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<document id="4C3A1FD79C07707C74645AEABD6A222B" ID-DOI="10.5281/zenodo.177085" ID-GBIF-Dataset="080ea21f-59f4-49ee-8045-04d275ce86e6" ID-ISSN="1175-5326" ID-Zenodo-Dep="177085" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459848528703" checkinUser="plazi" docAuthor="Stevens, Nicholas B. &amp; Austin, Andrew D." docDate="2007" docId="03F5879F527FD129FF684DC8F650F875" docLanguage="en" docName="zt01499p045.pdf" docOrigin="Zootaxa 1499" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Baeus dux Girault" docType="treatment" docVersion="11" lastPageNumber="18" masterDocId="FFCCFFE7526FD138FFFF4F05F336FFB5" masterDocTitle="Systematics, distribution and biology of the Australian ' micro-flea' wasps, Baeus spp. (Hymenoptera: Scelionidae): parasitoids of spider eggs" masterLastPageNumber="45" masterPageNumber="1" pageNumber="17" updateTime="1698225124475" updateUser="plazi">
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<mods:title id="67BEE2D50D2641AA9372605905B07186">Systematics, distribution and biology of the Australian ' micro-flea' wasps, Baeus spp. (Hymenoptera: Scelionidae): parasitoids of spider eggs</mods:title>
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<mods:namePart id="62DF5AF8C3575024562C8EA9908E0082">Stevens, Nicholas B.</mods:namePart>
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<mods:namePart id="EF47126030E73FF2695D33A38E9FB5D0">Austin, Andrew D.</mods:namePart>
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<treatment id="03F5879F527FD129FF684DC8F650F875" ID-DOI="http://doi.org/10.5281/zenodo.5690870" ID-GBIF-Taxon="127711457" ID-Zenodo-Dep="5690870" LSID="urn:lsid:plazi:treatment:03F5879F527FD129FF684DC8F650F875" httpUri="http://treatment.plazi.org/id/03F5879F527FD129FF684DC8F650F875" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C3466502527FD128FF684DC8F2AEFD52" box="[151,408,717,743]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BE33689527FD128FF684DC8F2AEFD52" blockId="16.[151,408,717,778]" box="[151,408,717,743]" pageId="16" pageNumber="17">
<heading id="D0AB81E5527FD128FF684DC8F2AEFD52" bold="true" box="[151,408,717,743]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B928EA9B527FD128FF684DC8F2AEFD52" bold="true" box="[151,408,717,743]" pageId="16" pageNumber="17">
2.
<taxonomicName id="4C5C4D0A527FD128FF4B4DC8F2AEFD52" ID-CoL="5WBQ7" authority="Girault" authorityName="Girault" box="[180,408,717,743]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dux">
<emphasis id="B928EA9B527FD128FF4B4DC8F205FD52" bold="true" box="[180,307,717,743]" italics="true" pageId="16" pageNumber="17">Baeus dux</emphasis>
Girault
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3466502527FD128FF684DF5F2B4FCBF" box="[151,386,752,778]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BE33689527FD128FF684DF5F2B4FCBF" blockId="16.[151,408,717,778]" box="[151,386,752,778]" pageId="16" pageNumber="17">
(
<figureCitation id="13672A0C527FD128FF604DF5F3D1FCBF" box="[159,231,752,778]" captionStart="FIGURE 8" captionStartId="17.[151,255,1533,1557]" captionTargetBox="[151,1436,194,1509]" captionTargetId="figure@17.[151,1436,194,1509]" captionTargetPageId="17" captionText="FIGURE 8. Female Baeus spp.: A &amp; B, B. dux, cresent shaped carina arrowed: A, lateral habitus; B, dorsal mesosoma. C &amp; D, B. glenysae: C, lateral habitus; D, dorsal habitus. E &amp; F, B. hallarakeri: E, lateral habitus; F, dorsal habitus. Scale lines, A F = 100 μm." httpUri="https://zenodo.org/record/177093/files/figure.png" pageId="16" pageNumber="17">Figs 8</figureCitation>
A &amp; B, 15A)
</paragraph>
</subSubSection>
<subSubSection id="C3466502527FD128FF684C40F77EFCE9" box="[151,1096,837,860]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BE33689527FD128FF684C40F77EFCE9" blockId="16.[151,1096,837,860]" box="[151,1096,837,860]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB4C11A7527FD128FF684C40F77EFCE9" box="[151,1096,837,860]" pageId="16" pageNumber="17">
<taxonomicName id="4C5C4D0A527FD128FF684C40F28FFCE9" ID-CoL="5WBQ7" authority="Girault, 1933: 2" authorityName="Girault" authorityPageNumber="2" authorityYear="1933" box="[151,441,837,860]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dux">
<emphasis id="B928EA9B527FD128FF684C40F233FCE9" box="[151,261,837,860]" italics="true" pageId="16" pageNumber="17">Baeus dux</emphasis>
Girault, 1933: 2
</taxonomicName>
: in Gordth
<emphasis id="B928EA9B527FD128FDCE4C40F151FCE9" box="[561,615,837,860]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
1979: 300;
<treatmentCitation id="0AFD1098527FD128FD194C40F0AAFCE9" author="Austin" box="[742,924,837,860]" page="89" pageId="16" pageNumber="17" year="1981">
<bibRefCitation id="EFCD4B78527FD128FD194C40F042FCE9" author="Austin" box="[742,884,837,860]" pageId="16" pageNumber="44" refString="Austin, A. D. 1981. The types of Australian species in the tribes Idrini, Baeini and Embidobiini (Hymenoptera: Scelionidae: Scelioninae). General and Applied Entomology, 13, 81 - 92." type="journal article" year="1981">Austin, 1981</bibRefCitation>
: 89
</treatmentCitation>
;
<bibRefCitation id="EFCD4B78527FD128FC584C40F772FCE9" author="Johnson" box="[935,1092,837,860]" pageId="16" pageNumber="44" refString="Johnson, N. 2004. Scelionidae. @ http: // atbi. biosci. ohio-state. edu: 210 / hymenoptera / eol _ scelionidae. home (2006)." type="book chapter" year="2004">Johnson, 2004</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3466502527FD128FF684C97F3CCFBF9" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BE33689527FD128FF684C97F760FC19" blockId="16.[151,1437,914,1900]" box="[151,1110,914,940]" pageId="16" pageNumber="17">
<typeStatus id="54E7882B527FD128FF684C97F231FC19" box="[151,263,914,940]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
, Ψ, 'Brisbane, garden on grass,
<date id="FFE21049527FD128FD884C97F03AFC19" box="[631,780,914,940]" pageId="16" pageNumber="17" value="1932-09-08">Sep. 8, 1932</date>
, A. R. Brimblecomb' (QM).
</paragraph>
<paragraph id="8BE33689527FD128FF3A4CBFF3CCFBF9" blockId="16.[151,1437,914,1900]" pageId="16" pageNumber="17">
Other material examined:
<emphasis id="B928EA9B527FD128FE064CBFF1A1FC61" bold="true" box="[505,663,954,980]" pageId="16" pageNumber="17">Queensland:</emphasis>
2 Ψ, Gatton, Qld,
<date id="FFE21049527FD128FC8C4CBFF712FC61" box="[883,1060,954,980]" pageId="16" pageNumber="17" value="1981-04-27" valueMax="1981-05-05" valueMin="1981-04-27">27.iv5.v.1981</date>
, yellow pan trap in potato crop, no collector (
<collectionCode id="ED4DAE4C527FD128FECC4CE7F2B7FC49" LSID="urn:lsid:biocol.org:col:32981" box="[307,385,994,1020]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="16" pageNumber="17">ANIC</collectionCode>
); 2 Ψ, same data,
<date id="FFE21049527FD128FDAA4CE7F1D3FC49" box="[597,741,994,1020]" pageId="16" pageNumber="17" value="1981-05-05" valueMax="1981-05-11" valueMin="1981-05-05">511.v.1981</date>
(
<collectionCode id="ED4DAE4C527FD128FD0A4CE7F075FC49" LSID="urn:lsid:biocol.org:col:32981" box="[757,835,994,1020]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="16" pageNumber="17">ANIC</collectionCode>
); 1 Ψ, same data,
<date id="FFE21049527FD128FBE94CE7F7F1FC49" box="[1046,1223,994,1020]" pageId="16" pageNumber="17" value="1981-05-25" valueMax="1981-06-03" valueMin="1981-05-25">25.v3.vi.1981</date>
(
<collectionCode id="ED4DAE4C527FD128FB284CE7F613FC49" LSID="urn:lsid:biocol.org:col:32981" box="[1239,1317,994,1020]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="16" pageNumber="17">ANIC</collectionCode>
);
<emphasis id="B928EA9B527FD128FAC54CE7F3D8FB91" bold="true" pageId="16" pageNumber="17">
<collectingCountry id="F34B7619527FD128FAC54CE7F3DCFB91" name="Norfolk Island" pageId="16" pageNumber="17">Norfolk Island</collectingCountry>
:
</emphasis>
1 Ψ,
<geoCoordinate id="EE68504E527FD128FEC04B0FF2A7FB91" box="[319,401,1034,1060]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="555" value="-29.07">29.07S</geoCoordinate>
<geoCoordinate id="EE68504E527FD128FE614B0FF132FB91" box="[414,516,1034,1060]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="167.57">167.57E</geoCoordinate>
, Phillip Island, Lower Long Valley,
<date id="FFE21049527FD128FC334B0FF744FB91" box="[972,1138,1034,1060]" pageId="16" pageNumber="17" value="1984-02-20" valueMax="1984-02-24" valueMin="1984-02-20">2024.ii.1984</date>
, T. A. Weir, pitfall trap (
<collectionCode id="ED4DAE4C527FD128FF604B37F3DBFBF9" LSID="urn:lsid:biocol.org:col:32981" box="[159,237,1074,1100]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="16" pageNumber="17">ANIC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3466502527FD129FF3A4B5FF194F965" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BE33689527FD128FF3A4B5FF77FFB29" blockId="16.[151,1437,914,1900]" pageId="16" pageNumber="17">
<emphasis id="B928EA9B527FD128FF3A4B5FF260FBC1" bold="true" box="[197,342,1114,1140]" pageId="16" pageNumber="17">Description</emphasis>
. Female. Mean length
<quantity id="4CA49B6C527FD128FDA04B5FF1FEFBC1" box="[607,712,1114,1140]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="16" pageNumber="17" unit="mm" value="0.64">0.64 mm</quantity>
(0.630.65; n = 3); body and head dark brown, nearly black, leg and antennae segments yellow with light brown markings on dorsal surfaces.
</paragraph>
<paragraph id="8BE33689527FD128FF3A4BAFF064F9B1" blockId="16.[151,1437,914,1900]" pageId="16" pageNumber="17">Head. 1.71 (1.651.82) x as wide as inter-ocular distance and 1.89 (1.752.07) x as wide as long; medial ocellus 10 μm in diameter, 80 μm from posterior head margin; lateral ocelli 5 μm from eye margin, 20 μm from posterior head margin; posterior ocellar line = inter-ocular distance; vertex imbricate, pilosity density gradates, anterior to posterior, from moderate to dense, and is medium length; eyes sub-triangular, eye height 0.44 (0.430.46) x head height, eye width 0.55 (0.50.64) x eye length, pilosity short; frontal carina prominent, reaching 0.55 (0.530.59) distance to medial ocellus; cristulations of malar region reaching to within 10 μm of eye margin; in postero-lateral view, anterior and posterior genal margins parallel medially; anterior genal margin in contact with 0.43 (0.40.5) of ventral eye margin length; posterior eye margin 20 μm from posterior head margin, not contacting hyperoccipital carina.</paragraph>
<paragraph id="8BE33689527FD128FF3A4917F6AAF8D9" blockId="16.[151,1437,914,1900]" pageId="16" pageNumber="17">
Mesosoma. Length 0.72 (0.710.73) x width; both mesoscutum and mesoscutellum imbricate, pilosity mostly dense, patches of moderate density can be present, and of medium length; propodeum glabrous mediodorsally; mesoscutum length 0.60 (0.570.63) x width, 0.73 (0.710.75) x mesosoma length and 3.71 (3.43 4.00) x mesoscutellum length; mesoscutellum length 2.56 (2.333.0) x propodeum length; dorso-lateral region of mesopleuron scrobiculate, sculpturing ending dorsally to dorsal metapleuron margin; dorso-lateral propodeum bearing a semi-circular ridge beginning near posterior margin of the propodeum and extending through ventral region of propodeal spiracle cone, before curving back to end near posterior margin of lateral propodeum (
<figureCitation id="13672A0C527FD128FED1482FF24DF8F1" box="[302,379,1834,1860]" captionStart="FIGURE 8" captionStartId="17.[151,255,1533,1557]" captionTargetBox="[151,1436,194,1509]" captionTargetId="figure@17.[151,1436,194,1509]" captionTargetPageId="17" captionText="FIGURE 8. Female Baeus spp.: A &amp; B, B. dux, cresent shaped carina arrowed: A, lateral habitus; B, dorsal mesosoma. C &amp; D, B. glenysae: C, lateral habitus; D, dorsal habitus. E &amp; F, B. hallarakeri: E, lateral habitus; F, dorsal habitus. Scale lines, A F = 100 μm." httpUri="https://zenodo.org/record/177093/files/figure.png" pageId="16" pageNumber="17">Figs 8</figureCitation>
A&amp;B); propodeal spiracle small and ovoid; posterior suture of metapleuron straight and short, dorsal extent of suture ending below the level of antero-lateral margin of T2; hind femoral spine absent.
</paragraph>
<caption id="DF236601527ED129FF684AF8F2FFF9EC" httpUri="https://zenodo.org/record/177093/files/figure.png" pageId="17" pageNumber="18" targetBox="[151,1436,194,1509]" targetPageId="17">
<paragraph id="8BE33689527ED129FF684AF8F2FFF9EC" blockId="17.[151,1437,1533,1984]" pageId="17" pageNumber="18">
<emphasis id="B928EA9B527ED129FF684AF8F22FF9A0" bold="true" box="[151,281,1533,1557]" pageId="17" pageNumber="18">FIGURE 8.</emphasis>
Female
<taxonomicName id="4C5C4D0A527ED129FE894AFBF281F9A0" box="[374,439,1534,1557]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA9B527ED129FE894AFBF281F9A0" box="[374,439,1534,1557]" italics="true" pageId="17" pageNumber="18">Baeus</emphasis>
</taxonomicName>
spp.:
<emphasis id="B928EA9B527ED129FE054AF8F13BF9A0" bold="true" box="[506,525,1533,1557]" pageId="17" pageNumber="18">A</emphasis>
&amp;
<emphasis id="B928EA9B527ED129FDCF4AF8F174F9A0" bold="true" box="[560,578,1533,1557]" pageId="17" pageNumber="18">B</emphasis>
,
<taxonomicName id="4C5C4D0A527ED129FDAF4AFBF1A2F9A0" box="[592,660,1534,1557]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dux">
<emphasis id="B928EA9B527ED129FDAF4AFBF1A2F9A0" box="[592,660,1534,1557]" italics="true" pageId="17" pageNumber="18">B. dux</emphasis>
</taxonomicName>
, cresent shaped carina arrowed:
<emphasis id="B928EA9B527ED129FC0A4AF8F73EF9A0" bold="true" box="[1013,1032,1533,1557]" pageId="17" pageNumber="18">A</emphasis>
, lateral habitus;
<emphasis id="B928EA9B527ED129FB434AF8F7F8F9A0" bold="true" box="[1212,1230,1533,1557]" pageId="17" pageNumber="18">B</emphasis>
, dorsal mesosoma.
<emphasis id="B928EA9B527ED129FF684925F39CF98D" bold="true" box="[151,170,1568,1592]" pageId="17" pageNumber="18">C</emphasis>
&amp;
<emphasis id="B928EA9B527ED129FF2E4925F3D2F98D" bold="true" box="[209,228,1568,1592]" pageId="17" pageNumber="18">D</emphasis>
,
<taxonomicName id="4C5C4D0A527ED129FF0B4925F247F982" box="[244,369,1568,1591]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="glenysae">
<emphasis id="B928EA9B527ED129FF0B4925F247F982" box="[244,369,1568,1591]" italics="true" pageId="17" pageNumber="18">B. glenysae</emphasis>
</taxonomicName>
:
<emphasis id="B928EA9B527ED129FE7E4925F2A2F98D" bold="true" box="[385,404,1568,1592]" pageId="17" pageNumber="18">C</emphasis>
, lateral habitus;
<emphasis id="B928EA9B527ED129FDB24925F156F98D" bold="true" box="[589,608,1568,1592]" pageId="17" pageNumber="18">D</emphasis>
, dorsal habitus.
<emphasis id="B928EA9B527ED129FCE84925F01FF98D" bold="true" box="[791,809,1568,1592]" pageId="17" pageNumber="18">E</emphasis>
&amp;
<emphasis id="B928EA9B527ED129FCAF4925F068F98D" bold="true" box="[848,862,1568,1592]" pageId="17" pageNumber="18">F</emphasis>
,
<taxonomicName id="4C5C4D0A527ED129FC924925F733F982" box="[877,1029,1568,1591]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hallarakeri">
<emphasis id="B928EA9B527ED129FC924925F733F982" box="[877,1029,1568,1591]" italics="true" pageId="17" pageNumber="18">B. hallarakeri</emphasis>
</taxonomicName>
:
<emphasis id="B928EA9B527ED129FBE94925F71EF98D" bold="true" box="[1046,1064,1568,1592]" pageId="17" pageNumber="18">E</emphasis>
, lateral habitus;
<emphasis id="B928EA9B527ED129FB1F4925F7D8F98D" bold="true" box="[1248,1262,1568,1592]" pageId="17" pageNumber="18">F</emphasis>
, dorsal habitus. Scale lines, A F = 100 μm.
</paragraph>
</caption>
<paragraph id="8BE33689527ED129FF3A4963F194F965" blockId="17.[151,1437,1533,1984]" pageId="17" pageNumber="18">Metasoma. T2 length 0.9 x width, anterior region imbricate to finely coriarious, to smooth posteriorly, pilosity dense anteriorly to moderately dense posteriorly, of medium length overall; T3 smooth, bearing one row of setae of medium length; T4 glabrous.</paragraph>
</subSubSection>
<subSubSection id="C3466502527ED129FF3A49D8F650F875" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BE33689527ED129FF3A49D8F650F875" blockId="17.[151,1437,1533,1984]" pageId="17" pageNumber="18">
<emphasis id="B928EA9B527ED129FF3A49D8F27AF942" bold="true" box="[197,332,1757,1783]" pageId="17" pageNumber="18">Comments</emphasis>
. The distinguishing feature for
<taxonomicName id="4C5C4D0A527ED129FD3349DBF02CF942" box="[716,794,1758,1783]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dux">
<emphasis id="B928EA9B527ED129FD3349DBF02CF942" box="[716,794,1758,1783]" italics="true" pageId="17" pageNumber="18">B. dux</emphasis>
</taxonomicName>
is its distinct semi-circular ridge on the dorso-lateral propodeum.
<taxonomicName id="4C5C4D0A527ED129FED34803F29FF8AA" box="[300,425,1798,1823]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="leai">
<emphasis id="B928EA9B527ED129FED34803F29FF8AA" box="[300,425,1798,1823]" italics="true" pageId="17" pageNumber="18">Baeus leai</emphasis>
</taxonomicName>
is the only other species to possess a similar structure. However, the two species are very different in many other characters, namely, the shape of the gena, sculpturing of the dorsal mesosomal, and smaller body and eye size.
<taxonomicName id="4C5C4D0A527ED129FDF24853F1BEF8DA" box="[525,648,1878,1903]" class="Insecta" family="Scelionidae" genus="Baeus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dux">
<emphasis id="B928EA9B527ED129FDF24853F1BEF8DA" box="[525,648,1878,1903]" italics="true" pageId="17" pageNumber="18">Baeus dux</emphasis>
</taxonomicName>
has been collected from south-eastern Queensland (
<figureCitation id="13672A0C527ED129FB0B4853F67FF8C5" box="[1268,1353,1878,1904]" captionStart="FIGURE 15" captionStartId="39.[151,255,1329,1353]" captionTargetBox="[158,1412,197,1279]" captionTargetId="figure@39.[157,1430,194,1305]" captionTargetPageId="39" captionText="FIGURE 15. Distribution maps: A B. arthuri, B. jenningsi, B. dux, B. glenysae; B B. hallarakeri, B. iqbali; C B. leai; D B. maryae." httpUri="https://zenodo.org/record/177100/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
A) and Philip Island near
<collectingCountry id="F34B7619527ED129FE93487BF12CF82D" box="[364,538,1918,1944]" name="Norfolk Island" pageId="17" pageNumber="18">Norfolk Island</collectingCountry>
. The habitats from which some specimens were collected include a suburban garden and a potato crop, indicating that the host(s) are tolerant of, or may prefer, heavily disturbed sites.
</paragraph>
</subSubSection>
</treatment>
</document>