collectingDateMin="1990-06-03" collectionCode="MAAS" collectorName="A. C. Deloya" country="Mexico" isTailEnumeration="true" location="Jumiltepec" municipality="Ocuituco" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Morelos" typeStatus="paratype">
<collectingMunicipality id="6BFCACE577007F2C8016A457FE05FD61" box="[302,410,619,645]" pageId="47" pageNumber="48">Ocuituco</collectingMunicipality>
,
<location id="8EF8604477007F2C809EA457FDBDFD61" LSID="urn:lsid:plazi:treatment:038E878977027F2C81AFA153FB4AFD61:8EF8604477007F2C809EA457FDBDFD61" box="[422,546,619,645]" municipality="Ocuituco" name="Jumiltepec" pageId="47" pageNumber="48">Jumiltepec</location>
, trampa de fruta,
<date id="FF99105F77007F2C83DEA457FCF2FD61" box="[742,877,619,645]" pageId="47" pageNumber="48" value="1998-06-22">
<collectingDate id="EFDDE9B777007F2C83DEA457FCF2FD61" box="[742,877,619,645]" pageId="47" pageNumber="48" value="1998-06-22">22-VI-1998</collectingDate>
</date>
,
<collectorName id="26D2534977007F2C824FA457FB94FD61" box="[887,1035,619,645]" pageId="47" pageNumber="48">A. C. Deloya</collectorName>
col.” (1,
<collectionCode id="ED36AE5A77007F2C854BA457FB56FD61" box="[1139,1225,619,645]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="47" pageNumber="48" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>