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<document id="8667E5218E82C24066BE9D840EA2DF91" ID-DOI="10.5281/zenodo.5183220" ID-ISSN="0253-116X" ID-Zenodo-Dep="5183220" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628561451650" checkinUser="felipe" docAuthor="Guilbert, Eric &amp; Heiss, Ernst" docDate="2016" docId="03F587E7FFBF2D72F69CFCE1FEE5FAA2" docLanguage="en" docName="LinzerbiolBeitr.48.2.1081-1090.pdf" docOrigin="Linzer biologische Beiträge 48 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Hovatlas invaginatus Rodrigues 1992" docType="treatment" docVersion="4" lastPageNumber="1085" masterDocId="FFCCFF9FFFBB2D76F62EFFA7FFCBFF86" masterDocTitle="First fossil records of Tingidae from Madagascan Copal with description of two new species (Hemiptera, Heteroptera)" masterLastPageNumber="1090" masterPageNumber="1081" pageNumber="1085" updateTime="1699046987057" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B27098DFFA54668C321CD7EFBE7D8805">First fossil records of Tingidae from Madagascan Copal with description of two new species (Hemiptera, Heteroptera)</mods:title>
</mods:titleInfo>
<mods:name id="AB537C910D512B2B4C9472DF442B6491" type="personal">
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<mods:namePart id="E8853142C6BB7DBD76E4FA9603B6043C">Guilbert, Eric</mods:namePart>
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<mods:name id="76A68F07E2992B73CD39662D510ACBB1" type="personal">
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<mods:roleTerm id="186168EDC0129813133BFA15F26A0597">Author</mods:roleTerm>
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<mods:namePart id="A77C0DB614AD34A137EED161F8007DE8">Heiss, Ernst</mods:namePart>
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<mods:title id="CFD9927BBD04F22B2FBE7612E087F123">Linzer biologische Beiträge</mods:title>
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<mods:date id="A1AD07D7B80DCEF33EB33EA6591B3A3C">2016</mods:date>
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<mods:number id="D537AD76622C5E66E0863A3C1CED043E">2016-12-19</mods:number>
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<mods:detail id="93649DFDED66355E02C4644AE15FC9A9" type="volume">
<mods:number id="29AD8C494EDACC588A366EF83B48F1B6">48</mods:number>
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<mods:number id="9796D6AD737AABCA780DEDEBC6445432">2</mods:number>
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<mods:classification id="A9A079CDE585F18EF0991F3F5305E213">journal article</mods:classification>
<mods:identifier id="F4154FBBD913D615A70A38B2B128512B" type="DOI">10.5281/zenodo.5183220</mods:identifier>
<mods:identifier id="5AA83C50AD75EC01B14EF051628AFFE2" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="C54E7237E548A253DB28F19B1ECA84BC" type="Zenodo-Dep">5183220</mods:identifier>
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<treatment id="03F587E7FFBF2D72F69CFCE1FEE5FAA2" ID-DOI="http://doi.org/10.5281/zenodo.5192096" ID-Zenodo-Dep="5192096" LSID="urn:lsid:plazi:treatment:03F587E7FFBF2D72F69CFCE1FEE5FAA2" httpUri="http://treatment.plazi.org/id/03F587E7FFBF2D72F69CFCE1FEE5FAA2" lastPageNumber="1085" pageId="4" pageNumber="1085">
<subSubSection id="C346657AFFBF2D72F69CFCE1FD94FC1C" pageId="4" pageNumber="1085" type="nomenclature">
<paragraph id="8BE336F1FFBF2D72F69CFCE1FCE2FCDA" blockId="4.[178,809,838,860]" box="[178,809,838,860]" pageId="4" pageNumber="1085">
<heading id="D0AB819DFFBF2D72F69CFCE1FCE2FCDA" box="[178,809,838,860]" fontSize="8" level="2" pageId="4" pageNumber="1085" reason="2">
<emphasis id="B928EAE3FFBF2D72F69CFCE1FCE2FCDA" bold="true" box="[178,809,838,860]" pageId="4" pageNumber="1085">
<taxonomicName id="4C5C4D72FFBF2D72F69CFCE1FE44FCDD" ID-CoL="3MR99" authorityName="Rodrigues" authorityYear="1992" box="[178,399,838,860]" class="Insecta" family="Tingidae" genus="Hovatlas" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="1085" phylum="Arthropoda" rank="species" species="invaginatus">
<emphasis id="B928EAE3FFBF2D72F69CFCE1FE44FCDD" bold="true" box="[178,399,838,860]" italics="true" pageId="4" pageNumber="1085">Hovatlas invaginatus</emphasis>
</taxonomicName>
<bibRefCitation id="EFCD4B00FFBF2D72F7B8FCE1FE25FCDD" author="DUARTE RODRIGUES P" box="[406,494,838,859]" pageId="4" pageNumber="1085" pagination="1 - 94" refId="ref3507" refString="DUARTE RODRIGUES P. (1992): Insectes Heteropteres Tingidae. - Faune de Madagascar, Fasc. 78: 1 - 94. Museum National d'Histoire Naturelle, Paris." type="book chapter" year="1992">
<collectingRegion id="4998F813FFBF2D72F7B8FCE1FE25FCDD" box="[406,494,838,859]" country="Dominican Republic" name="Duarte" pageId="4" pageNumber="1085">DUARTE</collectingRegion>
</bibRefCitation>
RODRIGUES, 1992 (
<figureCitation id="13672A74FFBF2D72F493FCE1FCEAFCDA" box="[701,801,838,860]" captionStart="Figs 7-11" captionStartId="5.[178,218,1309,1331]" captionTargetBox="[178,1085,212,1299]" captionTargetId="figure-75@5.[178,1085,212,1299]" captionTargetPageId="5" captionText="Figs 7-11: Tingidae in Madagascan Copal: (7) Hovatlas invaginatus, specimen 1, female, dorsal view; (8) ditto lateral view; (9) ditto ventral view; (10) Hovatlas invaginatus, specimen 2, female, dorsal view; (11) ditto ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5183224" httpUri="https://zenodo.org/record/5183224/files/figure.png" pageId="4" pageNumber="1085">Figs 9-11</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE336F1FFBF2D72F69CFCCBFD94FC1C" blockId="4.[178,1085,876,922]" pageId="4" pageNumber="1085">
M a t e r i a l e x a m i n e d: Two females, antennae and legs complete, specimen
<figureCitation id="13672A74FFBF2D72F5FEFCCBFBC4FC07" box="[976,1039,876,898]" captionStart="Figs 7-11" captionStartId="5.[178,218,1309,1331]" captionTargetBox="[178,1085,212,1299]" captionTargetId="figure-75@5.[178,1085,212,1299]" captionTargetPageId="5" captionText="Figs 7-11: Tingidae in Madagascan Copal: (7) Hovatlas invaginatus, specimen 1, female, dorsal view; (8) ditto lateral view; (9) ditto ventral view; (10) Hovatlas invaginatus, specimen 2, female, dorsal view; (11) ditto ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5183224" httpUri="https://zenodo.org/record/5183224/files/figure.png" pageId="4" pageNumber="1085">Fig. 10</figureCitation>
with damaged, dorsally depressed pronotal hood.
</paragraph>
</subSubSection>
<subSubSection id="C346657AFFBF2D72F69CFC00FEE5FAA2" pageId="4" pageNumber="1085" type="description">
<paragraph id="8BE336F1FFBF2D72F69CFC00FEF9FC73" blockId="4.[178,1085,935,1013]" pageId="4" pageNumber="1085">
The
<typeStatus id="54E78853FFBF2D72F6F1FC00FEF3FC3B" box="[223,312,935,957]" pageId="4" pageNumber="1085" type="holotype">holotype</typeStatus>
of
<emphasis id="B928EAE3FFBF2D72F774FC0FFE1BFC3B" box="[346,464,936,957]" italics="true" pageId="4" pageNumber="1085">invaginatus</emphasis>
is deposited in the Zoological Institute, Russian Academy of Sciences,
<collectingRegion id="4998F813FFBF2D72F70FFC64FE77FC5F" box="[289,444,963,985]" country="Russia" name="Saint Petersburg" pageId="4" pageNumber="1085">St. Petersburg</collectingRegion>
,
<collectingCountry id="F34B7661FFBF2D72F7FEFC64FDD3FC5F" box="[464,536,963,985]" name="Russia" pageId="4" pageNumber="1085">Russia</collectingCountry>
: HT, male,
<collectingCountry id="F34B7661FFBF2D72F485FC64FCEEFC5F" box="[683,805,963,985]" name="Madagascar" pageId="4" pageNumber="1085">Madagascar</collectingCountry>
Est, Périnet,
<date id="FFE21031FFBF2D72F5E5FC64FBF7FC5F" box="[971,1084,963,985]" pageId="4" pageNumber="1085" value="1935-03-13">13.III.1935</date>
(Olsoufieff).
</paragraph>
<paragraph id="8BE336F1FFBF2D72F69CFBA4FCD4FBB3" blockId="4.[178,1084,1027,1077]" pageId="4" pageNumber="1085">
D i s t r i b u t i o n: Endemic to
<collectingCountry id="F34B7661FFBF2D72F437FBA4FD5DFB9F" box="[537,662,1027,1049]" name="Madagascar" pageId="4" pageNumber="1085">Madagascar</collectingCountry>
, only known from Ranomafana National Park, Anabohazo forest and Bemahara Tsingy National Park.
</paragraph>
<paragraph id="8BE336F1FFBF2D72F69CFBE4FE69FBDF" blockId="4.[178,418,1091,1113]" box="[178,418,1091,1113]" pageId="4" pageNumber="1085">Host plant: unrecorded.</paragraph>
<paragraph id="8BE336F1FFBF2D72F69CFBC0FEE5FAA2" blockId="4.[178,1086,1127,1317]" pageId="4" pageNumber="1085">
D i s c u s s i o n: Several specimens belonging to the species
<taxonomicName id="4C5C4D72FFBF2D72F571FBC0FBF6FBFB" authorityName="Rodrigues" authorityYear="1992" box="[863,1085,1127,1149]" class="Insecta" family="Tingidae" genus="Hovatlas" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="1085" phylum="Arthropoda" rank="species" species="invaginatus">
<emphasis id="B928EAE3FFBF2D72F571FBC0FBF6FBFB" box="[863,1085,1127,1149]" italics="true" pageId="4" pageNumber="1085">Hovatlas invaginatus</emphasis>
</taxonomicName>
have an almost entirely biseriate costal area, while the
<typeStatus id="54E78853FFBF2D72F4C3FB24FC8DFB1F" box="[749,838,1155,1177]" pageId="4" pageNumber="1085" type="holotype">holotype</typeStatus>
of the species has been described as having the costal area anteriorly uniseriate. Specimen
<figureCitation id="13672A74FFBF2D72F54EFB38FC18FB32" box="[864,979,1183,1205]" captionStart="Figs 7-11" captionStartId="5.[178,218,1309,1331]" captionTargetBox="[178,1085,212,1299]" captionTargetId="figure-75@5.[178,1085,212,1299]" captionTargetPageId="5" captionText="Figs 7-11: Tingidae in Madagascan Copal: (7) Hovatlas invaginatus, specimen 1, female, dorsal view; (8) ditto lateral view; (9) ditto ventral view; (10) Hovatlas invaginatus, specimen 2, female, dorsal view; (11) ditto ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5183224" httpUri="https://zenodo.org/record/5183224/files/figure.png" pageId="4" pageNumber="1085">Figs 10, 11</figureCitation>
exhibits a costal area not entirely biseriate but uniseriate at the base. In addition, the rostrum of this specimen is shorter than in that of
<figureCitation id="13672A74FFBF2D72F436FB70FD9EFB6A" box="[536,597,1239,1261]" captionStart="Figs 7-11" captionStartId="5.[178,218,1309,1331]" captionTargetBox="[178,1085,212,1299]" captionTargetId="figure-75@5.[178,1085,212,1299]" captionTargetPageId="5" captionText="Figs 7-11: Tingidae in Madagascan Copal: (7) Hovatlas invaginatus, specimen 1, female, dorsal view; (8) ditto lateral view; (9) ditto ventral view; (10) Hovatlas invaginatus, specimen 2, female, dorsal view; (11) ditto ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5183224" httpUri="https://zenodo.org/record/5183224/files/figure.png" pageId="4" pageNumber="1085">Fig. 9</figureCitation>
and the rostral sulcus is wider. As the pronotal hood is damaged in specimen
<figureCitation id="13672A74FFBF2D72F7DEFB54FDA3FA8E" box="[496,616,1267,1289]" captionStart="Figs 7-11" captionStartId="5.[178,218,1309,1331]" captionTargetBox="[178,1085,212,1299]" captionTargetId="figure-75@5.[178,1085,212,1299]" captionTargetPageId="5" captionText="Figs 7-11: Tingidae in Madagascan Copal: (7) Hovatlas invaginatus, specimen 1, female, dorsal view; (8) ditto lateral view; (9) ditto ventral view; (10) Hovatlas invaginatus, specimen 2, female, dorsal view; (11) ditto ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5183224" httpUri="https://zenodo.org/record/5183224/files/figure.png" pageId="4" pageNumber="1085">Figs 10, 11</figureCitation>
, it is not obvious that it really belongs to
<taxonomicName id="4C5C4D72FFBF2D72F20AFB53FEE3FAA3" authorityName="Rodrigues" authorityYear="1992" class="Insecta" family="Tingidae" genus="Hovatlas" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="1085" phylum="Arthropoda" rank="species" species="invaginatus">
<emphasis id="B928EAE3FFBF2D72F20AFB53FEE3FAA3" italics="true" pageId="4" pageNumber="1085">H. invaginatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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