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<document id="E8AF289FA714E8012C21AB3FB279F17E" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFAAFF8CF3AF97A3FDC5B8C9" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus tonkinensis Brancucci 1983" docType="treatment" docVersion="8" lastPageNumber="325" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="324" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B01E7520D7161BABCF53B4A960AA5F47" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="795F302DCBABD32B99BB936C56C0E62D">
<mods:title id="2A6FC05B3BE975681B68E7B7AB823268">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="E0E4C6561691CBE79666B726CD5F340B" type="personal">
<mods:role id="26C3B0FEF3674B86EEFA6FE78E80B8C2">
<mods:roleTerm id="571634774CEF08F40491C6ED11529D5E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9ABDEF8A73BCC4D1A1FED80909F297B3">Hájek, Jiří</mods:namePart>
<mods:affiliation id="B433DBF145044E9DA5F1143725316095">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 – Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="275F5F618E3878E47EC617930A267501" type="email">jiri_hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="289AF478AFB5FC55970C75B25778BCB0" type="personal">
<mods:role id="247C6BDD80E8F29FDA65E85331D923B6">
<mods:roleTerm id="A030B734B06A05510CAC11AB755C9BD6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A84908BDDAF20818477130BB5FA809B7">Brancucci, Michel</mods:namePart>
<mods:affiliation id="0265D3B65C27790E943ABB7A951F5926">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="DC16E2E16CAD4CFF442CCFB8B78B530A">text</mods:typeOfResource>
<mods:relatedItem id="BFDD4BBE78EF80CF57F2D10A60E9D6BA" type="host">
<mods:titleInfo id="2F2C7E5EBC8B2473051357253950C7D5">
<mods:title id="5EC112DD1667F710280AC5720B2758DB">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="38CB3EE7B9BC37503CD081CC2D165A0C">
<mods:date id="F6AD0EDFD8BFFDE949AFC5A3556BDB6C">2015</mods:date>
<mods:detail id="5A4F9782A59853ED028A7202EED9BFD6" type="pubDate">
<mods:number id="CD4B56180EDF39694CEF242B52D5D68B">2015-08-06</mods:number>
</mods:detail>
<mods:detail id="7276DE9F6CCFFC5137A405CEE1F1AFA6" type="volume">
<mods:number id="8486C875ECD892E16A847907CEF7A75F">63</mods:number>
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<mods:start id="8B71606E8398AA2A2A82A29482491221">309</mods:start>
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<mods:classification id="C2C2D8C44EE059D554EB14837D139258">journal article</mods:classification>
<mods:identifier id="C7EF309BFDD78A7645849A6111E7B1CE" type="CLB-Dataset">55750</mods:identifier>
<mods:identifier id="85F984177287F3566D2321F571EA6E40" type="DOI">10.5281/zenodo.5385198</mods:identifier>
<mods:identifier id="10A54A572C6EBE306C9E914B3FB96AC9" type="GBIF-Dataset">c7f95b94-53f8-420d-97c5-4e27d7c211bf</mods:identifier>
<mods:identifier id="8A2F7350FFF2182539522F9AC8A7C276" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="934648A92EE629C13F034FFAA8DD86FD" type="Zenodo-Dep">5385198</mods:identifier>
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<treatment id="038A87C4FFAAFF8CF3AF97A3FDC5B8C9" ID-DOI="http://doi.org/10.5281/zenodo.6512187" ID-GBIF-Taxon="194955229" ID-Zenodo-Dep="6512187" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF8CF3AF97A3FDC5B8C9" httpUri="http://treatment.plazi.org/id/038A87C4FFAAFF8CF3AF97A3FDC5B8C9" lastPageId="16" lastPageNumber="325" pageId="15" pageNumber="324">
<subSubSection id="C3396559FFAAFF93F3AF97A3FA80BEAE" box="[901,1365,1498,1522]" pageId="15" pageNumber="324" type="nomenclature">
<paragraph id="8B9C36D2FFAAFF93F3AF97A3FA80BEAE" blockId="15.[901,1365,1498,1554]" box="[901,1365,1498,1522]" pageId="15" pageNumber="324">
<heading id="D0D481BEFFAAFF93F3AF97A3FA80BEAE" bold="true" box="[901,1365,1498,1522]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="324" reason="2">
<taxonomicName id="4C234D51FFAAFF93F3AF97A3FA80BEAE" ID-CoL="6NRN5" authority="Brancucci, 1983" authorityName="Brancucci" authorityYear="1983" box="[901,1365,1498,1522]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="324" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B957EAC0FFAAFF93F3AF97A3FA80BEAE" bold="true" box="[901,1365,1498,1522]" pageId="15" pageNumber="324">
<emphasis id="B957EAC0FFAAFF93F3AF97A3FB47BEAE" bold="true" box="[901,1170,1498,1522]" italics="true" pageId="15" pageNumber="324">Laccophilus tonkinensis</emphasis>
<bibRefCitation id="EFB24B23FFAAFF93F4B797A3FA80BEAE" author="Brancucci M" box="[1181,1365,1498,1522]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="15" pageNumber="324" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAAFF93F4089783FB63BD4E" box="[1058,1206,1530,1554]" pageId="15" pageNumber="324" type="description">
<paragraph id="8B9C36D2FFAAFF93F4089783FB63BD4E" blockId="15.[901,1365,1498,1554]" box="[1058,1206,1530,1554]" pageId="15" pageNumber="324">
(
<figureCitation id="13182A57FFAAFF93F4009783FB50BD4E" box="[1066,1157,1530,1554]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="15" pageNumber="324">Figs. 16</figureCitation>
,
<figureCitation id="13182A57FFAAFF93F4B89783FB7BBD4E" box="[1170,1198,1530,1554]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="15" pageNumber="324">34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAAFF93F3079440FB45BD37" pageId="15" pageNumber="324" type="reference_group">
<paragraph id="8B9C36D2FFAAFF93F3079440FB45BD37" blockId="15.[813,1452,1593,1643]" pageId="15" pageNumber="324">
<treatmentCitationGroup id="AB3311FCFFAAFF93F3079440FB45BD37" pageId="15" pageNumber="324">
<treatmentCitation id="0A8210C3FFAAFF93F3079440FB0EBD13" author="Brancucci M" box="[813,1243,1593,1615]" page="258" pageId="15" pageNumber="324" year="1983">
<taxonomicName id="4C234D51FFAAFF93F3079440FB0EBD13" ID-CoL="6NRN5" authority="Brancucci, 1983: 258" authorityName="Brancucci" authorityPageNumber="258" authorityYear="1983" box="[813,1243,1593,1615]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="324" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B957EAC0FFAAFF93F3079440FBDBBD12" box="[813,1038,1593,1614]" italics="true" pageId="15" pageNumber="324">Laccophilus tonkinensis</emphasis>
<bibRefCitation id="EFB24B23FFAAFF93F4399440FB0EBD13" author="Brancucci M" box="[1043,1243,1593,1615]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" page="258" pageId="15" pageNumber="324" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983: 258</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="0A8210C3FFAAFF93F378942CFC29BD37" author="Hajek J" box="[850,1020,1621,1643]" page="120" pageId="15" pageNumber="324" year="2003">
<bibRefCitation id="EFB24B23FFAAFF93F378942CFC29BD37" author="Hajek J" box="[850,1020,1621,1643]" editor="Jach MA &amp; Ji L" firstAuthor="Hajek" journalOrPublisher="Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien." page="120" pageId="15" pageNumber="324" pagination="115 - 123" refId="ref15355" refString="Hajek J (2003) Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera). In: Jach MA &amp; Ji L (eds.) Water Beetles of China, Volume III. Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien. Pp. 115 - 123." title="Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera)" type="book chapter" volumeTitle="Water Beetles of China, Volume III" year="2003">Hájek, 2003: 120</bibRefCitation>
</treatmentCitation>
(new records).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAAFF8CF30794EDFD98BA29" lastPageId="16" lastPageNumber="325" pageId="15" pageNumber="324" type="materials_examined">
<paragraph id="8B9C36D2FFAAFF93F30794EDFC4EBD97" blockId="15.[813,1451,1683,1740]" pageId="15" pageNumber="324">
<emphasis id="B957EAC0FFAAFF93F30794EDFC18BDF7" bold="true" box="[813,973,1683,1708]" pageId="15" pageNumber="324">
<typeStatus id="54988870FFAAFF93F30794EDFCBDBDF0" box="[813,872,1684,1708]" pageId="15" pageNumber="324">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3347642FFAAFF93F3F294EDFBFCBDF0" box="[984,1065,1684,1708]" name="Vietnam" pageId="15" pageNumber="324">Tonkin</collectingCountry>
,
<collectingRegion id="49E7F830FFAAFF93F41D94EDFB71BDF0" box="[1079,1188,1684,1708]" country="Vietnam" name="Hoa Binh" pageId="15" pageNumber="324">Hoa Binh</collectingRegion>
<materialsCitation id="3B4B3C8FFFAAFF93F48494EDFC40BD97" ID-GBIF-Occurrence="3760488363" country="Vietnam" isTailEnumeration="true" location="Hoa Binh" pageId="15" pageNumber="324" specimenCount="1" stateProvince="Hoa Binh" typeStatus="holotype">
[=
<collectingCountry id="F3347642FFAAFF93F4FB94EDFAE4BDF0" box="[1233,1329,1684,1708]" name="Vietnam" pageId="15" pageNumber="324">Vietnam</collectingCountry>
,
<collectingRegion id="49E7F830FFAAFF93F51594EDFC59BD90" country="Vietnam" name="Hoa Binh" pageId="15" pageNumber="324">Hoa Binh Province</collectingRegion>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8B9C36D2FFAAFF8CF307948AFD98BA29" blockId="15.[813,1452,1779,2092]" lastBlockId="16.[136,775,189,373]" lastPageId="16" lastPageNumber="325" pageId="15" pageNumber="324">
<emphasis id="B957EAC0FFAAFF93F307948AFBDFBC57" bold="true" box="[813,1034,1779,1803]" pageId="15" pageNumber="324">Material examined.</emphasis>
<specimenCount id="9D25FD5BFFAAFF93F425948DFB77BC50" box="[1039,1186,1780,1804]" pageId="15" pageNumber="324" type="generic">17 specimens</specimenCount>
.
<materialsCitation id="3B4B3C8FFFAAFF93F481948AFB15BC30" ID-GBIF-Occurrence="3760488359" collectionCode="NMPC, ZSMG" collectorName="M. Fikacek &amp; S. Zhao" country="China" elevation="650" location="Hainan" pageId="15" pageNumber="324" specimenCount="4" stateProvince="Hainan">
<emphasis id="B957EAC0FFAAFF93F481948AFAD1BC57" bold="true" box="[1195,1284,1779,1803]" pageId="15" pageNumber="324">
<collectingCountry id="F3347642FFAAFF93F481948AFAD1BC57" box="[1195,1284,1779,1803]" name="China" pageId="15" pageNumber="324">CHINA</collectingCountry>
</emphasis>
:
<emphasis id="B957EAC0FFAAFF93F53B948AFAB0BC57" bold="true" box="[1297,1381,1779,1803]" pageId="15" pageNumber="324">
<collectingRegion id="49E7F830FFAAFF93F53B948AFAB0BC57" box="[1297,1381,1779,1803]" country="China" name="Hainan" pageId="15" pageNumber="324">Hainan</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFAAFF93F558948DFA72BC57" box="[1394,1447,1780,1804]" pageId="15" pageNumber="324" type="generic">4 ex.</specimenCount>
,
<location id="8EFC6009FFAAFF93F307956DFC04BC77" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF8CF3AF97A3FDC5B8C9:8EFC6009FFAAFF93F307956DFC04BC77" box="[813,977,1812,1836]" country="China" name="Limushan Mts." pageId="15" pageNumber="324" stateProvince="Hainan">Limushan Mts.</location>
, exposed pools along road, partly connected with stream, 19°9.1–2′N, 109°45–46′E,
<quantity id="4CDB9B37FFAAFF93F4FE954AFA9BBC17" box="[1236,1358,1843,1867]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="15" pageNumber="324" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">
<elevation id="000ED1E1FFAAFF93F4FE954AFA9BBC17" box="[1236,1358,1843,1867]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="15" pageNumber="324" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">550–750 m</elevation>
</quantity>
, coll.
<collectorName id="26D65304FFAAFF93F5A7954DFC54BC30" pageId="15" pageNumber="324">M. Fikáček</collectorName>
&amp;
<collectorName id="26D65304FFAAFF93F382952DFC28BC30" box="[936,1021,1876,1900]" pageId="15" pageNumber="324">S. Zhao</collectorName>
(
<collectionCode id="ED32AE17FFAAFF93F427952DFB88BC30" box="[1037,1117,1876,1900]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="15" pageNumber="324">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFAAFF93F440952DFB6CBC30" box="[1130,1209,1876,1900]" pageId="15" pageNumber="324">ZSMG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFAAFF93F4E4952DFB35BCB0" ID-GBIF-Occurrence="3760488322" collectingDate="2011-05-10" collectionCode="NMPC" collectorName="M. Fikacek &amp; S. Zhao" country="China" county="Jianfengling Mts." elevation="885" location="Tianchi" municipality="Tianchi Lake" pageId="15" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
<specimenCount id="9D25FD5BFFAAFF93F4E4952DFACFBC30" box="[1230,1306,1876,1900]" pageId="15" pageNumber="324" type="male">1 male</specimenCount>
,
<collectingCounty id="62FD4E5EFFAAFF93F502952DFCB5BCD7" pageId="15" pageNumber="324">Jianfengling Mts.</collectingCounty>
,
<collectingMunicipality id="6BF8ACA8FFAAFF93F344950DFBD5BCD0" box="[878,1024,1908,1932]" pageId="15" pageNumber="324">Tianchi Lake</collectingMunicipality>
env., rd from
<location id="8EFC6009FFAAFF93F48D950DFB2CBCD0" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF8CF3AF97A3FDC5B8C9:8EFC6009FFAAFF93F48D950DFB2CBCD0" box="[1191,1273,1908,1932]" country="China" county="Jianfengling Mts." municipality="Tianchi Lake" name="Tianchi" pageId="15" pageNumber="324" stateProvince="Hainan">Tianchi</location>
vill.
<location id="8EFC6009FFAAFF93F512950DFA83BCD7" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF8CF3AF97A3FDC5B8C9:8EFC6009FFAAFF93F512950DFA83BCD7" box="[1336,1366,1908,1931]" country="China" county="Jianfengling Mts." municipality="Tianchi Lake" name="To" pageId="15" pageNumber="324" stateProvince="Hainan">To</location>
“sector 5”, pools in shaded drying-up stream in primary forest, 18°43.6–44.1′N, 108°52.1–5′E,
<quantity id="4CDB9B37FFAAFF93F4A295CAFAD0BC97" box="[1160,1285,1971,1995]" metricMagnitude="2" metricUnit="m" metricValue="8.85" metricValueMax="9.5" metricValueMin="8.2" pageId="15" pageNumber="324" unit="m" value="885.0" valueMax="950.0" valueMin="820.0">
<elevation id="000ED1E1FFAAFF93F4A295CAFAD0BC97" box="[1160,1285,1971,1995]" metricMagnitude="2" metricUnit="m" metricValue="8.85" metricValueMax="9.5" metricValueMin="8.2" pageId="15" pageNumber="324" unit="m" value="885.0" valueMax="950.0" valueMin="820.0">820–950 m</elevation>
</quantity>
,
<date id="FF9D1012FFAAFF93F53895CDFA7DBC90" box="[1298,1448,1972,1996]" pageId="15" pageNumber="324" value="2011-05-10">
<collectingDate id="EFD9E9FAFFAAFF93F53895CDFA7DBC90" box="[1298,1448,1972,1996]" pageId="15" pageNumber="324" value="2011-05-10">10 May 2011</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFAAFF93F34395ADFC3EBCB0" box="[873,1003,2004,2028]" pageId="15" pageNumber="324">M. Fikáček</collectorName>
&amp;
<collectorName id="26D65304FFAAFF93F43C95ADFBA4BCB0" box="[1046,1137,2004,2028]" pageId="15" pageNumber="324">S. Zhao</collectorName>
(
<collectionCode id="ED32AE17FFAAFF93F4A995ADFB0DBCB0" box="[1155,1240,2004,2028]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="15" pageNumber="324">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFAAFF8CF4C495AAFE07BB89" ID-GBIF-Occurrence="3760488317" collectingDate="2010-04-29" collectionCode="NMPC" collectorName="J. Hajek" country="Laos" elevation="665" lastPageId="16" lastPageNumber="325" latitude="17.81" location="No." longLatPrecision="127" longitude="107.39" pageId="15" pageNumber="324" specimenCount="3" specimenCount-male="3" stateProvince="Attapu">
<emphasis id="B957EAC0FFAAFF93F4C495AAFAEDBCB7" bold="true" box="[1262,1336,2003,2027]" pageId="15" pageNumber="324">
<collectingCountry id="F3347642FFAAFF93F4C495AAFAEDBCB7" box="[1262,1336,2003,2027]" name="Laos" pageId="15" pageNumber="324">LAOS</collectingCountry>
</emphasis>
:
<emphasis id="B957EAC0FFAAFF93F56795AAFC47B357" bold="true" pageId="15" pageNumber="324">
<collectingRegion id="49E7F830FFAAFF93F56795AAFC47B357" country="Laos" name="Attapu" pageId="15" pageNumber="324">Attapeu Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFAAFF93F388958DFC2CB350" box="[930,1017,2036,2060]" pageId="15" pageNumber="324" type="male">3 males</specimenCount>
, road
<location id="8EFC6009FFAAFF93F414958DFBB3B357" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF8CF3AF97A3FDC5B8C9:8EFC6009FFAAFF93F414958DFBB3B357" box="[1086,1126,2036,2059]" country="Laos" latitude="17.81" longLatPrecision="127" longitude="107.39" name="No." pageId="15" pageNumber="324" stateProvince="Attapu">No.</location>
18, km 80 E
<collectingRegion id="49E7F830FFAAFF93F52A958DFA82B350" box="[1280,1367,2036,2060]" country="Laos" name="Attapu" pageId="15" pageNumber="324">Attapeu</collectingRegion>
, stream near the road,
<geoCoordinate id="EE175015FFAAFF93F3E79A6BFBE8B377" box="[973,1085,2066,2091]" degrees="17" direction="north" minutes="48.6" orientation="latitude" pageId="15" pageNumber="324" precision="92" value="17.81">17°48.6′N</geoCoordinate>
,
<geoCoordinate id="EE175015FFAAFF93F4609A6BFB11B377" box="[1098,1220,2066,2091]" degrees="107" direction="east" minutes="23.4" orientation="longitude" pageId="15" pageNumber="324" precision="92" value="107.39">107°23.4′E</geoCoordinate>
, ca.
<quantity id="4CDB9B37FFAAFF93F4D39A6AFA94B377" box="[1273,1345,2067,2091]" metricMagnitude="2" metricUnit="m" metricValue="6.65" pageId="15" pageNumber="324" unit="m" value="665.0">
<elevation id="000ED1E1FFAAFF93F4D39A6AFA94B377" box="[1273,1345,2067,2091]" metricMagnitude="2" metricUnit="m" metricValue="6.65" pageId="15" pageNumber="324" unit="m" value="665.0">665 m</elevation>
</quantity>
,
<date id="FF9D1012FFAAFF8CF5659A6DFF14BB89" lastPageId="16" lastPageNumber="325" pageId="15" pageNumber="324" value="2010-04-29">
<collectingDate id="EFD9E9FAFFAAFF8CF5659A6DFF14BB89" lastPageId="16" lastPageNumber="325" pageId="15" pageNumber="324" value="2010-04-29">29 April 2010</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFB5FF8CF12392C7FEB3BB89" box="[265,358,189,213]" pageId="16" pageNumber="325">J. Hájek</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF15D92C4FE1FBB89" box="[375,458,189,213]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="325">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFB5FF8CF1CA92C4FDA2BA09" ID-GBIF-Occurrence="3760488353" collectingDate="2012-05-18" collectingDateMax="2012-05-21" collectingDateMin="2012-05-18" collectionCode="NHMB" collectorName="M. Brancucci &amp; M. Geiser &amp; K. Phanthavong &amp; S. Xayalath" elevation="675" latitude="17.950000" location="Ban Navang" longitude="105.216667" municipality="Nakai-Nam Theun" pageId="16" pageNumber="325" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Khammouan Province">
<emphasis id="B957EAC0FFB5FF8CF1CA92C4FD32BB89" bold="true" box="[480,743,189,213]" pageId="16" pageNumber="325">
<collectingRegion id="49E7F830FFB5FF8CF1CA92C4FD32BB89" box="[480,743,189,213]" country="Laos" name="Khammouan" pageId="16" pageNumber="325">Khammouan Province</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFB5FF8CF2D392C4FF69BBA9" pageId="16" pageNumber="325" type="male">1 male</specimenCount>
<specimenCount id="9D25FD5BFFB5FF8CF0E992A4FEF8BBA9" box="[195,301,221,245]" pageId="16" pageNumber="325" type="female">4 females</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFB5FF8CF11D92A4FE2BBBA9" box="[311,510,221,245]" pageId="16" pageNumber="325">Nakai-Nam Theun</collectingMunicipality>
NPA,
<location id="8EFC6009FFB5FF8CF26292A7FD19BBAA" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF8CF3AF97A3FDC5B8C9:8EFC6009FFB5FF8CF26292A7FD19BBAA" box="[584,716,222,246]" country="Laos" municipality="Nakai-Nam Theun" name="Ban Navang" pageId="16" pageNumber="325" stateProvince="Khammouan Province">Ban Navang</location>
env., 17°57–59′N, 105°13–16′E,
<quantity id="4CDB9B37FFB5FF8CF19C9284FDE0BA49" box="[438,565,253,277]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="16" pageNumber="325" unit="m" value="675.0" valueMax="750.0" valueMin="600.0">
<elevation id="000ED1E1FFB5FF8CF19C9284FDE0BA49" box="[438,565,253,277]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="16" pageNumber="325" unit="m" value="675.0" valueMax="750.0" valueMin="600.0">600–750 m</elevation>
</quantity>
,
<date id="FF9D1012FFB5FF8CF2699284FCD6BA49" box="[579,771,253,278]" pageId="16" pageNumber="325" value="2012-05-18" valueMax="2012-05-21" valueMin="2012-05-18">
<collectingDate id="EFD9E9FAFFB5FF8CF2699284FCD6BA49" box="[579,771,253,278]" pageId="16" pageNumber="325" value="2012-05-18" valueMax="2012-05-21" valueMin="2012-05-18">18–21 May 2012</collectingDate>
</date>
, NHMB Basel Expedition Laos 2012,
<collectorName id="26D65304FFB5FF8CF2169367FD0CBA69" box="[572,729,285,309]" pageId="16" pageNumber="325">M. Brancucci</collectorName>
,
<collectorName id="26D65304FFB5FF8CF2C29367FF04BA09" pageId="16" pageNumber="325">M. Geiser</collectorName>
,
<collectorName id="26D65304FFB5FF8CF0F09347FE52BA09" box="[218,391,317,341]" pageId="16" pageNumber="325">K. Phanthavong</collectorName>
,
<collectorName id="26D65304FFB5FF8CF1BB9344FDD9BA09" box="[401,524,317,341]" pageId="16" pageNumber="325">S. Xayalath</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF2339344FDBABA09" box="[537,623,317,341]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="325">NHMB</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B4B3C8FFFB5FF8CF25C9344FD9FBA29" ID-GBIF-Occurrence="3760488428" collectionCode="IRSNB" collectorName="de Cooman" country="Laos" location="Tonkin" pageId="16" pageNumber="325" specimenCount="4" stateProvince="Khammouan">
. V
<emphasis id="B957EAC0FFB5FF8CF2B89344FCD5BA09" bold="true" box="[658,768,317,341]" pageId="16" pageNumber="325">IETNAM</emphasis>
:
<specimenCount id="9D25FD5BFFB5FF8CF0A29324FF15BA29" box="[136,192,349,373]" pageId="16" pageNumber="325" type="generic">4 ex.</specimenCount>
,
<location id="8EFC6009FFB5FF8CF0E79324FEC8BA29" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF8CF3AF97A3FDC5B8C9:8EFC6009FFB5FF8CF0E79324FEC8BA29" box="[205,285,349,373]" country="Laos" name="Tonkin" pageId="16" pageNumber="325" stateProvince="Khammouan">Tonkin</location>
, coll.
<collectorName id="26D65304FFB5FF8CF14B9324FE08BA29" box="[353,477,349,373]" pageId="16" pageNumber="325">de Cooman</collectorName>
(
<collectionCode id="ED32AE17FFB5FF8CF1C79324FD94BA29" box="[493,577,349,373]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="16" pageNumber="325">IRSNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFB5FF8CF0A293E4FD12B969" pageId="16" pageNumber="325" type="diagnosis">
<paragraph id="8B9C36D2FFB5FF8CF0A293E4FD12B969" blockId="16.[136,775,413,565]" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF0A293E4FF2EBAE9" bold="true" box="[136,251,413,437]" pageId="16" pageNumber="325">Diagnosis.</emphasis>
Medium sized, elongate oval species (body length
<quantity id="4CDB9B37FFB5FF8CF0A293C4FEC5BA89" box="[136,272,445,469]" metricMagnitude="-3" metricUnit="m" metricValue="3.9499999999999997" metricValueMax="4.1" metricValueMin="3.8" pageId="16" pageNumber="325" unit="mm" value="3.9499999999999997" valueMax="4.1" valueMin="3.8">3.8–4.1 mm</quantity>
). Elytra with subbasal testaceous band broad, more or less connected with elytral base (
<figureCitation id="13182A57FFB5FF8CF27D93A4FD7FBAA9" box="[599,682,477,501]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="16" pageNumber="325">Fig. 16</figureCitation>
). Apical part of median lobe in lateral view with distinct bulge on dorsal side; apex elongate with pointed tip (
<figureCitation id="13182A57FFB5FF8CF2429064FD6FB969" box="[616,698,541,565]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="16" pageNumber="325">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFB5FF8CF0A29024FEEFB9A9" pageId="16" pageNumber="325" type="biology_ecology">
<paragraph id="8B9C36D2FFB5FF8CF0A29024FEEFB9A9" blockId="16.[136,776,605,757]" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF0A29024FF33B929" bold="true" box="[136,230,605,629]" pageId="16" pageNumber="325">Habitat.</emphasis>
In Hainan,
<taxonomicName id="4C234D51FFB5FF8CF1439024FDB9B929" authorityName="Brancucci" authorityYear="1983" box="[361,620,605,629]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="325" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B957EAC0FFB5FF8CF1439024FDB9B929" box="[361,620,605,629]" italics="true" pageId="16" pageNumber="325">Laccophilus tonkinensis</emphasis>
</taxonomicName>
was collected in pools in shaded drying up stream in primary mountane forest. In southern
<collectingCountry id="F3347642FFB5FF8CF17A90E7FE53B9E9" box="[336,390,670,693]" name="Laos" pageId="16" pageNumber="325">Laos</collectingCountry>
, it was collected in a pool in a small forest stream with a clay bottom and a layer of decaying leaves (
<figureCitation id="13182A57FFB5FF8CF0F190A4FEF8B9A9" box="[219,301,733,757]" captionStart="Figs" captionStartId="8.[135,179,2035,2057]" captionTargetBox="[187,1423,189,2020]" captionTargetId="figure-8@8.[187,1423,189,2020]" captionTargetPageId="8" captionText="Figs. 36–39. Habitats: 36, E Kalimantan, Muara Ritan (type locality of Laccophilus kalimantanensis, new species); 37, China, Guangxi, Shiwandashan (locality of L. komareki, new species); 38, E Kalimantan, Sungai Wain (locality of L. mahakamensis); 39, Laos, Attapeu (locality of L. tonkinensis)." figureDoi="http://doi.org/10.5281/zenodo.5385206" httpUri="https://zenodo.org/record/5385206/files/figure.png" pageId="16" pageNumber="325">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFB5FF8CF0A29164FDC5B8C9" pageId="16" pageNumber="325" type="distribution">
<paragraph id="8B9C36D2FFB5FF8CF0A29164FDC5B8C9" blockId="16.[136,775,797,917]" pageId="16" pageNumber="325">
<emphasis id="B957EAC0FFB5FF8CF0A29164FEC3B869" bold="true" box="[136,278,797,821]" pageId="16" pageNumber="325">Distribution.</emphasis>
A species described from northern
<collectingCountry id="F3347642FFB5FF8CF2A89164FD09B869" box="[642,732,797,821]" name="Vietnam" pageId="16" pageNumber="325">Vietnam</collectingCountry>
and subsequently recorded from
<collectingRegion id="49E7F830FFB5FF8CF1849144FE2FB809" box="[430,506,829,853]" country="China" name="Hainan" pageId="16" pageNumber="325">Hainan</collectingRegion>
Island,
<collectingCountry id="F3347642FFB5FF8CF2609144FD5CB809" box="[586,649,829,853]" name="China" pageId="16" pageNumber="325">China</collectingCountry>
(
<bibRefCitation id="EFB24B23FFB5FF8CF2BE9144FF15B829" author="Brancucci M" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="16" pageNumber="325" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
;
<bibRefCitation id="EFB24B23FFB5FF8CF0E79124FE80B829" author="Hajek J" box="[205,341,861,885]" editor="Jach MA &amp; Ji L" firstAuthor="Hajek" journalOrPublisher="Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien." pageId="16" pageNumber="325" pagination="115 - 123" refId="ref15355" refString="Hajek J (2003) Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera). In: Jach MA &amp; Ji L (eds.) Water Beetles of China, Volume III. Zoologisch-Botanische Gesellchaft in Osterreich and Wiener Coleopterologenverein, Wien. Pp. 115 - 123." title="Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera)" type="book chapter" volumeTitle="Water Beetles of China, Volume III" year="2003">Hájek, 2003</bibRefCitation>
). Here we present the first records from central and southern
<collectingCountry id="F3347642FFB5FF8CF1479107FE74B8C9" box="[365,417,894,917]" name="Laos" pageId="16" pageNumber="325">Laos</collectingCountry>
(
<figureCitation id="13182A57FFB5FF8CF19B9104FDD1B8C9" box="[433,516,893,917]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 40–41. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="16" pageNumber="325">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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