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<document id="D68B056E2D0F27D8D92FF583CBD8B1BF" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFA0FF99F0DC94B4FBC7B869" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus elegans Sharp 1882" docType="treatment" docVersion="8" lastPageNumber="314" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="314" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="7DA84ADA42E58E822D87F7D153BC0D9E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2A901646C7D050CFF7855BAE83B22C1D">
<mods:title id="8E038D9AB14622FBD570F8175F6D6B18">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="E5E0EA955E7F6658AF229F291B64801C" type="personal">
<mods:role id="E6AD144D16F8721F106C07A6690BC97D">
<mods:roleTerm id="DD73BAA79DEDE8A46A079CB687717E1F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70E69B9E2EB6754DCE273F61973C34F4">Hájek, Jiří</mods:namePart>
<mods:affiliation id="871DFB7D1E0EB30B80163A79DD2F947A">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 – Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="7B5EA25B0C96862A788246F212ADF9D1" type="email">jiri_hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="FDC0E27C0F25E7AFD25183E39FED8E18" type="personal">
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<mods:roleTerm id="B62376EFC22B4F13FD06DCB9D9959057">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="96820A239FBF456A8C89C6BE70B3F977">Brancucci, Michel</mods:namePart>
<mods:affiliation id="556B4E66658310AFA1D53823E7D88D21">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7BFA68E224E70AC3EA2E2DA337B1640E">text</mods:typeOfResource>
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<mods:title id="896C21BC6BC6450FA66B9F72E263FE76">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="837E6D578B9EF530D4022965B33D8888">
<mods:date id="4F890AB7B073DFC120F6CB87369A96F7">2015</mods:date>
<mods:detail id="65905B62058EE98631A57F3AF9BF87BC" type="pubDate">
<mods:number id="87C18568A034D44D1674154EC935537F">2015-08-06</mods:number>
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<mods:detail id="3D969DAF7D261DF0C8EC07D256F619F0" type="volume">
<mods:number id="59A4C0D4C6213C7EA40053F58440B4CC">63</mods:number>
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<mods:classification id="7F08ECBB7F02F193355FF20564765255">journal article</mods:classification>
<mods:identifier id="DF0D8A7A1ED2FD86981B8A9F11B88F79" type="CLB-Dataset">55750</mods:identifier>
<mods:identifier id="3B8B414C763EAF40F4DB102B4D31D85F" type="DOI">10.5281/zenodo.5385198</mods:identifier>
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<treatment id="038A87C4FFA0FF99F0DC94B4FBC7B869" ID-DOI="http://doi.org/10.5281/zenodo.6512179" ID-GBIF-Taxon="194955237" ID-Zenodo-Dep="6512179" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F0DC94B4FBC7B869" httpUri="http://treatment.plazi.org/id/038A87C4FFA0FF99F0DC94B4FBC7B869" lastPageNumber="314" pageId="5" pageNumber="314">
<subSubSection id="C3396559FFA0FF99F0DC94B4FDB8BDB9" box="[246,621,1741,1765]" pageId="5" pageNumber="314" type="nomenclature">
<paragraph id="8B9C36D2FFA0FF99F0DC94B4FDB8BDB9" blockId="5.[246,621,1741,1797]" box="[246,621,1741,1765]" pageId="5" pageNumber="314">
<heading id="D0D481BEFFA0FF99F0DC94B4FDB8BDB9" bold="true" box="[246,621,1741,1765]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="314" reason="2">
<taxonomicName id="4C234D51FFA0FF99F0DC94B4FDB8BDB9" ID-CoL="6NRMM" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[246,621,1741,1765]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B957EAC0FFA0FF99F0DC94B4FDB8BDB9" bold="true" box="[246,621,1741,1765]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F0DC94B4FE0CBDB9" bold="true" box="[246,473,1741,1765]" italics="true" pageId="5" pageNumber="314">Laccophilus elegans</emphasis>
<bibRefCitation id="EFB24B23FFA0FF99F1C994B4FDB8BDB9" author="Sharp D" box="[483,621,1741,1765]" firstAuthor="Sharp" journalOrPublisher="Scientific Transactions of the Royal Dublin Society" pageId="5" pageNumber="314" pagination="179 - 1003" part="2" refId="ref15851" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003, pls. 7 - 18." title="On aquatic carnivorous Coleoptera or Dytiscidae" type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F1449494FE21BC59" box="[366,500,1773,1797]" pageId="5" pageNumber="314" type="description">
<paragraph id="8B9C36D2FFA0FF99F1449494FE21BC59" blockId="5.[246,621,1741,1797]" box="[366,500,1773,1797]" pageId="5" pageNumber="314">
(
<figureCitation id="13182A57FFA0FF99F15F9494FE16BC59" box="[373,451,1773,1797]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 1–9. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="5" pageNumber="314">Figs. 4</figureCitation>
,
<figureCitation id="13182A57FFA0FF99F1FA9494FE3EBC59" box="[464,491,1773,1797]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="5" pageNumber="314">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F05B9555FEC8BCC4" pageId="5" pageNumber="314" type="reference_group">
<paragraph id="8B9C36D2FFA0FF99F05B9555FEC8BCC4" blockId="5.[113,753,1836,1944]" pageId="5" pageNumber="314">
<treatmentCitationGroup id="AB3311FCFFA0FF99F05B9555FEC8BCC4" pageId="5" pageNumber="314">
<treatmentCitation id="0A8210C3FFA0FF99F05B9555FDD6BC1D" author="Sharp D" box="[113,515,1836,1858]" page="302" pageId="5" pageNumber="314" year="1882">
<taxonomicName id="4C234D51FFA0FF99F05B9555FDD6BC1D" ID-CoL="6NRMM" authority="Sharp, 1882: 302" authorityName="Sharp" authorityPageNumber="302" authorityYear="1882" box="[113,515,1836,1858]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B957EAC0FFA0FF99F05B9555FE91BC1D" box="[113,324,1836,1857]" italics="true" pageId="5" pageNumber="314">Laccophilus elegans</emphasis>
<bibRefCitation id="EFB24B23FFA0FF99F1659555FDD6BC1D" author="Sharp D" box="[335,515,1836,1858]" firstAuthor="Sharp" journalOrPublisher="Scientific Transactions of the Royal Dublin Society" page="302" pageId="5" pageNumber="314" pagination="179 - 1003" part="2" refId="ref15851" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003, pls. 7 - 18." title="On aquatic carnivorous Coleoptera or Dytiscidae" type="journal article" year="1882">Sharp, 1882: 302</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="0A8210C3FFA0FF99F0B29530FEA8BC03" author="Regimbart M" box="[152,381,1865,1887]" page="266" pageId="5" pageNumber="314" year="1899">
<bibRefCitation id="EFB24B23FFA0FF99F0B29530FEA8BC03" author="Regimbart M" box="[152,381,1865,1887]" firstAuthor="Regimbart" journalOrPublisher="Annales de la Societe Entomologique de France" page="266" pageId="5" pageNumber="314" pagination="186 - 367" part="68" refId="ref15792" refString="Regimbart M (1899) Revision des Dytiscidae de la region Indo- Sino-Malaise. Annales de la Societe Entomologique de France, 68: 186 - 367." title="Revision des Dytiscidae de la region Indo- Sino-Malaise" type="journal article" year="1899">Régimbart, 1899: 266</bibRefCitation>
</treatmentCitation>
(description);
<treatmentCitation id="0A8210C3FFA0FF99F2349530FD25BC02" author="Vazirani TG" box="[542,752,1865,1887]" page="241" pageId="5" pageNumber="314" year="1969">
<bibRefCitation id="EFB24B23FFA0FF99F2349530FD25BC02" author="Vazirani TG" box="[542,752,1865,1887]" firstAuthor="Vazirani" journalOrPublisher="Oriental Insects" page="241" pageId="5" pageNumber="314" pagination="221 - 341" part="2" refId="ref15885" refString="Vazirani TG (1969) Contribution to the study of aquatic beetles (Coleoptera). 2. A Review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India. Oriental Insects, 2 (1968): 221 - 341." title="Contribution to the study of aquatic beetles (Coleoptera). 2. A Review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India" type="journal article" year="1969">Vazirani, 1969: 241</bibRefCitation>
</treatmentCitation>
(description, new records);
<treatmentCitation id="0A8210C3FFA0FF99F188951CFDA4BC27" author="Brancucci M" box="[418,625,1893,1915]" page="257" pageId="5" pageNumber="314" year="1983">
<bibRefCitation id="EFB24B23FFA0FF99F188951CFDA4BC27" author="Brancucci M" box="[418,625,1893,1915]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" page="257" pageId="5" pageNumber="314" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983: 257</bibRefCitation>
</treatmentCitation>
(description, new records).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F05B95B8FC26BAE9" pageId="5" pageNumber="314" type="materials_examined">
<paragraph id="8B9C36D2FFA0FF99F05B95B8FE0CBC84" blockId="5.[113,473,1984,2009]" box="[113,473,1984,2009]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F05B95B8FEDBBC84" bold="true" box="[113,270,1984,2009]" pageId="5" pageNumber="314">
<typeStatus id="54988870FFA0FF99F05B95B8FF7EBC85" box="[113,171,1985,2009]" pageId="5" pageNumber="314">Type</typeStatus>
locality.
</emphasis>
<location id="8EFC6009FFA0FF99F13C95B9FE00BC84" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F0DC94B4FBC7B869:8EFC6009FFA0FF99F13C95B9FE00BC84" box="[278,469,1984,2008]" name="Andaman Islands" pageId="5" pageNumber="314">Andaman Islands</location>
.
</paragraph>
<paragraph id="8B9C36D2FFA0FF99F05B9A79FC26BAE9" blockId="5.[113,752,2048,2105]" lastBlockId="5.[812,1452,189,437]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F05B9A79FE83B344" bold="true" box="[113,342,2048,2072]" pageId="5" pageNumber="314">Material examined.</emphasis>
<materialsCitation id="3B4B3C8FFFA0FF99F14B9A78FD3CB345" ID-GBIF-Occurrence="3760488366" box="[353,745,2048,2073]" country="India" location="Karnataka" pageId="5" pageNumber="314" specimenCount="10" stateProvince="Karnataka" typeStatus="holotype">
<specimenCount id="9D25FD5BFFA0FF99F14B9A78FE2FB344" box="[353,506,2048,2073]" pageId="5" pageNumber="314" type="generic">10 specimens</specimenCount>
.
<emphasis id="B957EAC0FFA0FF99F2229A79FD3CB345" bold="true" box="[520,745,2048,2073]" pageId="5" pageNumber="314">
<collectingCountry id="F3347642FFA0FF99F2229A79FD8EB344" box="[520,603,2048,2072]" name="India" pageId="5" pageNumber="314">INDIA</collectingCountry>
:
<collectingRegion id="49E7F830FFA0FF99F2419A78FD3CB345" box="[619,745,2049,2073]" country="India" name="Karnataka" pageId="5" pageNumber="314">Karnataka</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B4B3C8FFFA0FF99F05B9A58FC1DBBA9" ID-GBIF-Occurrence="3760488326" collectingDate="2002-05-16" collectingDateMax="2002-05-21" collectingDateMin="2002-05-16" collectorName="P. Pacholatko." country="India" elevation="350" latitude="12.47" location="Gundia" longLatPrecision="776" longitude="75.43" pageId="5" pageNumber="314" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka" typeStatus="holotype">
<specimenCount id="9D25FD5BFFA0FF99F05B9A58FF03B364" box="[113,214,2080,2105]" pageId="5" pageNumber="314" type="female">1 female</specimenCount>
,
<locationDeviation id="9A2E2FF0FFA0FF99F0CF9A58FE70B364" box="[229,421,2080,2105]" location="Shiradi" pageId="5" pageNumber="314">
<quantity id="4CDB9B37FFA0FF99F0CF9A58FEF8B364" box="[229,301,2080,2105]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="314" unit="km" value="18.0">18 km</quantity>
E Shiradi
</locationDeviation>
,
<location id="8EFC6009FFA0FF99F1999A59FDD3B364" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F0DC94B4FBC7B869:8EFC6009FFA0FF99F1999A59FDD3B364" box="[435,518,2080,2104]" country="India" latitude="12.47" longLatPrecision="776" longitude="75.43" name="Gundia" pageId="5" pageNumber="314" stateProvince="Karnataka">Gundia</location>
,
<geoCoordinate id="EE175015FFA0FF99F23F9A58FDB3B365" box="[533,614,2081,2105]" degrees="12.47" direction="north" orientation="latitude" pageId="5" pageNumber="314" precision="555" value="12.47">12.47N</geoCoordinate>
<geoCoordinate id="EE175015FFA0FF99F25B9A59FD14B364" box="[625,705,2080,2104]" degrees="75.43" direction="east" orientation="longitude" pageId="5" pageNumber="314" precision="555" value="75.43">75.43E</geoCoordinate>
, W. [Western]
<location id="8EFC6009FFA0FF99F3B492C4FC09BB89" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F0DC94B4FBC7B869:8EFC6009FFA0FF99F3B492C4FC09BB89" box="[926,988,189,213]" name="Ghats" pageId="5" pageNumber="314">Ghats</location>
[Mts.],
<quantity id="4CDB9B37FFA0FF99F41892C4FB78BB89" box="[1074,1197,189,213]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="5" pageNumber="314" unit="m" value="350.0" valueMax="500.0" valueMin="200.0">
<elevation id="000ED1E1FFA0FF99F41892C4FB78BB89" box="[1074,1197,189,213]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="5" pageNumber="314" unit="m" value="350.0" valueMax="500.0" valueMin="200.0">200–500 m</elevation>
</quantity>
,
<date id="FF9D1012FFA0FF99F49292C4FAA7BB89" box="[1208,1394,189,214]" pageId="5" pageNumber="314" value="2002-05-16" valueMax="2002-05-21" valueMin="2002-05-16">
<collectingDate id="EFD9E9FAFFA0FF99F49292C4FAA7BB89" box="[1208,1394,189,214]" pageId="5" pageNumber="314" value="2002-05-16" valueMax="2002-05-21" valueMin="2002-05-16">16–21 May 2002</collectingDate>
</date>
; coll.
<collectorName id="26D65304FFA0FF99F30792A7FC1DBBA9" box="[813,968,221,245]" pageId="5" pageNumber="314">P. Pacholátko.</collectorName>
</materialsCitation>
<materialsCitation id="3B4B3C8FFFA0FF99F3E492A4FC53BA69" ID-GBIF-Occurrence="3760488357" collectingDate="1993-12-06" collectingDateMax="1999-01-09" collectingDateMin="1993-12-06" collectionCode="NMPC, PMCB, NHMW" collectorName="P. Mazzoldi" county="Road Moolamattam - Idukki" elevation="650" latitude="10.05" location="Kallar valley" longLatPrecision="1298" longitude="76.96667" municipality="Panavu" pageId="5" pageNumber="314" specimenCount="9" specimenCount-female="3" stateProvince="Kerala">
<emphasis id="B957EAC0FFA0FF99F3E492A4FBC9BBA9" bold="true" box="[974,1052,221,245]" pageId="5" pageNumber="314">
<collectingRegion id="49E7F830FFA0FF99F3E492A4FBC9BBA9" box="[974,1052,221,245]" country="India" name="Kerala" pageId="5" pageNumber="314">Kerala</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFA0FF99F40192A4FBB7BBA9" box="[1067,1122,221,245]" pageId="5" pageNumber="314" type="generic">6 ex.</specimenCount>
,
<collectingCounty id="62FD4E5EFFA0FF99F44492A4FA72BBA9" box="[1134,1447,221,245]" pageId="5" pageNumber="314">Road Moolamattam – Idukki</collectingCounty>
, near
<collectingMunicipality id="6BF8ACA8FFA0FF99F34B9287FC65BA49" box="[865,944,254,277]" pageId="5" pageNumber="314">Panavu</collectingMunicipality>
stream,
<quantity id="4CDB9B37FFA0FF99F4269284FB52BA49" box="[1036,1159,253,277]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="314" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
<elevation id="000ED1E1FFA0FF99F4269284FB52BA49" box="[1036,1159,253,277]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="314" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600–700 m</elevation>
</quantity>
, coll.
<collectorName id="26D65304FFA0FF99F4E29287FA9FBA49" box="[1224,1354,253,277]" pageId="5" pageNumber="314">P. Mazzoldi</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F5729284FA7DBA49" box="[1368,1448,253,277]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="314">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFA0FF99F3079364FCAEBA69" box="[813,891,285,309]" pageId="5" pageNumber="314">PMCB</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4B3C8FFFA0FF99F3A19364FBDDBA29" ID-GBIF-Occurrence="3760488387" collectingDate="1993-12-06" collectingDateMax="1999-01-09" collectingDateMin="1993-12-06" collectionCode="NMPC, PMCB, NHMW" collectorName="D. Boukal &amp; Z. Kejval" county="Road Moolamattam - Idukki" elevation="650" latitude="10.05" location="Kallar valley" longLatPrecision="1298" longitude="76.96667" municipality="Panavu" pageId="5" pageNumber="314" specimenCount="9" specimenCount-female="3" stateProvince="Kerala">
<specimenCount id="9D25FD5BFFA0FF99F3A19364FC24BA69" box="[907,1009,285,309]" pageId="5" pageNumber="314" type="female">2 females</specimenCount>
,
<locationDeviation id="9A2E2FF0FFA0FF99F3D09364FB16BA69" box="[1018,1219,285,309]" location="Munnar" pageId="5" pageNumber="314">
<quantity id="4CDB9B37FFA0FF99F3D09364FBEEBA69" box="[1018,1083,285,309]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="314" unit="km" value="15.0">15 km</quantity>
SW Munnar
</locationDeviation>
,
<location id="8EFC6009FFA0FF99F4E69364FA80BA69" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F0DC94B4FBC7B869:8EFC6009FFA0FF99F4E69364FA80BA69" box="[1228,1365,285,309]" county="Road Moolamattam - Idukki" latitude="10.05" longLatPrecision="1298" longitude="76.96667" municipality="Panavu" name="Kallar valley" pageId="5" pageNumber="314" stateProvince="Kerala">Kallar valley</location>
,
<geoCoordinate id="EE175015FFA0FF99F5749364FA79BA69" box="[1374,1452,285,309]" degrees="10.02" direction="north" orientation="latitude" pageId="5" pageNumber="314" precision="555" value="10.02">10.02N</geoCoordinate>
<geoCoordinate id="EE175015FFA0FF99F3069344FCA9BA09" box="[812,892,317,341]" degrees="76.58" direction="east" orientation="longitude" pageId="5" pageNumber="314" precision="555" value="76.58">76.58E</geoCoordinate>
,
<quantity id="4CDB9B37FFA0FF99F3A29344FC0BBA09" box="[904,990,317,341]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="314" unit="m" value="1000.0">
<elevation id="000ED1E1FFA0FF99F3A29344FC0BBA09" box="[904,990,317,341]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="314" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FF9D1012FFA0FF99F3C19344FB0DBA09" box="[1003,1240,317,341]" pageId="5" pageNumber="314" value="1993-12-06" valueMax="1993-12-18" valueMin="1993-12-06">
<collectingDate id="EFD9E9FAFFA0FF99F3C19344FB0DBA09" box="[1003,1240,317,341]" pageId="5" pageNumber="314" value="1993-12-06" valueMax="1993-12-18" valueMin="1993-12-06">6–18 December 1993</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA0FF99F5379347FA58BA09" box="[1309,1421,317,341]" pageId="5" pageNumber="314">D. Boukal</collectorName>
&amp;
<collectorName id="26D65304FFA0FF99F3079327FC44BA29" box="[813,913,349,373]" pageId="5" pageNumber="314">Z. Kejval</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F3B59324FC28BA29" box="[927,1021,349,373]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="314" type="Museum">NHMW</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4B3C8FFFA0FF99F4249324FC3BBAE9" ID-GBIF-Occurrence="3760488379" collectingDate="1993-12-06" collectingDateMax="1999-01-09" collectingDateMin="1993-12-06" collectionCode="NMPC, PMCB, NHMW" collectorName="D. Boukal" county="Road Moolamattam - Idukki" elevation="650" latitude="10.05" location="Kallar valley" longLatPrecision="1298" longitude="76.96667" municipality="Panavu" pageId="5" pageNumber="314" specimenCount="9" specimenCount-female="3" stateProvince="Kerala">
<specimenCount id="9D25FD5BFFA0FF99F4249324FBB9BA29" box="[1038,1132,349,373]" pageId="5" pageNumber="314" type="female">1 female</specimenCount>
,
<locationDeviation id="9A2E2FF0FFA0FF99F45D9324FA8BBA29" box="[1143,1374,349,373]" location="Munnar" pageId="5" pageNumber="314">
<quantity id="4CDB9B37FFA0FF99F45D9324FB6FBA29" box="[1143,1210,349,373]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="314" unit="km" value="10.0">10 km</quantity>
WSW Munnar
</locationDeviation>
,
<location id="8EFC6009FFA0FF99F5439324FCA4BAC9" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F0DC94B4FBC7B869:8EFC6009FFA0FF99F5439324FCA4BAC9" county="Road Moolamattam - Idukki" latitude="10.05" longLatPrecision="1298" longitude="76.96667" municipality="Panavu" name="Kallar valley" pageId="5" pageNumber="314" stateProvince="Kerala">Kallar valley</location>
,
<quantity id="4CDB9B37FFA0FF99F3579304FC07BAC9" box="[893,978,381,405]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="314" unit="m" value="1100.0">
<elevation id="000ED1E1FFA0FF99F3579304FC07BAC9" box="[893,978,381,405]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="314" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<geoCoordinate id="EE175015FFA0FF99F3F49305FBEDBAC9" box="[990,1080,380,405]" degrees="10" direction="north" minutes="03" orientation="latitude" pageId="5" pageNumber="314" precision="925" value="10.05">10°03′N</geoCoordinate>
<geoCoordinate id="EE175015FFA0FF99F46A9305FB4CBAC9" box="[1088,1177,380,405]" degrees="76" direction="east" minutes="58" orientation="longitude" pageId="5" pageNumber="314" precision="925" value="76.96667">76°58′E</geoCoordinate>
,
<date id="FF9D1012FFA0FF99F48F9304FA9BBAC9" box="[1189,1358,381,406]" pageId="5" pageNumber="314" value="1999-01-09">
<collectingDate id="EFD9E9FAFFA0FF99F48F9304FA9BBAC9" box="[1189,1358,381,406]" pageId="5" pageNumber="314" value="1999-01-09">9 January 1999</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA0FF99F5BB9307FCAFBAE9" pageId="5" pageNumber="314">D. Boukal</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F3A093E4FC3DBAE9" box="[906,1000,413,437]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="314" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F30793A4FB02B929" pageId="5" pageNumber="314" type="diagnosis">
<paragraph id="8B9C36D2FFA0FF99F30793A4FB02B929" blockId="5.[812,1452,477,629]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F30793A4FC75BAA9" bold="true" box="[813,928,477,501]" pageId="5" pageNumber="314">Diagnosis.</emphasis>
Medium sized, elongate oval species (body length 3.8–4.0 mm). Elytra with subbasal testaceous band broad, more or less connected with elytral base (
<figureCitation id="13182A57FFA0FF99F52C9064FA99B969" box="[1286,1356,541,565]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 1–9. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="5" pageNumber="314">Fig. 4</figureCitation>
). Apical part of median lobe in lateral view with distinct bulge on dorsal side; apex short, blunt (
<figureCitation id="13182A57FFA0FF99F4539024FB1FB929" box="[1145,1226,605,629]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="5" pageNumber="314">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F30790E4FBC7B869" pageId="5" pageNumber="314" type="distribution">
<paragraph id="8B9C36D2FFA0FF99F30790E4FBC7B869" blockId="5.[813,1452,669,821]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F30790E4FC16B9E9" bold="true" box="[813,963,669,693]" pageId="5" pageNumber="314">Distribution.</emphasis>
Widespread in southern and eastern
<collectingCountry id="F3347642FFA0FF99F54690E4FA72B9E9" box="[1388,1447,669,693]" name="India" pageId="5" pageNumber="314">India</collectingCountry>
.
<bibRefCitation id="EFB24B23FFA0FF99F30790C4FC0DB989" author="Vazirani TG" box="[813,984,701,725]" firstAuthor="Vazirani" journalOrPublisher="Oriental Insects" pageId="5" pageNumber="314" pagination="221 - 341" part="2" refId="ref15885" refString="Vazirani TG (1969) Contribution to the study of aquatic beetles (Coleoptera). 2. A Review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India. Oriental Insects, 2 (1968): 221 - 341." title="Contribution to the study of aquatic beetles (Coleoptera). 2. A Review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India" type="journal article" year="1969">Vazirani (1969)</bibRefCitation>
and
<bibRefCitation id="EFB24B23FFA0FF99F42590C4FB1EB989" author="Brancucci M" box="[1039,1227,701,725]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="5" pageNumber="314" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci (1983)</bibRefCitation>
published data from
<collectingRegion id="49E7F830FFA0FF99F30790A4FBA2B9A9" box="[813,1143,733,757]" country="India" name="Andaman and Nicobar Islands" pageId="5" pageNumber="314">Andaman and Nicobar Islands</collectingRegion>
,
<collectingRegion id="49E7F830FFA0FF99F4A990A4FAE7B9A9" box="[1155,1330,733,757]" country="India" name="Andhra Pradesh" pageId="5" pageNumber="314">Andhra Pradesh</collectingRegion>
,
<collectingRegion id="49E7F830FFA0FF99F51590A4FA7DB9A9" box="[1343,1448,733,757]" country="India" name="Nagaland" pageId="5" pageNumber="314">Nagaland</collectingRegion>
,
<collectingRegion id="49E7F830FFA0FF99F3079084FCA4B849" box="[813,881,765,789]" country="India" name="Odisha" pageId="5" pageNumber="314">Orissa</collectingRegion>
and
<collectingRegion id="49E7F830FFA0FF99F3819087FBEEB849" box="[939,1083,765,789]" country="India" name="West Bengal" pageId="5" pageNumber="314">West Bengal</collectingRegion>
. We add records from
<collectingRegion id="49E7F830FFA0FF99F5149084FA79B849" box="[1342,1452,765,789]" country="India" name="Karnataka" pageId="5" pageNumber="314">Karnataka</collectingRegion>
and
<collectingRegion id="49E7F830FFA0FF99F3769164FC76B869" box="[860,931,797,821]" country="India" name="Kerala" pageId="5" pageNumber="314">Kerala</collectingRegion>
(
<figureCitation id="13182A57FFA0FF99F3999164FBD0B869" box="[947,1029,797,821]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 40–41. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="5" pageNumber="314">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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