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<mods:title id="858507F058084DCFCE4DB1232D82145A">The Dromiusina Bonelli, 1810 of southwestern Saudi Arabia with description of a new species (Coleoptera, Carabidae, Lebiini)</mods:title>
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<mods:namePart id="B40C18B879A4AB341A8090D5813EBB9B">Rasool, Iftekhar</mods:namePart>
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<mods:namePart id="387913D783BC88F69101B10B02E3C61E">Abdel-Dayem, Mahmoud S.</mods:namePart>
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<mods:namePart id="E9CABD3E057E6CB300725961E87E5D4B">Felix, Ron F. F. L.</mods:namePart>
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<mods:namePart id="CEDC1B772ADDB0B6B5234F3A11C97474">Aldhafer, Hathal M.</mods:namePart>
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<treatment id="034346A9E20733E24A5950381D43DB59" ID-GBIF-Taxon="146053649" LSID="urn:lsid:plazi:treatment:034346A9E20733E24A5950381D43DB59" httpUri="http://treatment.plazi.org/id/034346A9E20733E24A5950381D43DB59" lastPageNumber="73" pageId="0" pageNumber="73">
<subSubSection id="11983D233E8B025E8A512D3E76669A19" pageId="0" pageNumber="73" type="nomenclature">
<paragraph id="5AD646206F9788D81EB83B865FF4B232" pageId="0" pageNumber="73">
<taxonomicName id="AF8D3ACEB603BB25799C6AF03B69CAAA" ID-CoL="PY8L" authority="Bedel, 1907" authorityName="Bedel" authorityYear="1907" class="Insecta" family="Carabidae" genus="Calodromius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calodromius mayeti" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="mayeti">Calodromius mayeti (Bedel, 1907)</taxonomicName>
Figures 11, 22, 37, 49
</paragraph>
</subSubSection>
<subSubSection id="901104FDAA54C61A3C1FE026CDC44C02" pageId="0" pageNumber="73" type="reference_group">
<paragraph id="384802311EF3347108156C618D883597" pageId="0" pageNumber="73">
<taxonomicName id="553A00687E8E4E10FB18D8C72C02111A" class="Insecta" family="Carabidae" genus="Dromius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dromius mayeti" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="mayeti">Dromius mayeti</taxonomicName>
Bedel, 1907: 272.
</paragraph>
</subSubSection>
<subSubSection id="861D427FDC3A598F1730A53134158F80" pageId="0" pageNumber="73" type="type locality">
<paragraph id="7542A94277A586C3597345FE81591A09" pageId="0" pageNumber="73">Type locality.</paragraph>
<paragraph id="5C27F1E00F054C5D7EFDBA50471088F9" pageId="0" pageNumber="73">Tunisia.</paragraph>
</subSubSection>
<subSubSection id="E558CC40F1B61FE0FD753AFB777A00B2" pageId="0" pageNumber="73" type="type depository">
<paragraph id="CC835CDBEA4DA38D4825A7AEDB1585EE" pageId="0" pageNumber="73">Type depository.</paragraph>
<paragraph id="A853910681839FE1649A98C8C871BBF8" pageId="0" pageNumber="73">Holotype male in MNHN: Paratype in NHMB</paragraph>
</subSubSection>
<subSubSection id="200E4E992532AACA1F88D518BC8DE10A" pageId="0" pageNumber="73" type="materials_examined">
<paragraph id="05F81FFB3C502D697AE9EC77FCB1D9CB" pageId="0" pageNumber="73">Material examined.</paragraph>
<paragraph id="0F793D9570ECB98F2E69F850AAD15CF4" pageId="0" pageNumber="73">
Total 21 specimens: 1♀ &quot;[yellow label]&quot; / &quot;Saudi Arabia, W. Buttiker&quot; / Butayn, 21.IV.1981&quot;/ &quot;
<taxonomicName id="240915A8B935DFCC5314C302F5739017" class="Insecta" family="Carabidae" genus="Philorhizus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philorhizus mayeti" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="mayeti">Philorhizus mayeti</taxonomicName>
, J. Mateu det. 1983&quot;. [NHMB]. Al Baha: 1♀, &quot;KSA, Al Makhwa, Shada Al Aala, 19°52.598'N 41°18.672'E Alt. 892 m, 26.I.2015, (HP on light), I. Rasool&quot;. 2♂, &quot;16.II.2014, (LT), M.S. Abdel-Dayem &amp; I. Rasool&quot;. 1♂, 2♀, &quot;19°51.066'N 41°18.037'E Alt. 1325 m, 02.III.2015, (LT)., 1♀, &quot;19°51.066'N 41°18.037'E Alt. 1325 m, 17.X.2014, (LT)., 1♀, &quot;19°50.710'N 41°18.267'E Alt. 1474 m, (LT), H. Al Dhafer, M.S. Abdel-Dayem, H. H. Fadl &amp; I. Rasool&quot;. 1♂, 2♀, &quot;19°52.717'N 41°18.712'E Alt. 825 m, 15.XI.2015, (LT)., 1♂, 1♀, &quot;13.XI.2015, (LT)., 2♀, &quot;19°52.598'N 41°18.672'E Alt. 892 m, 13.XI.2015, (LT)., 1♂, 1♀, &quot;19°51.762'N 41°18.089'E Alt. 1225, 12.XI.2015, (LT)., 1♀, &quot;19°52.685'N 41°18.663'E Alt. 851 m, 15.XI.2015, (LT)., 1♂, &quot;19°51.066'N 41°18.037'E Alt. 1325 m, 14.XI.2015, (LT), H. Al Dhafer, M.S. Abdel-Dayem, H. H. Fadl,A. Elgarbawy, El Turkey and Soliman, A.&quot; Asir: 1♂, &quot;Asir, Abha, Rayda, 18°12.315'N 42°24.607'E Alt. 2578 m, 18.XI.2015, (LT), H. Al Dhafer, M.S. Abdel-Dayem, H. H. Fadl, A. Elgarbawy, El Turkey and Soliman&quot; [KSMA].
</paragraph>
</subSubSection>
<subSubSection id="D8464E21416447C1AB87C2BC02692032" pageId="0" pageNumber="73" type="description">
<paragraph id="85D69F4C60E3DFFA4E077CC490EDF733" pageId="0" pageNumber="73">Description.</paragraph>
<paragraph id="C453D9343D4FE249AE904B6221907600" pageId="0" pageNumber="73">Body form (Fig. 22), small species 3.60-3.90 mm. Color: Dorsum and ventrum of pronotum and abdomen, epipleurae, antennae, mouthparts and legs testaceous; head slightly darker than others; elytra pale testaceous with transverse black band in the middle, not reaching to the lateral margins, black band prolonged to the base along the suture, shortly extended towards the apex. Microsculpture: head, labrum, pronotum and elytra with isodiametric mesh pattern; sternite with transverse microlines. Head: almost as long as wide, HL 0.72-0.82 mm and HW 0.70-0.79 mm (Fig. 11). Pronotum: transverse, wider than long, PW 0.71-0.78 mm, and PL 0.56-0.64; sinuate posteriorly, base almost straight with acute angles (Fig. 11). Elytra: broadened posteriorly, EL 1.87-2.06 mm and EW 1.42-1.49 mm; apical margins transversely truncate; striae and intervals finely punctuate, provided with short brown pubescence. Claws smooth. Abdomen: All visible sternite with short pubescence, apical margin of last sternum rounded and 4-setose in both sexes. Aedeagus: Small and thick aedeagus (Fig. 37), 0.78 mm; basal side of aedeagus narrowed; very broad and depressed at apical lamina; apical end short and with tooth like tip.</paragraph>
<caption id="A9ACB57889B6A9760867DA104EF8D58E" pageId="0" pageNumber="73">
<paragraph id="57B90152DCCD0F941956ADFF060F7B16" pageId="0" pageNumber="73">
Figures 11-21. Dorsal view of head and pronotum of
<taxonomicName id="0545BC8AF6152D36AB7AA8A1560F81F9" lsidName="Dromiusina" pageId="0" pageNumber="73" rank="subTribe" subTribe="Dromiusina">Dromiusina</taxonomicName>
species: 11
<taxonomicName id="78662B789D1DA9A3A310C460C9FA83A0" class="Insecta" family="Carabidae" genus="Calodromius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calodromius mayeti" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="mayeti">Calodromius mayeti</taxonomicName>
(Bedel, 1907) 12
<taxonomicName id="83D17B348A11130C73089D5CF113509F" class="Insecta" family="Carabidae" genus="Dromius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dromius saudiarabicus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="saudiarabicus">Dromius saudiarabicus</taxonomicName>
sp. n. 13
<taxonomicName id="2B3115EB4D64243EE3DACED5BCCD6609" lsidName="D. buettikeri" pageId="0" pageNumber="73" rank="species" species="buettikeri">D. buettikeri</taxonomicName>
Mateu, 1990 14
<taxonomicName id="A64675184BDEE3960D565FD2BABDE64B" class="Insecta" family="Carabidae" genus="Metadromius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metadromius arabicus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="arabicus">Metadromius arabicus</taxonomicName>
Mateu, 1979 15
<taxonomicName id="08D5ED3A213BA2B173A39082CE8F2C23" lsidName="M. brittoni" pageId="0" pageNumber="73" rank="species" species="brittoni">M. brittoni</taxonomicName>
(Basilewsky, 1948) 16
<taxonomicName id="2F379A7C818299D68DF37C8369DA1EEE" class="Insecta" family="Carabidae" genus="Microlestes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microlestes discoidalis" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="discoidalis">Microlestes discoidalis</taxonomicName>
(Fairmaire, 1892) 17
<taxonomicName id="351A5E2EA2B85C5777E9A26374DDFC8B" lsidName="M. glabrellus" pageId="0" pageNumber="73" rank="species" species="glabrellus">M. glabrellus</taxonomicName>
(Reitter, 1901) 18
<taxonomicName id="A95BD7F691F9C0F9609D9B72AB2193B5" lsidName="M. infuscatus" pageId="0" pageNumber="73" rank="subSpecies" species="infuscatus" subSpecies="fragilis">M. infuscatus fragilis</taxonomicName>
Mateu, 1956 19
<taxonomicName id="0F2F2DB128321418D9CACD6E33D72862" class="Insecta" family="Carabidae" genus="Pseudomesolestes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudomesolestes brittoni" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="brittoni">Pseudomesolestes brittoni</taxonomicName>
Mateu, 1956 20
<taxonomicName id="DEA56932BBD6DC9D0C68188F5755E7AA" lsidName="P. quadriguttatus" pageId="0" pageNumber="73" rank="species" species="quadriguttatus">P. quadriguttatus</taxonomicName>
Mateu, 1979 21
<taxonomicName id="7731D759BAF5437A1B470D4E4C651E22" class="Insecta" family="Carabidae" genus="Zolotarevskyella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zolotarevskyella rhytidera" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="rhytidera">Zolotarevskyella rhytidera</taxonomicName>
(Chaudoir, 1876).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D6FEB9F03F1E1566148B2A557D6B4F01" pageId="0" pageNumber="73" type="ecological notes">
<paragraph id="5E79F21AEABB4C0BE324B40E150F94D6" pageId="0" pageNumber="73">Ecological notes.</paragraph>
<paragraph id="9ADAA13B4537243D0DE39B4D72149C33" pageId="0" pageNumber="73">
This species was collected in the natural habitat of mountains and valleys covered with variety of vegetation, sand, and stones. Species was distributed in elevation ranging from 892-1611 m (Fig. 49). Adult beetles were attracted to
<normalizedToken id="3C4D139DF25524A2D1021D03D0E46E18" originalValue="UVlight">UV-light</normalizedToken>
. In winters this species appears in low elevation while in summers it appears in high elevation.
</paragraph>
</subSubSection>
<subSubSection id="83C2E39899DF3C1D00454AAC5D1BA90D" pageId="0" pageNumber="73" type="distribution">
<paragraph id="B33BAEA2BB2CD60F87DB786D1CC02113" pageId="0" pageNumber="73">Geographical distribution.</paragraph>
<paragraph id="512BB2EB6C830735A9044885DAC5A95E" pageId="0" pageNumber="73">
This species is recorded from Iran, Libya, Morocco, Saudi Arabia, Tunisia, UAE (
<bibRefCitation id="F780EE860D0AB88D40DA4790E6BF473E" pageId="0" pageNumber="73">Kabak 2003</bibRefCitation>
,
<bibRefCitation id="4829D0C6CC706BB54D6E3F9D3F499532" pageId="0" pageNumber="73">2017</bibRefCitation>
,
<bibRefCitation id="2DD61062F2C4F44EF54427EB0ABA5CFE" author="Felix, RFFL" journalOrPublisher="Arthropod fauna of the UAE" pageId="0" pageNumber="73" pagination="66 - 141" title="Order Coleoptera, family Carabidae." volume="2" year="2009">Felix 2009</bibRefCitation>
). In current study it is collected from Al Baha and Asir regions of Saudi Arabia. It is a Mediterranean species that exemplifies Mediterranean-Sindian chorotype.
</paragraph>
</subSubSection>
</treatment>
</document>