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<document ID-DOI="10.1080/00222933.2022.2115950" ID-ISSN="1464-5262" ID-Zenodo-Dep="7156474" approvalRequired="9" approvalRequired_for_textStreams="9" checkinTime="1665136984522" checkinUser="plazi" docAuthor="Medeiros, Rone A. F., Seidel, Matthias &amp; Grossi, Paschoal C." docDate="2022" docId="03F09661FFFC806EFE0BE7267A265192" docLanguage="en" docName="JNATHIST.56.29-32.1315-1364.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 56 (29 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2022.2115950" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Byrsopolis Burmeister 1844" docType="treatment" docVersion="2" lastPageNumber="1320" masterDocId="FFC9EE19FFFF806BFFD0E116782B521F" masterDocTitle="Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay" masterLastPageNumber="1364" masterPageNumber="1315" pageNumber="1318" updateTime="1665137547684" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Medeiros, Rone A. F.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Seidel, Matthias</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Grossi, Paschoal C.</mods:namePart>
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<mods:title>Journal of Natural History</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="series">
<mods:title>J. Nat. Hist.</mods:title>
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<mods:number>2022-09-22</mods:number>
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<mods:detail type="volume">
<mods:number>56</mods:number>
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<mods:detail type="issue">
<mods:number>29 - 32</mods:number>
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<mods:start>1315</mods:start>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2022.2115950</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<subSubSection box="[475,840,1584,1611]" pageId="3" pageNumber="1318" type="nomenclature">
<paragraph blockId="3.[475,840,1584,1611]" box="[475,840,1584,1611]" pageId="3" pageNumber="1318">
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<taxonomicName authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" box="[475,840,1584,1611]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1318" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[475,840,1584,1611]" pageId="3" pageNumber="1318">
<emphasis bold="true" box="[475,607,1584,1611]" italics="true" pageId="3" pageNumber="1318">Byrsopolis</emphasis>
<bibRefCitation author="Burmeister HCC" box="[615,840,1584,1611]" pageId="3" pageNumber="1318" refId="ref25824" refString="Burmeister HCC. 1844. Besondere Entomologie, Fortsetzung. Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. xii. 588." type="book" year="1844">Burmeister, 1844</bibRefCitation>
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<subSubSection pageId="3" pageNumber="1318" type="type_taxon">
<paragraph blockId="3.[160,944,1631,1656]" box="[160,944,1631,1656]" pageId="3" pageNumber="1318">
<emphasis bold="true" box="[160,764,1631,1656]" pageId="3" pageNumber="1318">
<typeStatus box="[160,219,1631,1656]" pageId="3" pageNumber="1318">Type</typeStatus>
species:
<emphasis bold="true" box="[327,551,1631,1656]" italics="true" pageId="3" pageNumber="1318">Byrsopoliscastanea</emphasis>
<bibRefCitation author="Burmeister HCC" box="[559,764,1631,1656]" pageId="3" pageNumber="1318" refId="ref25824" refString="Burmeister HCC. 1844. Besondere Entomologie, Fortsetzung. Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. xii. 588." type="book" year="1844">Burmeister, 1844</bibRefCitation>
</emphasis>
[by monotypy].
</paragraph>
<paragraph blockId="3.[160,379,1702,1728]" box="[160,379,1702,1728]" pageId="3" pageNumber="1318">
<emphasis bold="true" box="[160,252,1702,1728]" italics="true" pageId="3" pageNumber="1318">Gender.</emphasis>
Feminine.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="1319" type="reference_group">
<paragraph blockId="4.[160,1157,142,757]" pageId="4" pageNumber="1319">
<treatmentCitationGroup pageId="4" pageNumber="1319">
<treatmentCitation author="Burmeister HCC" box="[192,554,142,167]" page="425" pageId="4" pageNumber="1319" year="1844">
<taxonomicName authority="Burmeister, 1844: 425" authorityName="Burmeister" authorityPageNumber="425" authorityYear="1844" box="[192,554,142,167]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1319" phylum="Arthropoda" rank="genus">
<emphasis box="[192,300,142,167]" italics="true" pageId="4" pageNumber="1319">Byrsopolis</emphasis>
<bibRefCitation author="Burmeister HCC" box="[309,554,142,167]" pageId="4" pageNumber="1319" refId="ref25824" refString="Burmeister HCC. 1844. Besondere Entomologie, Fortsetzung. Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. xii. 588." type="book" year="1844">Burmeister, 1844: 425</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation author="Blanchard E" box="[811,1041,142,167]" page="219" pageId="4" pageNumber="1319" year="1851">
<bibRefCitation author="Blanchard E" box="[811,1041,142,167]" pageId="4" pageNumber="1319" pagination="241" refId="ref25750" refString="Blanchard E. 1851. Gide et Baudry. In: Milne-Edwards H, Blanchard E, Lucas H, editors. Catalogue des collections Entomologiques du Museum d'Histoire naturelle de Paris, Classe des Insectes. Ordre des Coleopteres. Paris, France: Museum d'histoire naturelle de Paris. (1850) 1. (1850) 1128, (1851) 129240; p. 129240; p. 241." type="journal article" year="1851">Blanchard 1851: 219</bibRefCitation>
</treatmentCitation>
(citation);
<treatmentCitation author="Burmeister HCC" box="[160,409,177,202]" page="523" pageId="4" pageNumber="1319" year="1855">
<bibRefCitation author="Burmeister HCC" box="[160,409,177,202]" pageId="4" pageNumber="1319" pagination="570" refId="ref25867" refString="Burmeister HCC. 1855. Besondere Entomologie, Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. 570." type="book chapter" year="1855">Burmeister 1855: 523</bibRefCitation>
</treatmentCitation>
(female diagnosis);
<treatmentCitation author="Lacordaire JT" box="[650,894,177,202]" page="364" pageId="4" pageNumber="1319" year="1856">
<bibRefCitation author="Lacordaire JT" box="[650,894,177,202]" pageId="4" pageNumber="1319" pagination="594" refId="ref26568" refString="Lacordaire JT. 1856. Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme contenant les familles des pectinicornes et lamellicornes. Paris: Librairie Encyclopedique de Roret; p. 594." type="book chapter" year="1856">Lacordaire 1856: 364</bibRefCitation>
</treatmentCitation>
(redescription, key to genera);
<treatmentCitation author="Castelnau FL &amp; Lucas MH" box="[265,636,212,237]" page="130" pageId="4" pageNumber="1319" year="1857">
<bibRefCitation author="Castelnau FL &amp; Lucas MH" box="[265,636,212,237]" pageId="4" pageNumber="1319" pagination="473" refId="ref26028" refString="Castelnau FL, Lucas MH. 1857. Animaux nouveaux ou rares recueilles pendant l'Expedition dans les parties centrales de l'Amerique du Sud, de Rio de Janeiro a Lima et de Lima au Para sous la direction du Comte Francis de Castelnau. Chez P. Bertrand. Paris: Librareie-Editeur; p. 473." type="book chapter" year="1857">Castelnau and Lucas 1857: 130</bibRefCitation>
</treatmentCitation>
(distribution, illustrations);
<treatmentCitation author="Gemminger M &amp; Harold B" page="1226" pageId="4" pageNumber="1319" year="1869">
<bibRefCitation author="Gemminger M &amp; Harold B" pageId="4" pageNumber="1319" pagination="1240" refId="ref26271" refString="Gemminger M, Harold B. 1869. Catalogus Coleopterum: Scarabaeidae, Tom. IV. 4: 1240." type="journal article" year="1869">Gemminger and Harold 1869: 1226</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation author="Bates HW" box="[505,685,246,271]" page="290" pageId="4" pageNumber="1319" year="1888">
<bibRefCitation author="Bates HW" box="[505,685,246,271]" pageId="4" pageNumber="1319" pagination="456" refId="ref25676" refString="Bates HW. 1888. Biologia Centrali-Americana, Insecta, Coleoptera, Melolonthidae, Rutelidae, Dynastidae. Biologia Centrali-Americana. 2 (2): 456." type="journal article" year="1888">Bates 1888: 290</bibRefCitation>
</treatmentCitation>
(citation);
<treatmentCitation author="Heyne A &amp; Taschenberg O" page="88" pageId="4" pageNumber="1319" year="1908">
<bibRefCitation author="Heyne A &amp; Taschenberg O" pageId="4" pageNumber="1319" pagination="39" refId="ref26360" refString="Heyne A, Taschenberg O. 1908. Die Exotischen Kafer in Wort und Bild. Leipzig. vii + 262: 39." type="journal article" year="1908">Heyne and Taschenberg 1908: 88</bibRefCitation>
</treatmentCitation>
(citation), 356 (illustration);
<treatmentCitation author="Ohaus F" box="[510,701,281,306]" page="251" pageId="4" pageNumber="1319" year="1908">
<bibRefCitation author="Ohaus F" box="[510,701,281,306]" pageId="4" pageNumber="1319" pagination="239 - 262" refId="ref26967" refString="Ohaus F. 1908. Die Ruteliden meiner Sammelreisen in Sudamerika. (Col.). Deutsche entomologische Zeitschrift. 1908: 239 - 262." type="journal article" year="1908">Ohaus 1908: 251</bibRefCitation>
</treatmentCitation>
(citation); 1912: 312 (citation); 1915: 75 (citation);
<treatmentCitation author="Casey TL" box="[272,426,316,341]" page="1" pageId="4" pageNumber="1319" year="1915">
<bibRefCitation author="Casey TL" box="[272,426,316,341]" pageId="4" pageNumber="1319" pagination="1 - 394" refId="ref26000" refString="Casey TL. 1915. A revision of the American species of Rutelinae, Dynastinae, and Cetoniinae. Mem Coleoptera. 6: 1 - 394." type="journal article" year="1915">Casey 1915: 1</bibRefCitation>
</treatmentCitation>
(note);
<treatmentCitation author="Ohaus F" box="[513,686,316,341]" page="75" pageId="4" pageNumber="1319" year="1925">
<bibRefCitation author="Ohaus F" box="[513,686,316,341]" pageId="4" pageNumber="1319" pagination="75 - 83" refId="ref27086" refString="Ohaus F. 1925. XXI. Beitrag zur Kenntnis der Ruteliden (Col. Lamell.). Berlin, Germany: Deutsche entomologische Zeitschrift; pp. 75 - 83." type="book chapter" year="1925">Ohaus 1925: 75</bibRefCitation>
</treatmentCitation>
(citation); 1926: 225226 (citation);
<treatmentCitation author="Hardy A" page="235" pageId="4" pageNumber="1319" year="1971">
<bibRefCitation author="Hardy A" pageId="4" pageNumber="1319" pagination="235 - 242" refId="ref26328" refString="Hardy A. 1971. The North American Areodina with a description of a new genus from California. Pan- Pac Entomol. 47 (3): 235 - 242." type="journal article" year="1971">Hardy 1971: 235</bibRefCitation>
</treatmentCitation>
(status revalidation);
<treatmentCitation author="Machatschke JW" box="[502,733,350,375]" page="2" pageId="4" pageNumber="1319" year="1972">
<bibRefCitation author="Machatschke JW" box="[502,733,350,375]" pageId="4" pageNumber="1319" pagination="361" refId="ref26699" refString="Machatschke JW. 1972. Scarabaeoidea: Melolonthinae, Rutelinae. Coleopterorum Catalogus. Supplementa. 66: 361." type="journal article" year="1972">Machatschke 1972: 2</bibRefCitation>
</treatmentCitation>
(catalogue, distribution);
<treatmentCitation author="Moron MA" page="19" pageId="4" pageNumber="1319" year="1976">
<bibRefCitation author="Moron MA" pageId="4" pageNumber="1319" pagination="19 - 23" refId="ref26896" refString="Moron MA. 1976. Descripcion de una nueva especie Mexicana del genero Parabyrsopolis Ohaus, 1915. Anales del Instituto de Biologia de la Universidad Nacional Autonoma de Mexico. 47: 19 - 23." type="journal article" year="1976">Morón 1976: 19</bibRefCitation>
</treatmentCitation>
(citation);
<treatmentCitation author="Ratcliffe BC &amp; Jameson ML" box="[308,669,385,410]" page="135" pageId="4" pageNumber="1319" year="1989">
<bibRefCitation author="Ratcliffe BC &amp; Jameson ML" box="[308,669,385,410]" pageId="4" pageNumber="1319" pagination="135 - 144" refId="ref27184" refString="Ratcliffe BC, Jameson ML. 1989. A synopsis of the genus Areoda (Coleoptera: Scarabaeidae: Rutelinae). Coleopterists Bull. 43 (2): 135 - 144." type="journal article" year="1989">Ratcliffe and Jameson 1989: 135</bibRefCitation>
</treatmentCitation>
(citation);
<treatmentCitation author="Jameson ML" box="[789,1004,385,410]" page="377" pageId="4" pageNumber="1319" year="1990">
<bibRefCitation author="Jameson ML" box="[789,1004,385,410]" pageId="4" pageNumber="1319" pagination="377 - 422" refId="ref26416" refString="Jameson ML. 1990. Revision, phylogeny and biogeography of the genera Parabyrsopolis Ohaus and Viridimicus, new genus (Coleoptera: Scarabaeidae: Rutelinae). Coleopterists Bull. 44 (4): 377 - 422." type="journal article" year="1990">Jameson 1990: 377</bibRefCitation>
</treatmentCitation>
(citation, key to genera); 1998: 5 (citation);
<treatmentCitation author="Krajcik M" box="[488,662,420,445]" page="58" pageId="4" pageNumber="1319" year="2008">
<bibRefCitation author="Krajcik M" box="[488,662,420,445]" pageId="4" pageNumber="1319" pagination="142" refId="ref26488" refString="Krajcik M. 2008. Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). 2 nd ed. Plzen, Czech Republic: Milan Krajcik; p. 142." type="book chapter" year="2008">Krajcik 2008: 58</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation author="Smith" box="[793,974,420,445]" page="73" pageId="4" pageNumber="1319" year="2009">
<bibRefCitation author="Smith" box="[793,936,420,445]" pageId="4" pageNumber="1319" refId="ref27250" refString="Smith. 2009. Checklist and nomenclatural authority file of the Scarabaeoidea of the Neartic Realm. http: // unsm-ento. unl. edu / SSSA / Nearctic-Scarabs 4. pdf." type="url" year="2009">Smith (2009)</bibRefCitation>
: 73
</treatmentCitation>
(citation);
<bibRefCitation author="Soula M" pageId="4" pageNumber="1319" pagination="66" refId="ref27367" refString="Soula M. 2010 a. Les Coleopteres du Nouveau Monde. Volume 4: Rutelini 4. Revision des 4. Une revision des genres Catoclastus, Homonyx. Retour sur quelques especes ou groupe d'especes de Pelidnota. Addenda 2010. (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. Saintry: AECFT; p. 66." type="book chapter" year="2010">Soula 2010a</bibRefCitation>
(citation): 44; 2010b: 54 (key to species, illustration);
<treatmentCitation author="Grossi PC &amp; Vaz-de-Mello FZ" page="221" pageId="4" pageNumber="1319" year="2015">
<bibRefCitation author="Grossi PC &amp; Vaz-de-Mello FZ" pageId="4" pageNumber="1319" pagination="221 - 226" refId="ref26292" refString="Grossi PC, Vaz-de-Mello FZ. 2015. Moronius miguelangeli new genus and new species of Areodina from western Brazil (Melolonthidae, Rutelinae, Rutelini). Dugesiana. 22: 221 - 226." type="journal article" year="2015">Grossi and Vaz-de-Mello 2015: 221</bibRefCitation>
</treatmentCitation>
(biology, key to genera);
<treatmentCitation author="Moron MA &amp; Nogueira G" box="[497,828,489,514]" page="7" pageId="4" pageNumber="1319" year="2016">
<bibRefCitation author="Moron MA &amp; Nogueira G" box="[497,828,489,514]" pageId="4" pageNumber="1319" pagination="7 - 22" refId="ref26933" refString="Moron MA, Nogueira G. 2016. Actualizacion del genero Parabyrsopolis Ohaus (Coleoptera: Melolonthidae: Rutelinae). Boletin de la Sociedad Entomologica Aragonesa. 59: 7 - 22." type="journal article" year="2016">Morón and Nogueira 2016: 7</bibRefCitation>
</treatmentCitation>
(citation);
<treatmentCitation author="Ferreira ADS" box="[951,1155,489,514]" page="112" pageId="4" pageNumber="1319" year="2016">
<bibRefCitation author="Ferreira ADS" box="[951,1155,489,514]" pageId="4" pageNumber="1319" refId="ref26223" refString="Ferreira ADS. 2016. Besouros rutelineos: beleza escondida no Semiarido, 111 - 119. In: Bravo F, Calor AR, editors. Conhecendo os artropodes do Semiario. 1 st ed. Sao Paulo: Metis Producao Editorial; p. 192." type="journal volume" year="2016">Ferreira 2016: 112</bibRefCitation>
</treatmentCitation>
(illustration);
<treatmentCitation author="Moore MR &amp; Jameson ML &amp; Garner BH &amp; Audibert C &amp; Smith ABT &amp; Seidel M" box="[317,578,524,549]" page="49" pageId="4" pageNumber="1319" year="2017">
<bibRefCitation author="Moore MR &amp; Jameson ML &amp; Garner BH &amp; Audibert C &amp; Smith ABT &amp; Seidel M" box="[317,578,524,549]" pageId="4" pageNumber="1319" pagination="1 - 349" refId="ref26854" refString="Moore MR, Jameson ML, Garner BH, Audibert C, Smith ABT, Seidel M. 2017. Synopsis of the pelidnotine scarabs and annotated catalog of the species and subspecies. ZooKeys. 666: 1 - 349." type="journal article" year="2017">Moore et al. 2017: 49</bibRefCitation>
</treatmentCitation>
(citation);
<treatmentCitation author="Ferreira ADS &amp; Almeida LM &amp; Bravo F &amp; Grossi PC" box="[711,973,524,549]" page="4" pageId="4" pageNumber="1319" year="2018">
<bibRefCitation author="Ferreira ADS &amp; Almeida LM &amp; Bravo F &amp; Grossi PC" box="[711,973,524,549]" pageId="4" pageNumber="1319" pagination="20170476" refId="ref26150" refString="Ferreira ADS, Almeida LM, Bravo F, Grossi PC. 2018. A checklist of Rutelinae MacLeay, 1819 (Coleoptera, Melolonthidae) of Bahia, Brazil. Biota Neotrop. 18 (2): e 20170476. www. scielo. br / scielo. php? script = sci _ arttext &amp; pid = S 1676 - 06032018000200301 &amp; lng = en &amp; nrm = iso." type="journal article" year="2018">Ferreira et al. 2018: 4</bibRefCitation>
</treatmentCitation>
, 89 (checklist, distribution);
<treatmentCitation author="Carvalho TG &amp; Grossi PC" box="[322,692,558,583]" page="372" pageId="4" pageNumber="1319" year="2018">
<bibRefCitation author="Carvalho TG &amp; Grossi PC" box="[322,692,558,583]" pageId="4" pageNumber="1319" pagination="369 - 372" refId="ref25906" refString="Carvalho TG, Grossi PC. 2018. Description of a second species of the genus Moronius Grossi &amp; Vaz-deMello, 2015 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini). Zootaxa. 4434 (2): 369 - 372. doi: 10. 11646 / zootaxa. 4434.2.7." type="journal article" year="2018">Carvalho and Grossi 2018: 372</bibRefCitation>
</treatmentCitation>
(checklist, distribution);
<treatmentCitation author="Hielkema AJ &amp; Hielkema MA" page="126" pageId="4" pageNumber="1319" year="2019">
<bibRefCitation author="Hielkema AJ &amp; Hielkema MA" pageId="4" pageNumber="1319" pagination="1 - 306" refId="ref26384" refString="Hielkema AJ, Hielkema MA. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi. 732: 1 - 306." type="journal article" year="2019">Hielkema and Hielkema 2019: 126</bibRefCitation>
</treatmentCitation>
(checklist, distribution);
<treatmentCitation author="Menis FT &amp; Rodrigues SR" box="[671,1012,593,618]" page="3" pageId="4" pageNumber="1319" year="2021">
<bibRefCitation author="Menis FT &amp; Rodrigues SR" box="[671,1012,593,618]" pageId="4" pageNumber="1319" pagination="928" refId="ref26719" refString="Menis FT, Rodrigues SR. 2021. Occurrence of Phytophagous Scarabaeidae (Coleoptera) in a pasture area at the Balsamo municipality, Sao Paulo, Brazil. EntomoBrasilis, 14: e 928. doi: 10.12741 / ebrasi lis. v 14. e 928" type="journal article" year="2021">Menis and Rodrigues 2021: 3</bibRefCitation>
</treatmentCitation>
, 5 (citation);
<bibRefCitation author="Vaz-de-Mello FZ &amp; Grossi PC" box="[160,494,628,653]" pageId="4" pageNumber="1319" refId="ref27519" refString="Vaz-de-Mello FZ, Grossi PC. 2022. Melolonthidae in Catalogo Taxonomico da Fauna do Brasil. PNUD. http: // fauna. jbrj. gov. br / fauna / faunadobrasil / 127169." type="url" year="2022">Vaz-de-Mello and Grossi 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="4.[160,1157,142,757]" pageId="4" pageNumber="1319">
<taxonomicName authority="(Burmeister 1844)" baseAuthorityName="Burmeister" baseAuthorityYear="1844" box="[218,663,662,687]" class="Insecta" family="Rutelidae" genus="Cotalpa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1319" phylum="Arthropoda" rank="subGenus" subGenus="Byrsopolis">
<emphasis box="[218,305,662,687]" italics="true" pageId="4" pageNumber="1319">Cotalpa</emphasis>
(
<emphasis box="[324,435,662,687]" italics="true" pageId="4" pageNumber="1319">Byrsopolis</emphasis>
) (
<bibRefCitation author="Burmeister HCC" box="[461,654,662,687]" pageId="4" pageNumber="1319" refId="ref25824" refString="Burmeister HCC. 1844. Besondere Entomologie, Fortsetzung. Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. Handuch der Entomologie. Vierter Band. Berlin: T. C. F. Enslin; p. xii. 588." type="book" year="1844">Burmeister 1844</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation author="Ohaus F" box="[680,880,662,687]" pageId="4" pageNumber="1319" pagination="256 - 260" refId="ref27018" refString="Ohaus F. 1915. XVI Beitrag zur Kenntnis der Ruteliden. (Col. lamell.). Deutsche entomologische Zeitschrift. Berlin Beiheft; p. 256 - 260." type="book chapter" year="1915">Ohaus 1915: 256</bibRefCitation>
(status change, key to subgenera); 1918: 10 (checklist); 1934: 37 (subgenus redescription, checklist, distribution), 173 (illustration);
<bibRefCitation author="Blackwelder RE" box="[356,606,732,757]" pageId="4" pageNumber="1319" pagination="189 - 306" refId="ref25704" refString="Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexico, Central America, The West Indies, and South America. Washington: Smithsonian Institution, United States National Museum. Bulletin 185, Part 2; p. 189 - 306." type="journal article" year="1944">Blackwelder 1944: 235</bibRefCitation>
(checklist).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="1319" type="diagnosis">
<paragraph blockId="4.[160,1156,801,1138]" pageId="4" pageNumber="1319">
<emphasis bold="true" box="[160,282,801,827]" italics="true" pageId="4" pageNumber="1319">Diagnosis.</emphasis>
Frontoclypeal suture complete, almost linear or with curvilinear extremities; frons glabrous or with short setae; clypeus glabrous or setose; mandibles externally rounded with molar region striated, with or without projections; apex of prementum weakly to strongly emarginated; antennae with 10 antennomeres; antennal club longer than antennomeres IIVII combined, with marginal areas unconstricted. Thorax with mesosternal process not exceeding mesocoxae apex. Elytra punctate, with or without striae, striae with small punctures; apical calli distinct; hind wings with anterior margin (PC +C, ScA, ScP, RA
<subScript attach="left" box="[338,371,1055,1071]" fontSize="7" pageId="4" pageNumber="1319">1+2</subScript>
, RA
<subScript attach="left" box="[413,423,1055,1071]" fontSize="7" pageId="4" pageNumber="1319">3</subScript>
) glabrous, RA
<subScript attach="left" box="[578,588,1055,1071]" fontSize="7" pageId="4" pageNumber="1319">3</subScript>
margin very close to RA
<subScript attach="left" box="[858,868,1055,1071]" fontSize="7" pageId="4" pageNumber="1319">4</subScript>
+ RP
<subScript attach="left" box="[919,929,1055,1071]" fontSize="7" pageId="4" pageNumber="1319">1</subScript>
, but not reaching it; apex of Cu setose; AA
<subScript attach="left" box="[408,418,1089,1105]" fontSize="7" pageId="4" pageNumber="1319">3</subScript>
developed and converging to Cu. Parameres bifurcated, parallel, convergent or divergent; shape without projections.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="1320" pageId="4" pageNumber="1319" type="description">
<paragraph blockId="4.[160,1156,1182,1728]" lastBlockId="5.[160,1157,142,687]" lastPageId="5" lastPageNumber="1320" pageId="4" pageNumber="1319">
<emphasis bold="true" box="[160,326,1182,1208]" italics="true" pageId="4" pageNumber="1319">Redescription.</emphasis>
<emphasis box="[342,396,1183,1208]" italics="true" pageId="4" pageNumber="1319">Body</emphasis>
. Shape elongated, oval, sides parallel to rounded in dorsal view; dorsal surface usually convex, ventrally flattened in lateral view; females usually bigger than males and with a more distinctly convex dorsal surface. Colour very variable, from light yellow to completely black, with metallic reflections in some species, or with legs, head, pronotum and venter metallic green. Length
<quantity box="[730,917,1322,1347]" metricMagnitude="-2" metricUnit="m" metricValue="2.9105" metricValueMax="3.781" metricValueMin="2.04" pageId="4" pageNumber="1319" unit="mm" value="29.105" valueMax="37.81" valueMin="20.4">20.4037.81 mm</quantity>
; humeral width
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.5115" metricValueMax="2.073" metricValueMin="0.95" pageId="4" pageNumber="1319" unit="mm" value="15.115" valueMax="20.73" valueMin="9.5">9.50 20.73 mm</quantity>
, elytral width
<quantity box="[434,623,1357,1382]" metricMagnitude="-2" metricUnit="m" metricValue="1.671" metricValueMax="2.237" metricValueMin="1.105" pageId="4" pageNumber="1319" unit="mm" value="16.71" valueMax="22.37" valueMin="11.05">11.0522.37 mm</quantity>
.
<emphasis box="[633,690,1357,1382]" italics="true" pageId="4" pageNumber="1319">Head</emphasis>
. Clypeus rounded, trapezoidal, rectangular, semioval, or square; weakly to moderately concave, wider than long, widest at base; apex straight, truncated, emarginated, bilobed or trilobed; surface glabrous to hirsute; frontoclypeal suture clearly visible, almost straight or sinuous; clypeus and frons sometimes with isolated concavities, or a longitudinal midline formed by punctures; mandibles partially or totally covered by clypeus, glabrous or hirsute, smooth or punctate; maxillae symmetrical or asymmetrical; surface of prementum and area of insertion of palpigers excavated or not; premental apex with weak to strong emargination; labrum bilobed, with narrowly rounded to truncated lobes; male antennal club 1.53x longer than antennomeres IIVII combined, female antennal club smaller, about 1.21.5x longer than antennomeres IIVII combined.
<emphasis box="[456,531,1703,1728]" italics="true" pageId="4" pageNumber="1319">Thorax</emphasis>
. Pronotum with weak to strong concavities at isolated punctures or oblique lines near posterior angles; anterior angles obtuse, incurved; scutellar plate subtriangular, with punctures and setae varying in size from small to large, sparse to densely distributed; mesoventral process with apex rounded to oval, not exceeding mesocoxa.
<emphasis box="[415,476,246,271]" italics="true" pageId="5" pageNumber="1320">Elytra</emphasis>
. Humeri slightly concave, with small setae and small calli; elytral suture smooth to striate; disc with weak to strong striae, interstriae sparsely to densely punctate; epipleura sparsely to densely setose; hind wings with anterior margin near veins (PC+C, ScA, ScP, RA
<subScript attach="left" box="[503,535,361,377]" fontSize="7" pageId="5" pageNumber="1320">1+2</subScript>
, RA
<subScript attach="left" box="[579,589,361,377]" fontSize="7" pageId="5" pageNumber="1320">3</subScript>
) glabrous, vein AA
<subScript attach="left" box="[803,813,361,377]" fontSize="7" pageId="5" pageNumber="1320">3</subScript>
developed, converging to Cu; Cu setae only at apex; margin of RA
<subScript attach="left" box="[576,586,395,411]" fontSize="7" pageId="5" pageNumber="1320">3</subScript>
very close to RA
<subScript attach="left" box="[781,791,395,411]" fontSize="7" pageId="5" pageNumber="1320">4</subScript>
+ RP
<subScript attach="left" box="[845,855,395,411]" fontSize="7" pageId="5" pageNumber="1320">1</subScript>
, but not reaching it.
<emphasis box="[1102,1150,385,410]" italics="true" pageId="5" pageNumber="1320">Legs</emphasis>
. Protibia with three outer teeth; spur developed, at least 2x longer than wide, conical (spine-like) or flattened; females with protarsomeres IVV usually with longer setae than in males; apex of metatibiae with 1432 spine-like setae.
<emphasis box="[802,907,489,514]" italics="true" pageId="5" pageNumber="1320">Abdomen</emphasis>
. Tergite 8 striated or punctated, glabrous or hirsute.
<emphasis box="[519,624,524,549]" italics="true" pageId="5" pageNumber="1320">Aedeagus</emphasis>
. Parameres dorsally or laterally symmetrical or not, weakly to strongly bifurcated; rounded, parallel, divergent or convergent apex, weakly to strongly angular; lateral margins of parameres weakly to strongly excavated longitudinally, forming a small to long carina; right paramere sometimes thicker or more divergent.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="1320" type="distribution">
<paragraph blockId="5.[160,1156,745,909]" pageId="5" pageNumber="1320">
<emphasis bold="true" box="[160,307,745,771]" italics="true" pageId="5" pageNumber="1320">Distribution.</emphasis>
<taxonomicName authorityName="Burmeister" authorityYear="1844" box="[323,431,746,771]" class="Insecta" family="Rutelidae" genus="Byrsopolis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1320" phylum="Arthropoda" rank="genus">
<emphasis box="[323,431,746,771]" italics="true" pageId="5" pageNumber="1320">Byrsopolis</emphasis>
</taxonomicName>
has a disjunct distribution, with two species restricted to the coastal region of
<collectingCountry box="[359,523,780,805]" name="French Guiana" pageId="5" pageNumber="1320">French Guiana</collectingCountry>
(
<bibRefCitation author="Soula M" box="[536,676,780,805]" pageId="5" pageNumber="1320" pagination="66" refId="ref27367" refString="Soula M. 2010 a. Les Coleopteres du Nouveau Monde. Volume 4: Rutelini 4. Revision des 4. Une revision des genres Catoclastus, Homonyx. Retour sur quelques especes ou groupe d'especes de Pelidnota. Addenda 2010. (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. Saintry: AECFT; p. 66." type="book chapter" year="2010">Soula 2010a</bibRefCitation>
), and the others exclusive to the Brazilian Cerrado biome and Atlantic Forest found in Bahia, Distrito Federal, Goiás, Mato Grosso do Sul, Mato Grosso, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo and Paraná states, and
<collectingCountry box="[210,314,884,909]" name="Paraguay" pageId="5" pageNumber="1320">Paraguay</collectingCountry>
(Dept.
<collectingCountry box="[397,515,884,909]" name="Saint Pierre and Miquelon" pageId="5" pageNumber="1320">San Pedro</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>