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<document id="2DFA52038BB6D951EEADC39C41A6D73F" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFAEFF90F38C97E4FCBBBA49" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus komareki Hájek &amp; Brancucci 2015, new species" docType="treatment" docUuid="E1F0B209-C450-48DE-9870-A348E8B51335" docUuidSource="ZooBank" docVersion="10" lastPageNumber="321" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="320" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B965FE72A16206D5946320E45D386508" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4F5D65C6558B7D15CAED4CA90ED6CCE7">
<mods:title id="EB7EA7BBE2857ADC0A4F0AA3B96D0A47">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="9D57B03D5CDC2C1038ABB28482209F51" type="personal">
<mods:role id="7F37D4F76DA948FE6FD781D03EDD3A1E">
<mods:roleTerm id="559DD7160CA52D45C04A6F11A103ED2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7361B95307AB22A278595B693D40B6D9">Hájek, Jiří</mods:namePart>
<mods:affiliation id="226BDBC212357BA529BB53C102464C9C">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 – Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="46E4A84F17ADB5D64167BFFC745D2E13" type="email">jiri_hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="F13BDC0953A59DDFA838F4010FEEBB2C" type="personal">
<mods:role id="196F227E43385BB38CF19383898967F4">
<mods:roleTerm id="CC6949D9DB405FD52A859EC43E162FD1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA5ECF968F1B662E17D0CBD80A1A1937">Brancucci, Michel</mods:namePart>
<mods:affiliation id="B388E6D99C212612700C0515EB4DF220">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="06AF62DABE14D26DEDBB82B343EC6CD3">text</mods:typeOfResource>
<mods:relatedItem id="634B20E3E91F402FE3DEFACC537A6BB9" type="host">
<mods:titleInfo id="F731A6D5E63B647DDB97D0467CE59A62">
<mods:title id="3E25AC40B6FBDF2A7E9FFE4B3E3D68E4">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="7FC22D66762E718C6132BB6552771E3C">
<mods:date id="C67C78CA02C994B5304D745851DDFD65">2015</mods:date>
<mods:detail id="00C361242306DD8EC363CC4D4A33683E" type="pubDate">
<mods:number id="5B885ED1F5B0807792F0C50133B28280">2015-08-06</mods:number>
</mods:detail>
<mods:detail id="CBA3F517780D11561F850EEDDC531650" type="volume">
<mods:number id="29DB20E09ED89041E72FC6A2DF1A6337">63</mods:number>
</mods:detail>
<mods:extent id="3B97EFBF0137EE2AA16291D881510F93" unit="page">
<mods:start id="807C9F535CAC78399BC423EDDBEBABB4">309</mods:start>
<mods:end id="A36ECE75496303C13C7AAEBC8B7B154F">326</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FBBE5AB94E5C47F4E0DBAA1A3D7090EF">journal article</mods:classification>
<mods:identifier id="A18F7E1F255B8DC089AEA970D5929E8C" type="CLB-Dataset">55750</mods:identifier>
<mods:identifier id="36A3A21A669BC18A084DAC1992E46580" type="DOI">10.5281/zenodo.5385198</mods:identifier>
<mods:identifier id="AB1B2635049EFDA0AA54729D8F681365" type="GBIF-Dataset">c7f95b94-53f8-420d-97c5-4e27d7c211bf</mods:identifier>
<mods:identifier id="F3F40A01F8B0D79BB3C4E7E33273B0B0" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="DF2E031A40E7161151AA9AF1B183BD09" type="Zenodo-Dep">5385198</mods:identifier>
<mods:identifier id="EFF70963D8BAF74811C49ADB87C45CB0" type="ZooBank">AC7F1491-CD89-4F10-9D61-55354B55E368</mods:identifier>
</mods:mods>
<treatment id="038A87C4FFAEFF90F38C97E4FCBBBA49" ID-DOI="http://doi.org/10.5281/zenodo.5466893" ID-GBIF-Taxon="194955225" ID-Zenodo-Dep="5466893" LSID="urn:lsid:zoobank.org:act:E1F0B209-C450-48DE-9870-A348E8B51335" httpUri="http://treatment.plazi.org/id/038A87C4FFAEFF90F38C97E4FCBBBA49" lastPageId="12" lastPageNumber="321" pageId="11" pageNumber="320">
<subSubSection id="C3396559FFAEFF97F38C97E4FAE7BEE9" box="[934,1330,1437,1461]" pageId="11" pageNumber="320" type="nomenclature">
<paragraph id="8B9C36D2FFAEFF97F38C97E4FAE7BEE9" blockId="11.[934,1330,1437,1493]" box="[934,1330,1437,1461]" pageId="11" pageNumber="320">
<heading id="D0D481BEFFAEFF97F38C97E4FAE7BEE9" bold="true" box="[934,1330,1437,1461]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="320" reason="2">
<emphasis id="B957EAC0FFAEFF97F38C97E4FAE7BEE9" bold="true" box="[934,1330,1437,1461]" pageId="11" pageNumber="320">
<taxonomicName id="4C234D51FFAEFF97F38C97E4FB4BBEE9" ID-CoL="8LXXH" authority="Hájek &amp; Brancucci, 2015" authorityName="Hájek &amp; Brancucci" authorityYear="2015" box="[934,1182,1437,1461]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="komareki" status="sp. nov.">
<emphasis id="B957EAC0FFAEFF97F38C97E4FB4BBEE9" bold="true" box="[934,1182,1437,1461]" italics="true" pageId="11" pageNumber="320">Laccophilus komareki</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26457BBFFAEFF97F48797E4FAE7BEE9" box="[1197,1330,1437,1461]" pageId="11" pageNumber="320" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAEFF97F40397C4FB7ABE89" box="[1065,1199,1469,1493]" pageId="11" pageNumber="320" type="description">
<paragraph id="8B9C36D2FFAEFF97F40397C4FB7ABE89" blockId="11.[934,1330,1437,1493]" box="[1065,1199,1469,1493]" pageId="11" pageNumber="320">
(
<figureCitation id="13182A57FFAEFF97F41B97C4FBABBE89" box="[1073,1150,1469,1493]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 1–9. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="11" pageNumber="320">Figs. 9</figureCitation>
,
<figureCitation id="13182A57FFAEFF97F4A697C4FB7DBE89" box="[1164,1192,1469,1493]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="11" pageNumber="320">27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAEFF97F3079785FAC5BD4D" box="[813,1296,1532,1554]" pageId="11" pageNumber="320" type="reference_group">
<paragraph id="8B9C36D2FFAEFF97F3079785FAC5BD4D" blockId="11.[813,1296,1532,1554]" box="[813,1296,1532,1554]" pageId="11" pageNumber="320">
<treatmentCitationGroup id="AB3311FCFFAEFF97F3079785FAC5BD4D" box="[813,1296,1532,1554]" pageId="11" pageNumber="320">
<treatmentCitation id="0A8210C3FFAEFF97F3079785FB67BD4D" author="Hajek J" box="[813,1202,1532,1554]" page="120" pageId="11" pageNumber="320" year="2003">
<taxonomicName id="4C234D51FFAEFF97F3079785FB67BD4D" ID-CoL="3RMCZ" authority=": Hajek, 2003: 120" authorityName=": Hajek" authorityPageNumber="120" authorityYear="2003" box="[813,1202,1532,1554]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="B957EAC0FFAEFF97F3079785FC2FBD4E" box="[813,1018,1532,1554]" italics="true" pageId="11" pageNumber="320">Laccophilus wittmeri</emphasis>
:
<bibRefCitation id="EFB24B23FFAEFF97F4229785FB67BD4D" author="Hajek J" box="[1032,1202,1532,1554]" editor="Jach MA &amp; Ji L" firstAuthor="Hajek" journalOrPublisher="Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien." page="120" pageId="11" pageNumber="320" pagination="115 - 123" refId="ref15355" refString="Hajek J (2003) Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera). In: Jach MA &amp; Ji L (eds.) Water Beetles of China, Volume III. Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien. Pp. 115 - 123." title="Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera)" type="book chapter" volumeTitle="Water Beetles of China, Volume III" year="2003">Hájek, 2003: 120</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAEFF97F3079443FA5CB34E" pageId="11" pageNumber="320" type="materials_examined">
<paragraph id="8B9C36D2FFAEFF97F3079443FA79BD2E" blockId="11.[812,1452,1594,1650]" pageId="11" pageNumber="320">
<emphasis id="B957EAC0FFAEFF97F3079443FC14BD0E" bold="true" box="[813,961,1594,1618]" pageId="11" pageNumber="320">
<typeStatus id="54988870FFAEFF97F3079443FCB3BD0E" box="[813,870,1594,1618]" pageId="11" pageNumber="320">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3347642FFAEFF97F3EF9443FBD3BD0E" box="[965,1030,1594,1618]" name="China" pageId="11" pageNumber="320">China</collectingCountry>
,
<collectingRegion id="49E7F830FFAEFF97F4249443FB3EBD0E" box="[1038,1259,1594,1618]" country="China" name="Guangdong" pageId="11" pageNumber="320">Guangdong Province</collectingRegion>
,
<location id="8EFC6009FFAEFF97F4DE9443FBE2BD2E" LSID="urn:lsid:plazi:treatment:038A87C4FFAEFF90F38C97E4FCBBBA49:8EFC6009FFAEFF97F4DE9443FBE2BD2E" country="China" latitude="23.166666" longLatPrecision="1256" longitude="112.53333" name="Dinghu Mountain National Nature Reserve" pageId="11" pageNumber="320" stateProvince="Guangdong">Dinghu Mountain National Nature Reserve</location>
, ca.
<geoCoordinate id="EE175015FFAEFF97F44F9420FB14BD2E" box="[1125,1217,1625,1650]" degrees="23" direction="north" minutes="10" orientation="latitude" pageId="11" pageNumber="320" precision="925" value="23.166666">23°10′N</geoCoordinate>
,
<geoCoordinate id="EE175015FFAEFF97F4E69420FAE7BD2E" box="[1228,1330,1625,1650]" degrees="112" direction="east" minutes="32" orientation="longitude" pageId="11" pageNumber="320" precision="925" value="112.53333">112°32′E</geoCoordinate>
, ca.
<specimenCount id="9D25FD5BFFAEFF97F54B9423FA79BD2E" box="[1377,1452,1626,1650]" pageId="11" pageNumber="320" type="generic">
<quantity id="4CDB9B37FFAEFF97F54B9423FA7DBD2E" box="[1377,1448,1626,1650]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="320" unit="m" value="250.0">
<elevation id="000ED1E1FFAEFF97F54B9423FA7DBD2E" box="[1377,1448,1626,1650]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="320" unit="m" value="250.0">250 m</elevation>
</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8B9C36D2FFAEFF97F30794E3FA5CB34E" blockId="11.[813,1452,1690,2066]" pageId="11" pageNumber="320">
<emphasis id="B957EAC0FFAEFF97F30794E3FBCCBDEE" bold="true" box="[813,1049,1690,1714]" pageId="11" pageNumber="320">Material examined.</emphasis>
<materialsCitation id="3B4B3C8FFFAEFF97F40E94E3FACFBDAE" ID-GBIF-Occurrence="3760488437" collectingDate="2001-10-28" collectionCode="CASS" collectorName="Jach &amp; Komarek" country="China" elevation="250" location="Dinghu Nat. Res." pageId="11" pageNumber="320" specimenCode="CWBS 449" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong Prov." typeStatus="holotype">
<typeStatus id="54988870FFAEFF97F40E94E3FB59BDEE" box="[1060,1164,1690,1714]" pageId="11" pageNumber="320" type="holotype">Holotype</typeStatus>
<specimenCount id="9D25FD5BFFAEFF97F4B294E3FB04BDEE" box="[1176,1233,1690,1714]" pageId="11" pageNumber="320" type="male">male</specimenCount>
, labelled:
<collectingCountry id="F3347642FFAEFF97F56694E3FA72BDEE" box="[1356,1447,1690,1714]" name="China" pageId="11" pageNumber="320">CHINA</collectingCountry>
:
<collectingRegion id="49E7F830FFAEFF97F30794C3FC39BD8E" box="[813,1004,1722,1746]" country="China" name="Guangdong" pageId="11" pageNumber="320">Guangdong Prov.</collectingRegion>
/
<location id="8EFC6009FFAEFF97F42994C3FB15BD8E" LSID="urn:lsid:plazi:treatment:038A87C4FFAEFF90F38C97E4FCBBBA49:8EFC6009FFAEFF97F42994C3FB15BD8E" box="[1027,1216,1722,1746]" country="China" name="Dinghu Nat. Res." pageId="11" pageNumber="320" stateProvince="Guangdong Prov.">Dinghu Nat. Res.</location>
/
<date id="FF9D1012FFAEFF97F4FD94C3FA84BD8E" box="[1239,1361,1722,1746]" pageId="11" pageNumber="320" value="2001-10-28">
<collectingDate id="EFD9E9FAFFAEFF97F4FD94C3FA84BD8E" box="[1239,1361,1722,1746]" pageId="11" pageNumber="320" value="2001-10-28">28.10.2001</collectingDate>
</date>
, ca.
<quantity id="4CDB9B37FFAEFF97F5AE94C3FC97BDAE" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="320" unit="m" value="250.0">
<elevation id="000ED1E1FFAEFF97F5AE94C3FC97BDAE" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="320" unit="m" value="250.0">250 m</elevation>
</quantity>
/
<collectorName id="26D65304FFAEFF97F37794A3FC58BDAE" box="[861,909,1754,1778]" pageId="11" pageNumber="320">Jäch</collectorName>
&amp;
<collectorName id="26D65304FFAEFF97F39C94A3FBCFBDAE" box="[950,1050,1754,1778]" pageId="11" pageNumber="320">Komarek</collectorName>
(
<specimenCode id="DB859EA9FFAEFF97F40194A3FB78BDAE" box="[1067,1197,1754,1778]" collectionCode="CWBS" pageId="11" pageNumber="320">CWBS 449</specimenCode>
) (
<collectionCode id="ED32AE17FFAEFF97F4EC94A3FADBBDAE" box="[1222,1294,1754,1778]" country="China" httpUri="http://grbio.org/cool/3ndm-kie9" name="Chinese Academy of Sciences, Shenyang" pageId="11" pageNumber="320">CASS</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4B3C8FFFAEFF97F50E94A2FC21BC6E" ID-GBIF-Occurrence="3760488420" collectingDate="2001-10-28" collectionCode="NHMB, NMPC, NHMW" collectorName="Jach &amp; Komarek" country="China" elevation="250" location="Dinghu Nat. Res." pageId="11" pageNumber="320" specimenCode="CWBS 449" specimenCount="13" specimenCount-female="5" specimenCount-male="8" stateProvince="Guangdong Prov." typeStatus="paratype">
<typeStatus id="54988870FFAEFF97F50E94A2FA44BDAF" box="[1316,1425,1755,1779]" pageId="11" pageNumber="320" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D25FD5BFFAEFF97F5B494A3FCA5BC4E" pageId="11" pageNumber="320" type="male">8 males</specimenCount>
,
<specimenCount id="9D25FD5BFFAEFF97F3559483FC24BC4E" box="[895,1009,1786,1810]" pageId="11" pageNumber="320" type="female">5 females</specimenCount>
: same label data as holotype (
<collectionCode id="ED32AE17FFAEFF97F57B9483FA72BC4E" box="[1361,1447,1786,1810]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="11" pageNumber="320">NHMB</collectionCode>
,
<collectionCode id="ED32AE17FFAEFF97F3079563FCA8BC6E" box="[813,893,1818,1842]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="320">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFAEFF97F3A19563FC3CBC6E" box="[907,1001,1818,1842]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="320" type="Museum">NHMW</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4B3C8FFFAEFF97F3D59563FC3DBCEE" ID-GBIF-Occurrence="3760488348" collectingDate="2013-04-05" collectingDateMax="2013-04-09" collectingDateMin="2013-04-05" collectionCode="NMPC, PMCB, ZSMG" collectorName="M. Fikacek &amp; J. Hajek &amp; J. Ruzicka" country="China" elevation="300" latitude="21.906666" location="Shiwandashan National Forest" longLatPrecision="125" longitude="107.903336" municipality="Shiwandashan National Forest Park" pageId="11" pageNumber="320" specimenCount="15" specimenCount-female="7" specimenCount-male="8" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="9D25FD5BFFAEFF97F3D59563FB8CBC6E" box="[1023,1113,1818,1842]" pageId="11" pageNumber="320" type="male">8 males</specimenCount>
,
<specimenCount id="9D25FD5BFFAEFF97F4429562FB0CBC6E" box="[1128,1241,1818,1842]" pageId="11" pageNumber="320" type="female">7 females</specimenCount>
:
<collectingCountry id="F3347642FFAEFF97F4CD9563FA95BC6E" box="[1255,1344,1818,1842]" name="China" pageId="11" pageNumber="320">CHINA</collectingCountry>
,
<collectingRegion id="49E7F830FFAEFF97F5649563FA79BC6E" box="[1358,1452,1818,1842]" country="China" name="Guangxi" pageId="11" pageNumber="320">Guangxi</collectingRegion>
A.R.,
<date id="FF9D1012FFAEFF97F35B9543FC3ABC0E" box="[881,1007,1850,1874]" pageId="11" pageNumber="320" value="2013-04-05" valueMax="2013-04-09" valueMin="2013-04-05">
<collectingDate id="EFD9E9FAFFAEFF97F35B9543FC3ABC0E" box="[881,1007,1850,1874]" pageId="11" pageNumber="320" value="2013-04-05" valueMax="2013-04-09" valueMin="2013-04-05">5-9.iv.2013</collectingDate>
</date>
/
<collectingMunicipality id="6BF8ACA8FFAEFF97F4219543FA4FBC0E" box="[1035,1434,1850,1874]" pageId="11" pageNumber="320">
<location id="8EFC6009FFAEFF97F4219543FA8BBC0E" LSID="urn:lsid:plazi:treatment:038A87C4FFAEFF90F38C97E4FCBBBA49:8EFC6009FFAEFF97F4219543FA8BBC0E" box="[1035,1374,1850,1874]" country="China" latitude="21.906666" longLatPrecision="125" longitude="107.903336" municipality="Shiwandashan National Forest Park" name="Shiwandashan National Forest" pageId="11" pageNumber="320" stateProvince="Guangxi">Shiwandashan National Forest</location>
Park
</collectingMunicipality>
/ (small pools in drying-up stream) /
<geoCoordinate id="EE175015FFAEFF97F49B9520FAF4BC2E" box="[1201,1313,1881,1906]" degrees="21" direction="north" minutes="54.4" orientation="latitude" pageId="11" pageNumber="320" precision="92" value="21.906666">21°54.4′N</geoCoordinate>
,
<geoCoordinate id="EE175015FFAEFF97F5049520FA72BC2E" box="[1326,1447,1881,1906]" degrees="107" direction="east" minutes="54.2" orientation="longitude" pageId="11" pageNumber="320" precision="92" value="107.903336">107°54.2′E</geoCoordinate>
,
<quantity id="4CDB9B37FFAEFF97F3079503FCA3BCCE" box="[813,886,1914,1938]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="320" unit="m" value="300.0">
<elevation id="000ED1E1FFAEFF97F3079503FCA3BCCE" box="[813,886,1914,1938]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="320" unit="m" value="300.0">300 m</elevation>
</quantity>
/
<collectorName id="26D65304FFAEFF97F3BE9503FBC5BCCE" box="[916,1040,1914,1938]" pageId="11" pageNumber="320">M.Fikáček</collectorName>
,
<collectorName id="26D65304FFAEFF97F4359503FBA1BCCE" box="[1055,1140,1914,1938]" pageId="11" pageNumber="320">J.Hájek</collectorName>
&amp;
<collectorName id="26D65304FFAEFF97F4B59503FAD8BCCE" box="[1183,1293,1914,1938]" pageId="11" pageNumber="320">J.Růžička</collectorName>
leg. (
<collectionCode id="ED32AE17FFAEFF97F57E9503FA73BCCE" box="[1364,1446,1914,1938]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="320">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFAEFF97F30795E3FCA8BCEE" box="[813,893,1946,1970]" pageId="11" pageNumber="320">PMCB</collectionCode>
,
<collectionCode id="ED32AE17FFAEFF97F3A795E3FC35BCEE" box="[909,992,1946,1970]" pageId="11" pageNumber="320">ZSMG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFAEFF97F3DD95E3FA5CB34E" ID-GBIF-Occurrence="3760488344" collectingDate="2013-04-07" collectingDateMax="2013-04-08" collectingDateMin="2013-04-07" collectionCode="NMPC" collectorName="M. Fikacek &amp; J. Hajek &amp; J. Ruzicka" country="China" elevation="280" latitude="21.918333" location="Shiwandashan National Forest" longLatPrecision="125" longitude="107.915" municipality="Shiwandashan National Forest Park" pageId="11" pageNumber="320" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="9D25FD5BFFAEFF97F3DD95E3FB80BCEE" box="[1015,1109,1946,1970]" pageId="11" pageNumber="320" type="male">2 males</specimenCount>
,
<specimenCount id="9D25FD5BFFAEFF97F44F95E3FB05BCEE" box="[1125,1232,1946,1970]" pageId="11" pageNumber="320" type="female">1 female</specimenCount>
:
<collectingCountry id="F3347642FFAEFF97F4F595E3FAEFBCEE" box="[1247,1338,1946,1970]" name="China" pageId="11" pageNumber="320">CHINA</collectingCountry>
,
<collectingRegion id="49E7F830FFAEFF97F56195E3FA7EBCEE" box="[1355,1451,1946,1970]" country="China" name="Guangxi" pageId="11" pageNumber="320">Guangxi</collectingRegion>
A.R.,
<date id="FF9D1012FFAEFF97F34595C3FC3FBC8E" box="[879,1002,1978,2002]" pageId="11" pageNumber="320" value="2013-04-07" valueMax="2013-04-08" valueMin="2013-04-07">
<collectingDate id="EFD9E9FAFFAEFF97F34595C3FC3FBC8E" box="[879,1002,1978,2002]" pageId="11" pageNumber="320" value="2013-04-07" valueMax="2013-04-08" valueMin="2013-04-07">7-8.iv.2013</collectingDate>
</date>
/ stream valley ca. 2 NE north gate of /
<collectingMunicipality id="6BF8ACA8FFAEFF97F30795A3FB71BCAE" box="[813,1188,2010,2034]" pageId="11" pageNumber="320">
<location id="8EFC6009FFAEFF97F30795A3FBBBBCAE" LSID="urn:lsid:plazi:treatment:038A87C4FFAEFF90F38C97E4FCBBBA49:8EFC6009FFAEFF97F30795A3FBBBBCAE" box="[813,1134,2010,2034]" country="China" latitude="21.918333" longLatPrecision="125" longitude="107.915" municipality="Shiwandashan National Forest Park" name="Shiwandashan National Forest" pageId="11" pageNumber="320" stateProvince="Guangxi">Shiwandashan National Forest</location>
Park
</collectingMunicipality>
/
<geoCoordinate id="EE175015FFAEFF97F49F95A0FAF1BCAE" box="[1205,1316,2009,2034]" degrees="21" direction="north" minutes="55.1" orientation="latitude" pageId="11" pageNumber="320" precision="92" value="21.918333">21°55.1′N</geoCoordinate>
,
<geoCoordinate id="EE175015FFAEFF97F50495A0FA72BCAE" box="[1326,1447,2009,2034]" degrees="107" direction="east" minutes="54.9" orientation="longitude" pageId="11" pageNumber="320" precision="92" value="107.915">107°54.9′E</geoCoordinate>
,
<quantity id="4CDB9B37FFAEFF97F3079583FCA7B34E" box="[813,882,2042,2066]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="320" unit="m" value="280.0">
<elevation id="000ED1E1FFAEFF97F3079583FCA7B34E" box="[813,882,2042,2066]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="320" unit="m" value="280.0">280 m</elevation>
</quantity>
/
<collectorName id="26D65304FFAEFF97F3A19583FBD5B34E" box="[907,1024,2042,2066]" pageId="11" pageNumber="320">M.Fikáček</collectorName>
,
<collectorName id="26D65304FFAEFF97F4279583FB8BB34E" box="[1037,1118,2042,2066]" pageId="11" pageNumber="320">J.Hájek</collectorName>
&amp;
<collectorName id="26D65304FFAEFF97F4AE9583FB39B34E" box="[1156,1260,2042,2066]" pageId="11" pageNumber="320">J.Růžička</collectorName>
leg. (
<collectionCode id="ED32AE17FFAEFF97F5069583FAABB34E" box="[1324,1406,2042,2066]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="320">NMPC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F0A292C4FE37BAC9" pageId="12" pageNumber="321" type="diagnosis">
<paragraph id="8B9C36D2FFA9FF90F0A292C4FE37BAC9" blockId="12.[136,775,189,405]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F0A292C4FF28BB89" bold="true" box="[136,253,189,213]" pageId="12" pageNumber="321">Diagnosis.</emphasis>
Large, broadly oval species (body length
<quantity id="4CDB9B37FFA9FF90F29D92C4FF63BBA9" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="4.9" metricValueMin="4.5" pageId="12" pageNumber="321" unit="mm" value="4.7" valueMax="4.9" valueMin="4.5">4.5–4.9 mm</quantity>
). A species closely related to
<taxonomicName id="4C234D51FFA9FF90F22B92A7FDACBBA9" authorityName=": Hajek" authorityYear="2003" box="[513,633,222,245]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="B957EAC0FFA9FF90F22B92A7FDACBBA9" box="[513,633,222,245]" italics="true" pageId="12" pageNumber="321">L. wittmeri</emphasis>
</taxonomicName>
, from which it can be distinguished by constantly testaceous colouration of medial part of elytra, yellowish subbasal and preapical elytral band bordered with thin blackish stripes, and by different shape of the apical part of the median lobe, which is straighter than in
<taxonomicName id="4C234D51FFA9FF90F14E9307FE0EBAC9" authorityName=": Hajek" authorityYear="2003" box="[356,475,382,405]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="B957EAC0FFA9FF90F14E9307FE0EBAC9" box="[356,475,382,405]" italics="true" pageId="12" pageNumber="321">L. wittmeri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F0A293C4FE22BDC9" pageId="12" pageNumber="321" type="description">
<paragraph id="8B9C36D2FFA9FF90F0A293C4FCDDB909" blockId="12.[136,776,445,597]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F0A293C4FEC6BA89" bold="true" box="[136,275,445,469]" pageId="12" pageNumber="321">Description.</emphasis>
Colouration: Head and pronotum testaceous. Elytra largely testaceous, sinuous transverse subbasal and preapical bands only slightly paler than ground colour of elytra, but well recognisable because of their distinct blackish bordering (
<figureCitation id="13182A57FFA9FF90F0D59044FE97B909" box="[255,322,573,597]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 1–9. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="12" pageNumber="321">Fig. 9</figureCitation>
). Appendages and ventral part testaceous.
</paragraph>
<paragraph id="8B9C36D2FFA9FF90F0A29004FCDDBFE9" blockId="12.[136,777,637,1205]" pageId="12" pageNumber="321">Surface structures and sculpture: Head reticulation simple, composed of large irregular polygonal meshes, which are moderately deeply impressed. Frons between eyes, and clypeus with several large punctures. Reticulation of pronotum like that of head. Pronotum along margins with numerous large setigerous punctures. Reticulation of elytra simple, composed of large irregular polygonal meshes similar to those of head or pronotum. Meshes mostly complete, inner space often with several small punctures. Each elytron with indistinct, sutural, discal and lateral row of large serial setigerous punctures. Ventral part reticulated with fine elongate meshes. Prosternal apophysis lanceolate. Abdominal ventrites with numerous oblique grooves and several large setigerous punctures, lateral margin bordered. Apical ventrite rounded posteriorly, with large setigerous punctures and transverse grooves on disc. Male pro- and mesotarsomeres 1–3 weakly dilated with bundle of setae on ventral side. Claws of pro- and mesotarsi simple and equal.</paragraph>
<paragraph id="8B9C36D2FFA9FF90F0A296A4FF2FBE49" blockId="12.[136,775,1245,1301]" pageId="12" pageNumber="321">
Male genitalia: Median lobe of aedeagus in lateral view as in
<figureCitation id="13182A57FFA9FF90F08F9684FF23BE49" box="[165,246,1277,1301]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="12" pageNumber="321">Fig. 27</figureCitation>
.
</paragraph>
<paragraph id="8B9C36D2FFA9FF90F0A29744FED6BE29" blockId="12.[136,775,1341,1397]" pageId="12" pageNumber="321">Female genitalia: Similar to male; pro- and mesotarsomeres not dilated.</paragraph>
<paragraph id="8B9C36D2FFA9FF90F0A297E4FEE2BD49" blockId="12.[136,775,1437,1557]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F0A297E4FED9BEE9" bold="true" box="[136,268,1437,1461]" pageId="12" pageNumber="321">Variability.</emphasis>
Specimens of
<typeStatus id="54988870FFA9FF90F18697E7FE0FBEEA" box="[428,474,1438,1462]" pageId="12" pageNumber="321">type</typeStatus>
series varying considerably in body colouration, in some specimens, longitudinal blackish stripe divides the subbasal band into two separate yellowish spots.
</paragraph>
<paragraph id="8B9C36D2FFA9FF90F0A29447FE22BDC9" blockId="12.[136,775,1597,1685]" pageId="12" pageNumber="321">
Measurements (N= 18): Body length
<quantity id="4CDB9B37FFA9FF90F2189444FD6DBD09" box="[562,696,1597,1621]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="4.9" metricValueMin="4.5" pageId="12" pageNumber="321" unit="mm" value="4.7" valueMax="4.9" valueMin="4.5">4.5–4.9 mm</quantity>
(mean value
<quantity id="4CDB9B37FFA9FF90F0E39424FECBBD29" box="[201,286,1629,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="12" pageNumber="321" unit="mm" value="4.7">4.7 mm</quantity>
,
<typeStatus id="54988870FFA9FF90F1009424FE52BD29" box="[298,391,1629,1653]" pageId="12" pageNumber="321" type="holotype">holotype</typeStatus>
<quantity id="4CDB9B37FFA9FF90F1A49424FE30BD29" box="[398,485,1629,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="12" pageNumber="321" unit="mm" value="4.8">4.8 mm</quantity>
); width
<quantity id="4CDB9B37FFA9FF90F2169424FD68BD29" box="[572,701,1629,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="12" pageNumber="321" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
(mean value
<quantity id="4CDB9B37FFA9FF90F0E09404FEF5BDC9" box="[202,288,1661,1685]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="12" pageNumber="321" unit="mm" value="2.7">2.7 mm</quantity>
,
<typeStatus id="54988870FFA9FF90F1079404FE5FBDC9" box="[301,394,1661,1685]" pageId="12" pageNumber="321" type="holotype">holotype</typeStatus>
<quantity id="4CDB9B37FFA9FF90F1B99404FE3EBDC9" box="[403,491,1661,1685]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="321" unit="mm" value="2.8">2.8 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F0A294C4FE98BC49" pageId="12" pageNumber="321" type="etymology">
<paragraph id="8B9C36D2FFA9FF90F0A294C4FE98BC49" blockId="12.[136,776,1725,1813]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F0A294C4FEDABD89" bold="true" box="[136,271,1725,1749]" pageId="12" pageNumber="321">Etymology.</emphasis>
The new species is dedicated to one of its collectors, Albrecht Komarek (Mödling,
<collectingCountry id="F3347642FFA9FF90F21994A4FD50BDA9" box="[563,645,1757,1781]" name="Austria" pageId="12" pageNumber="321">Austria</collectingCountry>
), a specialist on
<taxonomicName id="4C234D51FFA9FF90F0819484FE9CBC49" authorityName="Latreille" authorityYear="1802" box="[171,329,1789,1813]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F0A29544FE8CB369" pageId="12" pageNumber="321" type="biology_ecology">
<paragraph id="8B9C36D2FFA9FF90F0A29544FE8CB369" blockId="12.[136,776,1853,2101]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F0A29544FF3DBC09" bold="true" box="[136,232,1853,1877]" pageId="12" pageNumber="321">Habitat.</emphasis>
At the
<typeStatus id="54988870FFA9FF90F1699547FEA7BC0A" box="[323,370,1854,1878]" pageId="12" pageNumber="321">type</typeStatus>
locality,
<taxonomicName id="4C234D51FFA9FF90F1F59544FD06BC09" authorityName="Hájek &amp; Brancucci" authorityYear="2015" box="[479,723,1853,1877]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="komareki">
<emphasis id="B957EAC0FFA9FF90F1F59544FD06BC09" box="[479,723,1853,1877]" italics="true" pageId="12" pageNumber="321">Laccophilus komareki</emphasis>
</taxonomicName>
was collected in a stream, ca.
<quantity id="4CDB9B37FFA9FF90F1819524FDD2BC29" box="[427,519,1885,1909]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="12" pageNumber="321" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 m</quantity>
wide, flowing through dense primary forest, geology: sandstone (
<bibRefCitation id="EFB24B23FFA9FF90F2649504FD29BCC9" author="Jach MA &amp; Ji L" box="[590,764,1917,1941]" editor="Jach MA &amp; Ji L" firstAuthor="Jach" journalOrPublisher="Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien." pageId="12" pageNumber="321" pagination="1 - 20" refId="ref15460" refString="Jach MA &amp; Ji L (2003) China water beetles survey (1999 - 2001). In: Jach MA &amp; Ji L (eds.) Water Beetles of China, Volume III. Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien. Pp. 1 - 20." title="China water beetles survey (1999 - 2001)" type="book chapter" volumeTitle="Water Beetles of China, Volume III" year="2003">Jäch &amp; Ji, 2003</bibRefCitation>
). In
<collectingRegion id="49E7F830FFA9FF90F08095E4FED9BCE9" box="[170,268,1949,1973]" country="China" name="Guangxi" pageId="12" pageNumber="321">Guangxi</collectingRegion>
, the new species was collected in a shaded deep remnant pool (diameter ca.
<quantity id="4CDB9B37FFA9FF90F1C495C4FDCCBC89" box="[494,537,1981,2005]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="321" unit="m" value="1.0">1 m</quantity>
) of a drying up forest stream, the bottom with a thick layer of decaying leaves; and in small rocky pools (diameter ca.
<quantity id="4CDB9B37FFA9FF90F2149584FD51B349" box="[574,644,2045,2069]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="321" unit="cm" value="30.0">30 cm</quantity>
) in a forest streamlet (
<figureCitation id="13182A57FFA9FF90F0D09A64FE99B369" box="[250,332,2077,2101]" captionStart="Figs" captionStartId="8.[135,179,2035,2057]" captionTargetBox="[187,1423,189,2020]" captionTargetId="figure-8@8.[187,1423,189,2020]" captionTargetPageId="8" captionText="Figs. 36–39. Habitats: 36, E Kalimantan, Muara Ritan (type locality of Laccophilus kalimantanensis, new species); 37, China, Guangxi, Shiwandashan (locality of L. komareki, new species); 38, E Kalimantan, Sungai Wain (locality of L. mahakamensis); 39, Laos, Attapeu (locality of L. tonkinensis)." figureDoi="http://doi.org/10.5281/zenodo.5385206" httpUri="https://zenodo.org/record/5385206/files/figure.png" pageId="12" pageNumber="321">Fig. 37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F36992C4FCBBBA49" pageId="12" pageNumber="321" type="distribution">
<paragraph id="8B9C36D2FFA9FF90F36992C4FCBBBA49" blockId="12.[835,1475,189,277]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F36992C4FC09BB89" bold="true" box="[835,988,189,213]" pageId="12" pageNumber="321">Distribution.</emphasis>
So far known only from two localities in southern
<collectingCountry id="F3347642FFA9FF90F38392A4FC3CBBA9" box="[937,1001,221,245]" name="China" pageId="12" pageNumber="321">China</collectingCountry>
(
<collectingRegion id="49E7F830FFA9FF90F3D392A4FBA2BBA9" box="[1017,1143,221,245]" country="China" name="Guangdong" pageId="12" pageNumber="321">Guangdong</collectingRegion>
and
<collectingRegion id="49E7F830FFA9FF90F49A92A4FAD9BBA9" box="[1200,1292,221,245]" country="China" name="Guangxi" pageId="12" pageNumber="321">Guangxi</collectingRegion>
provinces) (
<figureCitation id="13182A57FFA9FF90F5BD92A4FCB7BA49" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 40–41. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="12" pageNumber="321">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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