353 lines
75 KiB
XML
353 lines
75 KiB
XML
<document id="1B72FC2D695C70619BA388918C256BF3" ID-CLB-Dataset="5899" ID-DOI="10.11646/zootaxa.4981.3.5" ID-GBIF-Dataset="fce50f60-b834-424f-9a19-cf4601ffd2a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921345" ID-ZooBank="3F94E1C8-2F49-4B85-86E3-48341F17A358" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623281889193" checkinUser="plazi" docAuthor="Rafael, José Albertino & Marques, Dayse Willkenia A." docDate="2021" docId="03F5878CFFC9FFF275D1AC68C7BAADF6" docLanguage="en" docName="zootaxa.4981.3.5.pdf" docOrigin="Zootaxa 4981 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrostomus chelicercus Rafael & Marques 2021, sp. nov." docType="treatment" docUuid="EFD706BA-BAD1-4542-85C0-2A78DED937A3" docUuidSource="ZooBank" docVersion="8" lastPageNumber="517" masterDocId="FFCCFFF4FFC3FFFF7546AA0DC66DAC5B" masterDocTitle="Four new species of Macrostomus Wiedemann and a checklist of Empididae s. str (Diptera) from Peru" masterLastPageNumber="530" masterPageNumber="506" pageNumber="516" updateTime="1699009141239" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="D05A0F86FA01BCC4AA0D59C5669B9A6F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="5F52B03DB64A3B527597665098E8BBD4">
|
||
<mods:title id="2BB5969E8A3CBC056A10A99E61A48F5A">Four new species of Macrostomus Wiedemann and a checklist of Empididae s. str (Diptera) from Peru</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="36FF02D3B943C397EA74122D79AB36C3" type="personal">
|
||
<mods:role id="588D1139536D48F4FB0B9996492B89DF">
|
||
<mods:roleTerm id="CD2099C11C05B3E54E6EB1083EF3BAAD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A92CC9C44CA712E3A70FA3A50CC74F67">Rafael, José Albertino</mods:namePart>
|
||
<mods:nameIdentifier id="50328DAD29336E874A2D9A4F68B12BCF" type="ORCID">0000-0002-0170-0514</mods:nameIdentifier>
|
||
<mods:affiliation id="52BB4E49CA2AD90F346AFEE1388B5F84">Instituto Nacional de Pesquisas da Amazônia, Coordenação de Biodiversidade, Manaus, Amazonas, Brazil. jarafael @ inpa. gov. br; https: // orcid. org / 0000 - 0002 - 0170 - 0514</mods:affiliation>
|
||
<mods:nameIdentifier id="FBBE7BC5DC5C58D8DB5C16DDBA2EB347" type="email">jarafael@inpa.gov.br</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="6570DE8D55F814DA8AD3CFFC501A7B64" type="personal">
|
||
<mods:role id="24CFAC984A5D3A7DD7B1E349177C09AF">
|
||
<mods:roleTerm id="7A7BEFE21742EC4E3587A59951CE35CE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7E90E1111F142970775F1691F993D6B6">Marques, Dayse Willkenia A.</mods:namePart>
|
||
<mods:nameIdentifier id="ECA3778F259F05DEE75701908EEBB49D" type="ORCID">0000-0001-7260-5760</mods:nameIdentifier>
|
||
<mods:affiliation id="0C2096952D62BBC0AC034E2A45EA6EBF">Instituto Nacional de Pesquisas da Amazônia, Programa de Pós-Graduação em Entomologia, Manaus, Amazonas, Brazil. willkenia @ gmail. com; https: // orcid. org / 0000 - 0001 - 7260 - 5760</mods:affiliation>
|
||
<mods:nameIdentifier id="B091ECE6651F167465B1F00D45F70CC1" type="email">willkenia@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="74A1D34B4E8B781B4906407696B37788">text</mods:typeOfResource>
|
||
<mods:relatedItem id="21DA33F9C83142B67F97ED9AAC12E1CA" type="host">
|
||
<mods:titleInfo id="AE75CB820DE4BE24426B169D40AA3CCC">
|
||
<mods:title id="59B5E1F4145BAC9EF58D9D07527331EA">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="36ADF13EDDD39F7C871B2EC9A36EBDF5">
|
||
<mods:date id="E4658C1886C7C64B777C0E3E14EE7787">2021</mods:date>
|
||
<mods:detail id="49B53EE5D329629F3BA027C9917BBDC7" type="pubDate">
|
||
<mods:number id="EA5E1263F5A343191D8496F0FE93894D">2021-06-08</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D228CE1923CAA87E5A4F5715DCAF5D15" type="volume">
|
||
<mods:number id="221516D34B55BFFE00C87E32D15FE1C0">4981</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="02E60BA9E5AF6DDBFDF40A375464097A" type="issue">
|
||
<mods:number id="1F8D124E78887C2CD92AFBF21E04375A">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="D14D19B2CB36616CDBE13498C224C910" unit="page">
|
||
<mods:start id="D68DB8333A05BAF6C4FE4D5C18C18E25">506</mods:start>
|
||
<mods:end id="C4B630DED9CA47C4B27E6AB1B1EA2C35">530</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="6497998B4F62A7F59FACDF19984F2BDF">journal article</mods:classification>
|
||
<mods:identifier id="D02011C857DE65FBB1484BA212FB6E4D" type="CLB-Dataset">5899</mods:identifier>
|
||
<mods:identifier id="778738E1FF8ED6778C4ECE78EBE4179D" type="DOI">10.11646/zootaxa.4981.3.5</mods:identifier>
|
||
<mods:identifier id="2B09D7826F3A2C1D1F43BA27FE508E86" type="GBIF-Dataset">fce50f60-b834-424f-9a19-cf4601ffd2a3</mods:identifier>
|
||
<mods:identifier id="E5EEFFB70822B5309937427775490DF8" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="5E83BC4540A2985FC4E085F92F9969A9" type="Zenodo-Dep">4921345</mods:identifier>
|
||
<mods:identifier id="4E1FC6AA54948F2DA17C5DB5340612B8" type="ZooBank">3F94E1C8-2F49-4B85-86E3-48341F17A358</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F5878CFFC9FFF275D1AC68C7BAADF6" ID-DOI="http://doi.org/10.5281/zenodo.5047207" ID-GBIF-Taxon="183228122" ID-Zenodo-Dep="5047207" LSID="urn:lsid:zoobank.org:act:EFD706BA-BAD1-4542-85C0-2A78DED937A3" httpUri="http://treatment.plazi.org/id/03F5878CFFC9FFF275D1AC68C7BAADF6" lastPageId="13" lastPageNumber="517" pageId="10" pageNumber="516">
|
||
<subSubSection id="C3466511FFC9FFF575D1AC68C447AA24" box="[151,554,1636,1663]" pageId="10" pageNumber="516" type="nomenclature">
|
||
<paragraph id="8BE3369AFFC9FFF575D1AC68C447AA24" blockId="10.[151,1151,1636,1728]" box="[151,554,1636,1663]" pageId="10" pageNumber="516">
|
||
<heading id="D0AB81F6FFC9FFF575D1AC68C447AA24" bold="true" box="[151,554,1636,1663]" fontSize="11" level="1" pageId="10" pageNumber="516" reason="1">
|
||
<emphasis id="B928EA88FFC9FFF575D1AC68C447AA24" bold="true" box="[151,554,1636,1663]" pageId="10" pageNumber="516">
|
||
<taxonomicName id="4C5C4D19FFC9FFF575D1AC68C7A5AA24" ID-CoL="9HTKZ" authority="Rafael & Marques, 2021" authorityName="Rafael & Marques" authorityYear="2021" box="[151,456,1636,1663]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="516" phylum="Arthropoda" rank="species" species="chelicercus" status="sp. nov.">
|
||
<emphasis id="B928EA88FFC9FFF575D1AC68C7A5AA24" bold="true" box="[151,456,1636,1663]" italics="true" pageId="10" pageNumber="516">Macrostomus chelicercus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21B57F3FFC9FFF57489AC68C447AA24" box="[463,554,1637,1663]" pageId="10" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466511FFC9FFF575D1AC84C212AA9B" pageId="10" pageNumber="516" type="description">
|
||
<paragraph id="8BE3369AFFC9FFF575D1AC84C718AAF8" blockId="10.[151,1151,1636,1728]" box="[151,373,1673,1699]" pageId="10" pageNumber="516">
|
||
(
|
||
<figureCitation id="13672A1FFFC9FFF575D9AC84C772AAF8" box="[159,287,1673,1699]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="10" pageNumber="516">Figs 8A–N</figureCitation>
|
||
,
|
||
<figureCitation id="13672A1FFFC9FFF5746CAC84C700AAF8" box="[298,365,1673,1699]" captionStart="FIGURE 9" captionStartId="13.[151,250,1016,1040]" captionTargetBox="[156,1429,461,990]" captionTargetId="figure-138@13.[156,1430,453,993]" captionTargetPageId="13" captionText="FIGURE 9. Macrostomus chelicercus sp. nov., paratype ♀. A. Tergite 8, tergite 9+10, sternite 8, sternite 10 and cercus, lateral view; B. Tergite 8, tergite 9+10 and cerci, dorsal view; C. Sternite 8, sternite 10 and cerci, ventral view; D. Tergite 8, tergite 9+10 and cerci, dorsal view (inferior pieces), sternite 8, ventral view; E. Genital fork and genital chamber, ventral view. Scale bars, figures A–E = 0.2 mm. Abbreviations: gen ch, genital chamber; gen fk, genital fork; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921373" httpUri="https://zenodo.org/record/4921373/files/figure.png" pageId="10" pageNumber="516">9A–E</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BE3369AFFC9FFF575D1ACA5C212AA9B" blockId="10.[151,1151,1636,1728]" box="[151,1151,1704,1728]" pageId="10" pageNumber="516">
|
||
http://zoobank.org/
|
||
<uri id="FFCD3A98FFC9FFF57421ACA5C212AA9B" box="[359,1151,1704,1728]" pageId="10" pageNumber="516">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFFA0C4FFFC9FFF5772FACA5C212AA9B" box="[617,1151,1704,1728]" pageId="10" pageNumber="516">EFD706BA-BAD1-4542-85C0-2A78DED937A3</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466511FFC9FFF575D1ACE3C5A5ABC3" pageId="10" pageNumber="516" type="diagnosis">
|
||
<paragraph id="8BE3369AFFC9FFF575D1ACE3C5A5ABC3" blockId="10.[151,1437,1774,2017]" pageId="10" pageNumber="516">
|
||
<emphasis id="B928EA88FFC9FFF575D1ACE3C736AB53" bold="true" box="[151,347,1774,1800]" pageId="10" pageNumber="516">Diagnosis. Male.</emphasis>
|
||
Thorax with mesonotum, scutellum and mediotergite shiny brown to black; postpronotal lobe and mesopleuron shiny yellow; pronotum and notopleuron partially shiny brown to black and partially yellow; mid and hind femora and tibiae slightly curved; hind femur clavate from basal 2/3; tergite 8 short, with shallow basal sinus, sinuose posteriorly; anterior cercus with U-shaped cercal bridge; posterior cercus cheliform, distinctly inward and forward curved, bifid at apex; phallus distinctly longer than hypandrium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466511FFC9FFF47581ADAFC41CA9C6" lastPageId="11" lastPageNumber="517" pageId="10" pageNumber="516" type="description">
|
||
<paragraph id="8BE3369AFFC9FFF47581ADAFC2CFA802" blockId="10.[151,1437,1774,2017]" lastBlockId="11.[151,1437,151,2013]" lastPageId="11" lastPageNumber="517" pageId="10" pageNumber="516">
|
||
<emphasis id="B928EA88FFC9FFF57581ADAFC734ABE7" bold="true" box="[199,345,1954,1980]" pageId="10" pageNumber="516">Description.</emphasis>
|
||
<typeStatus id="54E78838FFC9FFF57427ADAFC7A4ABE7" box="[353,457,1954,1980]" pageId="10" pageNumber="516" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D5AFD13FFC9FFF57494ADAEC78AABE6" box="[466,487,1955,1981]" pageId="10" pageNumber="516" type="male">♂</specimenCount>
|
||
(
|
||
<figureCitation id="13672A1FFFC9FFF574BEADAFC439ABE7" box="[504,596,1954,1980]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="10" pageNumber="516">Fig. 8A</figureCitation>
|
||
). Body length
|
||
<quantity id="4CA49B7FFFC9FFF577BBADAFC535ABE6" box="[765,856,1954,1981]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="516" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; wing length 3.0 mm. Head dichoptic. Frons shiny black narrower than width of anterior ocellus, narrower ventrally. One pair of proclinate ocellar setae. Face as wide as frons ventrally, less shiny black than frons, slightly gray pruinose. Postcranium shiny black, sparsely gray pruinose, except dorsal postocular margin shiny without pruinescence. Inner and outer vertical setae broken. Gena with 2 small setae and postgena with 1–2 minute setae. Antenna velvety brown to black with postpedicel short setulose. Proboscis mainly yellow. Palpus brown to black. Thorax (
|
||
<figureCitation id="13672A1FFFC8FFF47665AB0EC516AD46" box="[803,891,259,285]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8A</figureCitation>
|
||
) shiny brown to black dorsally, partially on pronotum, entirely on mesonotum, partially on notopleuron, entirely on scutellum and mediotergite; shiny yellow on postpronotal lobe, partially on notopleuron and entirely on mesopleuron. Thoracic chaetotaxy: 3 antepronotals; 1 postpronotal stout and 1–2 minute; 4 dorsocentrals, with posterior pair stouter; 1 presutural supra-alar; 0 postsutural supra-alar; 1 postalar stout, 1 minute; 2–3 small proepisternals; 1 notopleural stout, 1 weaker; 2 pairs of scutellars, outer pair weaker; 4 laterotergitals. Legs (
|
||
<figureCitation id="13672A1FFFC8FFF47733ABBAC498AD8A" box="[629,757,439,465]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Figs 8A, B</figureCitation>
|
||
) shiny yellow except apices of mid and hind femora, distal half of hind tibia, mid and hind tarsi brown to black. Mid and hind femora and tibiae slightly curved. Hind femur clavate from basal 2/3. Mid tibia with slender anterodorsal distal seta; hind tibia with 1 short seta at basal fourth and 2 longer setae at distal fourth; hind tarsomere 1 with 3 longer dorsal setae at distal half. Wing (
|
||
<figureCitation id="13672A1FFFC8FFF4719BA82EC358AE66" box="[1245,1333,547,573]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8A</figureCitation>
|
||
) hyaline, slightly brown infuscate at pterostigma and apex of cell r
|
||
<subScript id="17D834DFFFC8FFF47657A854C547AE3C" attach="right" box="[785,810,601,615]" fontSize="6" pageId="11" pageNumber="517">2+3</subScript>
|
||
. Veins M
|
||
<subScript id="17D834DFFFC8FFF476D3A854C5F0AE3C" attach="left" box="[917,925,601,615]" fontSize="6" pageId="11" pageNumber="517">1</subScript>
|
||
, M
|
||
<subScript id="17D834DFFFC8FFF47685A854C5A6AE3C" attach="left" box="[963,971,601,615]" fontSize="6" pageId="11" pageNumber="517">2</subScript>
|
||
and CuA+CuP evanescent. Distal section of vein M
|
||
<subScript id="17D834DFFFC8FFF4744EA870C77DAED0" attach="left" box="[264,272,637,651]" fontSize="6" pageId="11" pageNumber="517">4</subScript>
|
||
longer than vein dm–m length. Cell dm elongate, somewhat truncate. Halter yellow with knob brown. Abdomen (
|
||
<figureCitation id="13672A1FFFC8FFF47452A882C706AEF2" box="[276,363,655,681]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8A</figureCitation>
|
||
) with tergite 1 and base of tergite 2 shiny yellow, remaining shiny brown to black. Tergite 8 (
|
||
<figureCitation id="13672A1FFFC8FFF4702DA882C6B5AE96" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Figs 8C–E</figureCitation>
|
||
) short, with shallow basal sinus, sinuose posteriorly. Sternite 8 (
|
||
<figureCitation id="13672A1FFFC8FFF476E4A8BEC275AE96" box="[930,1048,691,717]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Figs 8E, F</figureCitation>
|
||
) with posterolateral saddle-shaped projection. Tergite 8 narrowly articulated anterolaterally with sternite 8 (
|
||
<figureCitation id="13672A1FFFC8FFF4769EA8DAC25FAEAA" box="[984,1074,727,753]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8E</figureCitation>
|
||
). Epandrium (
|
||
<figureCitation id="13672A1FFFC8FFF47190A8DAC335AEAA" box="[1238,1368,727,753]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Figs 8G, H</figureCitation>
|
||
) with posterodorsal lobe higher than anterodorsal lobe, latter with small rounded mesial lobe and longer setae posteriorly and dorsally, without anterodorsal nor anteroventral cleft. Anterior cercus (
|
||
<figureCitation id="13672A1FFFC8FFF476AFA912C22FAF62" box="[1001,1090,799,825]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8G</figureCitation>
|
||
) subtriangular, wider dorsally, with slightly longer setae dorsally and triangular mesial process of anterior cercus touching each other but not fused (
|
||
<figureCitation id="13672A1FFFC8FFF475D9A96AC760AFDA" box="[159,269,871,897]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Figs 8I, J</figureCitation>
|
||
); cercal bridge U-shaped (
|
||
<figureCitation id="13672A1FFFC8FFF4777BA96AC4E0AFDA" box="[573,653,871,897]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8I</figureCitation>
|
||
). Median cercus (
|
||
<figureCitation id="13672A1FFFC8FFF4761CA96AC5DEAFDA" box="[858,947,871,897]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8G</figureCitation>
|
||
) somewhat translucent, subsquare, upward directed with longer marginal setae; posterior cercus (
|
||
<figureCitation id="13672A1FFFC8FFF47646A986C534AFFE" box="[768,857,907,933]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8G</figureCitation>
|
||
) cheliform, distinctly inward and forward directed with apex bifid. Ventral projection of cercus (
|
||
<figureCitation id="13672A1FFFC8FFF477E0A9A2C49BAF92" box="[678,758,943,969]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8J</figureCitation>
|
||
) short, transversely positioned, with 3–4 stout distal setae. Dorsal hypoproctal lobe (
|
||
<figureCitation id="13672A1FFFC8FFF474FDA9DEC459AFB6" box="[443,564,979,1005]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Figs 8J, K</figureCitation>
|
||
) large, with slender marginal setae; ventral hypoproctal lobe inconspicuous. Hypandrium (
|
||
<figureCitation id="13672A1FFFC8FFF47472A9FAC7D8A84A" box="[308,437,1015,1041]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Figs 8L, M</figureCitation>
|
||
) in lateral view slightly wider distally, apex with two protuberances, anterior shorter and reclined; posterior margin of hypandrium with 7 uniseriate setae. Ejaculatory apodeme (
|
||
<figureCitation id="13672A1FFFC8FFF47120AE16C2D4A86E" box="[1126,1209,1051,1077]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8L</figureCitation>
|
||
) with dorsal lamella slightly higher than ventral lamella. Phallus (
|
||
<figureCitation id="13672A1FFFC8FFF477C8AE32C563A802" box="[654,782,1087,1113]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Figs 8L, M</figureCitation>
|
||
) distinctly longer than hypandrium.
|
||
</paragraph>
|
||
<paragraph id="8BE3369AFFC8FFF47581AE6EC24BA922" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="517">
|
||
<emphasis id="B928EA88FFC8FFF47581AE6EC748A826" bold="true" box="[199,293,1123,1149]" pageId="11" pageNumber="517">Female.</emphasis>
|
||
One specimen in poor condition appears to be conspecific with male specimens. Body and wing length not measured. Frons shiny black slightly wider than anterior ocellus width, narrower ventrally. Face slightly narrower than frons, slightly gray pruinose. Postcranium shiny black, entirely sparsely gray pruinose. Legs shiny yellow with apices of distal tarsomeres brown. All legs not curved, nor clavate. Wings damaged. Abdomen entirely shiny black. Tergite 8 damaged, apex with deep distal sinus (
|
||
<figureCitation id="13672A1FFFC8FFF47786AEFEC52EA956" box="[704,835,1267,1293]" captionStart="FIGURE 9" captionStartId="13.[151,250,1016,1040]" captionTargetBox="[156,1429,461,990]" captionTargetId="figure-138@13.[156,1430,453,993]" captionTargetPageId="13" captionText="FIGURE 9. Macrostomus chelicercus sp. nov., paratype ♀. A. Tergite 8, tergite 9+10, sternite 8, sternite 10 and cercus, lateral view; B. Tergite 8, tergite 9+10 and cerci, dorsal view; C. Sternite 8, sternite 10 and cerci, ventral view; D. Tergite 8, tergite 9+10 and cerci, dorsal view (inferior pieces), sternite 8, ventral view; E. Genital fork and genital chamber, ventral view. Scale bars, figures A–E = 0.2 mm. Abbreviations: gen ch, genital chamber; gen fk, genital fork; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921373" httpUri="https://zenodo.org/record/4921373/files/figure.png" pageId="11" pageNumber="517">Figs 9B, D</figureCitation>
|
||
). Tergite 9+10 (
|
||
<figureCitation id="13672A1FFFC8FFF476BCAEFEC217A956" box="[1018,1146,1267,1293]" captionStart="FIGURE 9" captionStartId="13.[151,250,1016,1040]" captionTargetBox="[156,1429,461,990]" captionTargetId="figure-138@13.[156,1430,453,993]" captionTargetPageId="13" captionText="FIGURE 9. Macrostomus chelicercus sp. nov., paratype ♀. A. Tergite 8, tergite 9+10, sternite 8, sternite 10 and cercus, lateral view; B. Tergite 8, tergite 9+10 and cerci, dorsal view; C. Sternite 8, sternite 10 and cerci, ventral view; D. Tergite 8, tergite 9+10 and cerci, dorsal view (inferior pieces), sternite 8, ventral view; E. Genital fork and genital chamber, ventral view. Scale bars, figures A–E = 0.2 mm. Abbreviations: gen ch, genital chamber; gen fk, genital fork; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921373" httpUri="https://zenodo.org/record/4921373/files/figure.png" pageId="11" pageNumber="517">Figs 9A, B</figureCitation>
|
||
) short with distinct basal sinus. Sternite 8 (
|
||
<figureCitation id="13672A1FFFC8FFF4741EAF1AC7B9A96A" box="[344,468,1303,1329]" captionStart="FIGURE 9" captionStartId="13.[151,250,1016,1040]" captionTargetBox="[156,1429,461,990]" captionTargetId="figure-138@13.[156,1430,453,993]" captionTargetPageId="13" captionText="FIGURE 9. Macrostomus chelicercus sp. nov., paratype ♀. A. Tergite 8, tergite 9+10, sternite 8, sternite 10 and cercus, lateral view; B. Tergite 8, tergite 9+10 and cerci, dorsal view; C. Sternite 8, sternite 10 and cerci, ventral view; D. Tergite 8, tergite 9+10 and cerci, dorsal view (inferior pieces), sternite 8, ventral view; E. Genital fork and genital chamber, ventral view. Scale bars, figures A–E = 0.2 mm. Abbreviations: gen ch, genital chamber; gen fk, genital fork; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921373" httpUri="https://zenodo.org/record/4921373/files/figure.png" pageId="11" pageNumber="517">Figs 9A, C</figureCitation>
|
||
) somewhat truncate distally, with incision laterally and mid-longitudinal hyaline stripe ventrally, stripe almost reaching apex. Sternite 10 with basal sinus. Cercus cylindrical. Genital fork (
|
||
<figureCitation id="13672A1FFFC8FFF471A9AF36C329A90E" box="[1263,1348,1339,1365]" captionStart="FIGURE 9" captionStartId="13.[151,250,1016,1040]" captionTargetBox="[156,1429,461,990]" captionTargetId="figure-138@13.[156,1430,453,993]" captionTargetPageId="13" captionText="FIGURE 9. Macrostomus chelicercus sp. nov., paratype ♀. A. Tergite 8, tergite 9+10, sternite 8, sternite 10 and cercus, lateral view; B. Tergite 8, tergite 9+10 and cerci, dorsal view; C. Sternite 8, sternite 10 and cerci, ventral view; D. Tergite 8, tergite 9+10 and cerci, dorsal view (inferior pieces), sternite 8, ventral view; E. Genital fork and genital chamber, ventral view. Scale bars, figures A–E = 0.2 mm. Abbreviations: gen ch, genital chamber; gen fk, genital fork; st, sternite; t, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921373" httpUri="https://zenodo.org/record/4921373/files/figure.png" pageId="11" pageNumber="517">Fig. 9E</figureCitation>
|
||
) simple rod-shaped, slightly wider distally. Genital chamber predominantly membranous.
|
||
</paragraph>
|
||
<paragraph id="8BE3369AFFC8FFF47581AF8FC41CA9C6" blockId="11.[151,1437,151,2013]" box="[199,625,1410,1437]" pageId="11" pageNumber="517">
|
||
<emphasis id="B928EA88FFC8FFF47581AF8FC7A3A9C6" bold="true" box="[199,462,1410,1437]" pageId="11" pageNumber="517">Geographical records.</emphasis>
|
||
<collectingCountry id="F34B760AFFC8FFF47493AF8EC465A9C6" box="[469,520,1411,1437]" name="Peru" pageId="11" pageNumber="517">Peru</collectingCountry>
|
||
(
|
||
<collectingRegion id="4998F878FFC8FFF4775EAF8EC408A9C6" box="[536,613,1411,1437]" country="Peru" name="Cusco" pageId="11" pageNumber="517">Cuzco</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466511FFC8FFF47581AFAAC695AAC2" pageId="11" pageNumber="517" type="materials_examined">
|
||
<paragraph id="8BE3369AFFC8FFF47581AFAAC461AA0A" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="517">
|
||
<emphasis id="B928EA88FFC8FFF47581AFAAC790A99B" bold="true" box="[199,509,1446,1473]" pageId="11" pageNumber="517">Type Material examined.</emphasis>
|
||
<materialsCitation id="3B343CC7FFC8FFF4774FAFABC422AA52" ID-GBIF-Occurrence="3312253315" collectingDate="2012-08-22" collectingDateMax="2012-08-26" collectingDateMin="2012-08-22" collectionCode="MUSM" collectorName="J. A. Rafael & R. R. Cavichioli" country="Peru" elevation="1000" latitude="-13.3550005" location="Rio Araza" longLatPrecision="21" longitude="-70.88944" municipality="Quincemil" pageId="11" pageNumber="517" specimenCount="2" specimenCount-male="2" stateProvince="Cusco" typeStatus="holotype">
|
||
<emphasis id="B928EA88FFC8FFF4774FAFABC4C5A99B" bold="true" box="[521,680,1446,1472]" pageId="11" pageNumber="517">
|
||
<typeStatus id="54E78838FFC8FFF4774FAFABC4C5A99B" box="[521,680,1446,1472]" pageId="11" pageNumber="517" type="holotype">HOLOTYPE</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D5AFD13FFC8FFF477F2AFAAC4A6A99A" box="[692,715,1447,1473]" pageId="11" pageNumber="517" type="male">♂</specimenCount>
|
||
, labelled: “
|
||
<collectingCountry id="F34B760AFFC8FFF47614AFAAC5F2A99A" box="[850,927,1447,1473]" name="Peru" pageId="11" pageNumber="517">PERU</collectingCountry>
|
||
,
|
||
<collectingRegion id="4998F878FFC8FFF476E9AFAAC59AA99A" box="[943,1015,1447,1473]" country="Peru" name="Cusco" pageId="11" pageNumber="517">Cuzco</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B87ACE0FFC8FFF4714EAFAAC213A99A" box="[1032,1150,1447,1473]" pageId="11" pageNumber="517">Quincemil</collectingMunicipality>
|
||
,
|
||
<location id="8E836041FFC8FFF471C8AFAAC36EA99A" LSID="urn:lsid:plazi:treatment:03F5878CFFC9FFF275D1AC68C7BAADF6:8E836041FFC8FFF471C8AFAAC36EA99A" box="[1166,1283,1447,1473]" country="Peru" latitude="-13.3550005" longLatPrecision="21" longitude="-70.88944" municipality="Quincemil" name="Rio Araza" pageId="11" pageNumber="517" stateProvince="Cusco">Rio Araza</location>
|
||
,
|
||
<geoCoordinate id="EE68505DFFC8FFF47055AFA5C3E1A998" box="[1299,1420,1448,1475]" degrees="13" direction="south" minutes="21" orientation="latitude" pageId="11" pageNumber="517" precision="15" seconds="18" value="-13.3550005">13º21′18″S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="EE68505DFFC8FFF475D1AFC1C770A9BC" box="[151,285,1484,1511]" degrees="70" direction="west" minutes="53" orientation="longitude" pageId="11" pageNumber="517" precision="15" seconds="22" value="-70.88944">70º53′22″W</geoCoordinate>
|
||
,
|
||
<quantity id="4CA49B7FFFC8FFF47461AFC6C716A9BE" box="[295,379,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="517" unit="m" value="1000.0">
|
||
<elevation id="0071D1A9FFC8FFF47461AFC6C716A9BE" box="[295,379,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="517" unit="m" value="1000.0">1000 m</elevation>
|
||
</quantity>
|
||
” / “
|
||
<date id="FFE2105AFFC8FFF474E6AFC6C43CA9BE" box="[416,593,1483,1509]" pageId="11" pageNumber="517" value="2012-08-22" valueMax="2012-08-26" valueMin="2012-08-22">
|
||
<collectingDate id="EFA6E9B2FFC8FFF474E6AFC6C43CA9BE" box="[416,593,1483,1509]" pageId="11" pageNumber="517" value="2012-08-22" valueMax="2012-08-26" valueMin="2012-08-22">22–26.viii.2012</collectingDate>
|
||
</date>
|
||
, sweep,
|
||
<collectorName id="26A9534CFFC8FFF477EDAFC6C54AA9BE" box="[683,807,1483,1509]" pageId="11" pageNumber="517">J.A. Rafael</collectorName>
|
||
,
|
||
<collectorName id="26A9534CFFC8FFF47676AFC6C5B6A9BE" box="[816,987,1483,1509]" pageId="11" pageNumber="517">R.R. Cavichioli</collectorName>
|
||
” / “
|
||
<typeStatus id="54E78838FFC8FFF47146AFC6C205A9BE" box="[1024,1128,1483,1509]" pageId="11" pageNumber="517" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D5AFD13FFC8FFF47128AFC6C2EEA9BE" box="[1134,1155,1483,1509]" pageId="11" pageNumber="517" type="male">♂</specimenCount>
|
||
<taxonomicName id="4C5C4D19FFC8FFF471CEAFC6C708AA52" authority="Rafael & Marques" authorityName="Rafael & Marques" authorityYear="2021" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="chelicercus">
|
||
<emphasis id="B928EA88FFC8FFF471CEAFC6C3F1A9BE" box="[1160,1436,1483,1509]" italics="true" pageId="11" pageNumber="517">Macrostomus chelicercus</emphasis>
|
||
Rafael & Marques
|
||
</taxonomicName>
|
||
” (
|
||
<figureCitation id="13672A1FFFC8FFF47439AFE2C7B8AA52" box="[383,469,1519,1545]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[251,1325,202,1668]" captionTargetId="figure-17@12.[246,1340,201,1668]" captionTargetPageId="12" captionText="FIGURE 8. Macrostomus chelicercus sp. nov., holotype ♂. A. Habitus, lateral view before abdomen dissected; B. Mid and hind legs, lateral view; C. Apex of abdomen, lateral view; D. Tergite 8, dorsal view; E. Tergite and sternite 8, lateral view; F. Sternite 8, ventral view; G. Epandrium and cerci, lateral view; H. Epandrium detached, lateral view; I. Anterior cercus and subepandrial sclerite, anterior view; J. Posterior mesial process of anterior cercus, dorsal hypoproct lobe and apex of ventral projection of cercus, posterior view; K. Mesial process of anterior cercus, dorsal hypoproct lobe and cercal bridge, dorsal view; L. Hypandrium, phallus and ejaculatory apodeme, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: a ce, anterior cercus; ad l anterodorsal lobe of epandrium; c brd, cercal bridge; d hyprct l, dorsal hypoproctal lobe; ej apod, ejaculatory apodeme; hypd, hypandrium; m ce, median cercus; mpac, mesial process of anterior cercus; p ce, posterior cercus; pd l, posterodorsal lobe of epandrium; ph, phallus; sbepand scl, subepandrial sclerite; st, sternite; t, tergite, vpc, ventral process of cerci." figureDoi="http://doi.org/10.5281/zenodo.4921371" httpUri="https://zenodo.org/record/4921371/files/figure.png" pageId="11" pageNumber="517">Fig. 8N</figureCitation>
|
||
) (
|
||
<collectionCode id="ED4DAE5FFFC8FFF474ABAFE2C42AAA52" box="[493,583,1519,1545]" pageId="11" pageNumber="517">MUSM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="B928EA88FFC8FFF4771FAFE3C497AA53" bold="true" box="[601,762,1518,1544]" pageId="11" pageNumber="517">
|
||
<typeStatus id="54E78838FFC8FFF4771FAFE3C497AA53" box="[601,762,1518,1544]" pageId="11" pageNumber="517" type="paratype">PARATYPES</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D5AFD13FFC8FFF4764CAFE2C541AA52" box="[778,812,1519,1545]" pageId="11" pageNumber="517" type="male">1♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D5AFD13FFC8FFF47625AFE2C5E8AA52" box="[867,901,1519,1545]" pageId="11" pageNumber="517" type="female">1♀</specimenCount>
|
||
,
|
||
<materialsCitation id="3B343CC7FFC8FFF476D2AFE2C465AA0A" ID-GBIF-Occurrence="3312253304" collectingDate="2013-12-12" collectingDateMax="2013-12-16" collectingDateMin="2013-12-12" collectionCode="INPA, MUSM, CNC" collectorName=" A. L. Norrbom & B. D. Sutton" country="Peru" elevation="572" latitude="-1.89111" location="Estacion Biologica Villa Carmen" longLatPrecision="1" longitude="-71.41046" municipality="Quincemil" pageId="11" pageNumber="517" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Cusco" typeStatus="paratype">
|
||
same data as holotype (
|
||
<specimenCount id="9D5AFD13FFC8FFF471DEAFE2C2D4AA52" box="[1176,1209,1519,1545]" pageId="11" pageNumber="517" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED4DAE5FFFC8FFF47187AFE2C36CAA52" box="[1217,1281,1519,1545]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="11" pageNumber="517">INPA</collectionCode>
|
||
,
|
||
<specimenCount id="9D5AFD13FFC8FFF4704BAFE2C35DAA52" box="[1293,1328,1519,1545]" pageId="11" pageNumber="517" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED4DAE5FFFC8FFF47071AFE2C3FDAA52" box="[1335,1424,1519,1545]" pageId="11" pageNumber="517">MUSM</collectionCode>
|
||
);
|
||
<collectingRegion id="4998F878FFC8FFF475D1AC1EC6B6AA76" box="[151,219,1555,1581]" country="Peru" name="Cusco" pageId="11" pageNumber="517">Cusco</collectingRegion>
|
||
<location id="8E836041FFC8FFF475A3AC1EC43BAA76" LSID="urn:lsid:plazi:treatment:03F5878CFFC9FFF275D1AC68C7BAADF6:8E836041FFC8FFF475A3AC1EC43BAA76" box="[229,598,1555,1581]" latitude="-1.89111" longLatPrecision="1" longitude="-71.41046" name="Estacion Biologica Villa Carmen" pageId="11" pageNumber="517" stateProvince="Cusco">Estacion Biologica Villa Carmen</location>
|
||
,
|
||
<collectorName id="26A9534CFFC8FFF47722AC1EC4A3AA76" box="[612,718,1555,1581]" pageId="11" pageNumber="517">Oroya, S.</collectorName>
|
||
side,
|
||
<geoCoordinate id="EE68505DFFC8FFF47652AC1EC5E9AA76" box="[788,900,1555,1581]" degrees="1.89111" direction="south" orientation="latitude" pageId="11" pageNumber="517" precision="1" value="-1.89111">1.89111°S</geoCoordinate>
|
||
<geoCoordinate id="EE68505DFFC8FFF476C8AC1EC274AA76" box="[910,1049,1555,1581]" degrees="71.41046" direction="west" orientation="longitude" pageId="11" pageNumber="517" precision="1" value="-71.41046">71.41046°W</geoCoordinate>
|
||
,
|
||
<quantity id="4CA49B7FFFC8FFF47161AC1FC21FAA76" box="[1063,1138,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="5.72" pageId="11" pageNumber="517" unit="m" value="572.0">
|
||
<elevation id="0071D1A9FFC8FFF47161AC1FC21FAA76" box="[1063,1138,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="5.72" pageId="11" pageNumber="517" unit="m" value="572.0">572 m</elevation>
|
||
</quantity>
|
||
, MT,
|
||
<date id="FFE2105AFFC8FFF471F1AC1EC330AA76" box="[1207,1373,1555,1581]" pageId="11" pageNumber="517" value="2013-12-12" valueMax="2013-12-16" valueMin="2013-12-12">
|
||
<collectingDate id="EFA6E9B2FFC8FFF471F1AC1EC330AA76" box="[1207,1373,1555,1581]" pageId="11" pageNumber="517" value="2013-12-12" valueMax="2013-12-16" valueMin="2013-12-12">12–16.xii.2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A9534CFFC8FFF4702CAC1EC693AA0A" pageId="11" pageNumber="517">A.L. Norrbom</collectorName>
|
||
,
|
||
<collectorName id="26A9534CFFC8FFF4744FAC3AC7E7AA0A" box="[265,394,1591,1617]" pageId="11" pageNumber="517">B.D. Sutton</collectorName>
|
||
(
|
||
<specimenCount id="9D5AFD13FFC8FFF474DCAC3AC7D1AA0A" box="[410,444,1591,1617]" pageId="11" pageNumber="517" type="male">1♂</specimenCount>
|
||
<collectionCode id="ED4DAE5FFFC8FFF47485AC3AC46DAA0A" box="[451,512,1591,1617]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="11" pageNumber="517">CNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE3369AFFC8FFF47581AC56C695AAC2" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="517">
|
||
<emphasis id="B928EA88FFC8FFF47581AC56C7C1AA2F" bold="true" box="[199,428,1626,1653]" pageId="11" pageNumber="517">
|
||
<typeStatus id="54E78838FFC8FFF47581AC56C75CAA2E" box="[199,305,1627,1653]" pageId="11" pageNumber="517" type="holotype">Holotype</typeStatus>
|
||
condition.
|
||
</emphasis>
|
||
Pinned, glued on small card, in good condition; abdomen dissected, stored in microvial with glycerin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466511FFC8FFF47581ACAEC6A9AABA" pageId="11" pageNumber="517" type="etymology">
|
||
<paragraph id="8BE3369AFFC8FFF47581ACAEC6A9AABA" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="517">
|
||
<emphasis id="B928EA88FFC8FFF47581ACAEC723AAE6" bold="true" box="[199,334,1699,1725]" pageId="11" pageNumber="517">Etymology.</emphasis>
|
||
From Greek
|
||
<emphasis id="B928EA88FFC8FFF474A1ACAEC44FAAE6" box="[487,546,1699,1725]" italics="true" pageId="11" pageNumber="517">chele</emphasis>
|
||
= claw and
|
||
<emphasis id="B928EA88FFC8FFF477F6ACAEC495AAE7" box="[688,760,1699,1724]" italics="true" pageId="11" pageNumber="517">cercus</emphasis>
|
||
= cercus, refers to the cheliform shape of the posterior cercus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3466511FFC8FFF27581ACE6C7BAADF6" lastPageId="13" lastPageNumber="519" pageId="11" pageNumber="517" type="discussion">
|
||
<paragraph id="8BE3369AFFC8FFF27581ACE6C7BAADF6" blockId="11.[151,1437,151,2013]" lastBlockId="13.[151,1437,151,429]" lastPageId="13" lastPageNumber="519" pageId="11" pageNumber="517">
|
||
<emphasis id="B928EA88FFC8FFF47581ACE6C756AB5E" bold="true" box="[199,315,1771,1797]" pageId="11" pageNumber="517">Remarks.</emphasis>
|
||
<taxonomicName id="4C5C4D19FFC8FFF47405ACE6C40DAB5E" authority="Rafael & Marques, 2021" authorityName="Rafael & Marques" authorityYear="2021" box="[323,608,1771,1797]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="chelicercus" status="sp. nov.">
|
||
<emphasis id="B928EA88FFC8FFF47405ACE6C40DAB5E" box="[323,608,1771,1797]" italics="true" pageId="11" pageNumber="517">Macrostomus chelicercus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA88FFC8FFF4772EACE6C4AFAB5E" bold="true" box="[616,706,1771,1797]" pageId="11" pageNumber="517">
|
||
<taxonomicNameLabel id="A21B57F3FFC8FFF4772EACE6C4AFAB5E" box="[616,706,1771,1797]" pageId="11" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is best placed in the
|
||
<taxonomicName id="4C5C4D19FFC8FFF476F4ACE6C20AAB5F" authorityName="Rafael & Cumming" authorityYear="2015" box="[946,1127,1771,1796]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="amazonensis">
|
||
<emphasis id="B928EA88FFC8FFF476F4ACE6C20AAB5F" box="[946,1127,1771,1796]" italics="true" pageId="11" pageNumber="517">M. amazonensis</emphasis>
|
||
</taxonomicName>
|
||
species-group as described by
|
||
<bibRefCitation id="EFCD4B6BFFC8FFF475FFAD02C78BAB72" author="Rafael, J. A. & Cumming, J. M." box="[185,486,1806,1833]" pageId="11" pageNumber="517" pagination="109 - 121" refId="ref17462" refString="Rafael, J. A. & Cumming, J. M. (2015) Revision of the genus Macrostomus Wiedemann (Diptera, Empididae, Empidinae). IV. The amazonensis species-group. Zootaxa, 3947 (1), 109 - 121. https: // doi. org / 10.11646 / zootaxa. 3947.1.7" type="journal article" year="2015">Rafael & Cumming (2015)</bibRefCitation>
|
||
. In the key presented by
|
||
<bibRefCitation id="EFCD4B6BFFC8FFF477B2AD02C272AB72" author="Rafael, J. A. & Cumming, J. M." box="[756,1055,1806,1833]" pageId="11" pageNumber="517" pagination="109 - 121" refId="ref17462" refString="Rafael, J. A. & Cumming, J. M. (2015) Revision of the genus Macrostomus Wiedemann (Diptera, Empididae, Empidinae). IV. The amazonensis species-group. Zootaxa, 3947 (1), 109 - 121. https: // doi. org / 10.11646 / zootaxa. 3947.1.7" type="journal article" year="2015">Rafael & Cumming (2015)</bibRefCitation>
|
||
it runs to couplet 2 of
|
||
<taxonomicName id="4C5C4D19FFC8FFF47055AD02C7D5AB16" authority="Rafael & Cumming, 2015" authorityName="Rafael & Cumming" authorityYear="2015" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="acreanus">
|
||
<emphasis id="B928EA88FFC8FFF47055AD02C3F0AB73" box="[1299,1437,1807,1832]" italics="true" pageId="11" pageNumber="517">M. acreanus</emphasis>
|
||
<bibRefCitation id="EFCD4B6BFFC8FFF475D1AD3EC7D5AB16" author="Rafael, J. A. & Cumming, J. M." box="[151,440,1842,1869]" pageId="11" pageNumber="517" pagination="109 - 121" refId="ref17462" refString="Rafael, J. A. & Cumming, J. M. (2015) Revision of the genus Macrostomus Wiedemann (Diptera, Empididae, Empidinae). IV. The amazonensis species-group. Zootaxa, 3947 (1), 109 - 121. https: // doi. org / 10.11646 / zootaxa. 3947.1.7" type="journal article" year="2015">Rafael & Cumming, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
by having the mesopleuron yellow. Both species have a short tergite 8 with shallow basal sinus.
|
||
<taxonomicName id="4C5C4D19FFC8FFF4759BAD5AC795AB2A" authority="Rafael & Marques, 2021" authorityName="Rafael & Marques" authorityYear="2021" box="[221,504,1879,1905]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="chelicercus" status="sp. nov.">
|
||
<emphasis id="B928EA88FFC8FFF4759BAD5AC795AB2A" box="[221,504,1879,1905]" italics="true" pageId="11" pageNumber="517">Macrostomus chelicercus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA88FFC8FFF474B8AD5AC438AB2A" bold="true" box="[510,597,1879,1905]" pageId="11" pageNumber="517">
|
||
<taxonomicNameLabel id="A21B57F3FFC8FFF474B8AD5AC438AB2A" box="[510,597,1879,1905]" pageId="11" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs by the cercal bridge U-shaped (versus dome-shaped in
|
||
<taxonomicName id="4C5C4D19FFC8FFF47045AD5AC3E1AB2B" authorityName="Rafael & Cumming" authorityYear="2015" box="[1283,1420,1879,1904]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="acreanus">
|
||
<emphasis id="B928EA88FFC8FFF47045AD5AC3E1AB2B" box="[1283,1420,1879,1904]" italics="true" pageId="11" pageNumber="517">M. acreanus</emphasis>
|
||
</taxonomicName>
|
||
), anterior cercus subtriangular (versus longitudinally elongate), posterior cercus inward and forward directed with apex bifid (versus vertically elongate with apex simple). Nevertheless,
|
||
<taxonomicName id="4C5C4D19FFC8FFF476F8AD92C2B6ABE2" authority="Rafael & Marques, 2021" authorityName="Rafael & Marques" authorityYear="2021" box="[958,1243,1951,1977]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="chelicercus" status="sp. nov.">
|
||
<emphasis id="B928EA88FFC8FFF476F8AD92C2B6ABE2" box="[958,1243,1951,1977]" italics="true" pageId="11" pageNumber="517">Macrostomus chelicercus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA88FFC8FFF471A2AD92C352ABE2" bold="true" box="[1252,1343,1951,1977]" pageId="11" pageNumber="517">
|
||
<taxonomicNameLabel id="A21B57F3FFC8FFF471A2AD92C352ABE2" box="[1252,1343,1951,1977]" pageId="11" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appears to be more related with
|
||
<taxonomicName id="4C5C4D19FFC8FFF474E4ADCEC552AB86" authority="Rafael & Cumming, 2015" authorityName="Rafael & Cumming" authorityYear="2015" box="[418,831,1986,2013]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="517" phylum="Arthropoda" rank="species" species="xavieri">
|
||
<emphasis id="B928EA88FFC8FFF474E4ADCEC478AB87" box="[418,533,1987,2012]" italics="true" pageId="11" pageNumber="517">M. xavieri</emphasis>
|
||
<bibRefCitation id="EFCD4B6BFFC8FFF4775AADCEC552AB86" author="Rafael, J. A. & Cumming, J. M." box="[540,831,1986,2013]" pageId="11" pageNumber="517" pagination="109 - 121" refId="ref17462" refString="Rafael, J. A. & Cumming, J. M. (2015) Revision of the genus Macrostomus Wiedemann (Diptera, Empididae, Empidinae). IV. The amazonensis species-group. Zootaxa, 3947 (1), 109 - 121. https: // doi. org / 10.11646 / zootaxa. 3947.1.7" type="journal article" year="2015">Rafael & Cumming, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
by the short tergite 8 with shallow basal sinus, mesial process of anterior cercus small and posterior cercus inwardly curved.
|
||
<taxonomicName id="4C5C4D19FFCEFFF276EBAA9AC2A4ACEA" authority="Rafael & Marques, 2021" authorityName="Rafael & Marques" authorityYear="2021" box="[941,1225,151,177]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="519" phylum="Arthropoda" rank="species" species="chelicercus" status="sp. nov.">
|
||
<emphasis id="B928EA88FFCEFFF276EBAA9AC2A4ACEA" box="[941,1225,151,177]" italics="true" pageId="13" pageNumber="519">Macrostomus chelicercus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA88FFCEFFF27196AA9AC347ACEA" bold="true" box="[1232,1322,151,177]" pageId="13" pageNumber="519">
|
||
<taxonomicNameLabel id="A21B57F3FFCEFFF27196AA9AC347ACEA" box="[1232,1322,151,177]" pageId="13" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs by cercal bridge U-shaped (versus somewhat bilobate in
|
||
<taxonomicName id="4C5C4D19FFCEFFF277B8AAB6C51EAC8F" authorityName="Rafael & Cumming" authorityYear="2015" box="[766,883,187,212]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="519" phylum="Arthropoda" rank="species" species="xavieri">
|
||
<emphasis id="B928EA88FFCEFFF277B8AAB6C51EAC8F" box="[766,883,187,212]" italics="true" pageId="13" pageNumber="519">M. xavieri</emphasis>
|
||
</taxonomicName>
|
||
), anterior cercus subtriangular (versus vertically elongate), posterior cercus inwardly and forwardly directed with apex bifid (versus inwardly curved with apex simple). Additionally,
|
||
<taxonomicName id="4C5C4D19FFCEFFF274D5AB0EC45AAD46" authority="Rafael & Marques, 2021" authorityName="Rafael & Marques" authorityYear="2021" box="[403,567,259,285]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="519" phylum="Arthropoda" rank="species" species="chelicercus" status="sp. nov.">
|
||
<emphasis id="B928EA88FFCEFFF274D5AB0EC45AAD46" box="[403,567,259,285]" italics="true" pageId="13" pageNumber="519">M. chelicercus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA88FFCEFFF27779AB0EC4F4AD46" bold="true" box="[575,665,259,285]" pageId="13" pageNumber="519">
|
||
<taxonomicNameLabel id="A21B57F3FFCEFFF27779AB0EC4F4AD46" box="[575,665,259,285]" pageId="13" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from both species by the distinctly crooked hind femur, with denser, short, stouter dorsal and ventral setae at curved area and by the slightly curved mid and hind femora and tibiae. In CNC, there is an additional new species which keys to
|
||
<taxonomicName id="4C5C4D19FFCEFFF2762AAB46C27DAD3E" authority="Rafael & Marques, 2021" authorityName="Rafael & Marques" authorityYear="2021" box="[876,1040,331,357]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="519" phylum="Arthropoda" rank="species" species="chelicercus" status="sp. nov.">
|
||
<emphasis id="B928EA88FFCEFFF2762AAB46C27DAD3E" box="[876,1040,331,357]" italics="true" pageId="13" pageNumber="519">M. chelicercus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928EA88FFCEFFF27151AB46C21CAD3E" bold="true" box="[1047,1137,331,357]" pageId="13" pageNumber="519">
|
||
<taxonomicNameLabel id="A21B57F3FFCEFFF27151AB46C21CAD3E" box="[1047,1137,331,357]" pageId="13" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(B.J. Sinclair pers. comm. 2020). The specimens are from
|
||
<collectingCountry id="F34B760AFFCEFFF27741AB62C450ADD2" box="[519,573,367,393]" name="Peru" pageId="13" pageNumber="519">Peru</collectingCountry>
|
||
, Manu Wildlife Centre, Bamboo Trial,
|
||
<date id="FFE2105AFFCEFFF27144AB62C218ADD2" box="[1026,1141,367,393]" pageId="13" pageNumber="519" value="2006-10-30">30.x.2006</date>
|
||
,
|
||
<geoCoordinate id="EE68505DFFCEFFF271C5AB7DC369ADD0" box="[1155,1284,368,395]" degrees="12" direction="south" minutes="17" orientation="latitude" pageId="13" pageNumber="519" precision="15" seconds="08" value="-12.285556">12°17′08″S</geoCoordinate>
|
||
<geoCoordinate id="EE68505DFFCEFFF2704AAB7DC3F5ADD0" box="[1292,1432,368,395]" degrees="70" direction="west" minutes="54" orientation="longitude" pageId="13" pageNumber="519" precision="15" seconds="14" value="-70.90389">70°54′14″W</geoCoordinate>
|
||
, J.H. Skevington (
|
||
<specimenCount id="9D5AFD13FFCEFFF2741DAB9EC7ECADF6" box="[347,385,403,429]" pageId="13" pageNumber="519" type="male">4♂</specimenCount>
|
||
, CNC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |