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<document id="33708977B29B7FDB3D0B167BEF964B46" ID-DOI="10.11646/zootaxa.5530.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14023149" ID-ZooBank="41376D87-B401-4301-9DDC-54606653881F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730454251080" checkinUser="plazi" docAuthor="Kovac, Damir, Kameneva, Elena P., Korneyev, Severyn V., Araújo, Alexandre Santos, Savaris, Marcoandre, Smit, John T., Schneider, Alexander, Schreiber, Robert &amp; Korneyev, Valery A." docDate="2024" docId="03F029281E49FFDFFF215257FD77FBF5" docLanguage="en" docName="zootaxa.5530.1.1.pdf" docOrigin="Zootaxa 5530 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5530.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Aspistomella garleppi V. Korneyev &amp; Kameneva 2024, sp. nov." docType="treatment" docUuid="539952F5-EBD0-4671-828B-92BB1BF92327" docUuidSource="ZooBank" docVersion="2" lastPageNumber="44" masterDocId="FFC951501E60FFF4FFB6514DFFDAFFC0" masterDocTitle="Revision of the Aspistomella group of genera (Diptera: Ulidiidae: Pterocallinae: Lipsanini)" masterLastPageNumber="117" masterPageNumber="1" pageNumber="42" updateTime="1730455055018" updateUser="ExternalLinkService">
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<mods:title id="DF3270CF1A0A332CC29BFC97CB5137DA">Revision of the Aspistomella group of genera (Diptera: Ulidiidae: Pterocallinae: Lipsanini)</mods:title>
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<mods:namePart id="FD2D617A04BE682D2A10E30A220E010D">Kovac, Damir</mods:namePart>
<mods:affiliation id="A88F99C2F51B762A1B5FAB918A125EBB">Forschungsinstitut Senckenberg, Senckenberganlage 25, Frankfurt am Main, 60325 Germany https: // orcid. org / 0009 - 0000 - 2202 - 4052 https: // orcid. org / 0000 - 0001 - 8451 - 5500</mods:affiliation>
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<mods:namePart id="8C32518DBF15CDE0BE4C964875953D07">Kameneva, Elena P.</mods:namePart>
<mods:affiliation id="9E374057075134F3E367514B6C471232">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 &amp; Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
<mods:nameIdentifier id="90294D36AC9E34234C5480920398CFF3" type="email">valery.korneyev@gmail.com</mods:nameIdentifier>
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<mods:namePart id="96C3942B09CD981015FC1D1A68FFF120">Korneyev, Severyn V.</mods:namePart>
<mods:affiliation id="C21DF1BF4D33EB612438ADD3AC2B4A8E">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 &amp; University of California, Davis, Department of Entomology and Nematology, College of Agricultural and Environmental Sciences, One Shields Avenue, Davis, California, 95616 USA https: // orcid. org / 0000 - 0001 - 8599 - 7695 &amp; California Department of Food and Agriculture, Plant Pest Diagnostics Branch, 3294 Meadowview Road, Sacramento, California, 95832 - 1448 USA</mods:affiliation>
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<mods:namePart id="6D30A1D245CB4BF5858E696CCBC0F27B">Araújo, Alexandre Santos</mods:namePart>
<mods:affiliation id="C84B0CD59091043294F56600E4C5E8A9">University of São Paulo, “ Luiz de Queiroz ” College of Agriculture, Av. Pádua Dias, 11, Piracicaba, São Paulo, Brazil https: // orcid. org / 0000 - 0003 - 0245 - 763 X https: // orcid. org / 0000 - 0002 - 9145 - 6059</mods:affiliation>
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<mods:namePart id="9035D9233377D61E761B2BDBEB0D4AA7">Savaris, Marcoandre</mods:namePart>
<mods:affiliation id="BF6E0D78C23960922CA123DC1227A0C5">University of São Paulo, “ Luiz de Queiroz ” College of Agriculture, Av. Pádua Dias, 11, Piracicaba, São Paulo, Brazil https: // orcid. org / 0000 - 0003 - 0245 - 763 X https: // orcid. org / 0000 - 0002 - 9145 - 6059</mods:affiliation>
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<mods:namePart id="CF65B391A0BB0064BF1CEC2E5AAA634F">Smit, John T.</mods:namePart>
<mods:affiliation id="E42AECA24DF4C037DAF702B4BE1E9058">EIS Kenniscentrum Insecten en andere ongewervelden p / a Naturalis Biodiversity Center Postbus 9517 2300 RA Leiden, The Netherlands https: // orcid. org / 0000 - 0002 - 1568 - 5183</mods:affiliation>
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<mods:namePart id="78E0C316A59825C2B1F73A68665BF994">Schneider, Alexander</mods:namePart>
<mods:affiliation id="906BB54434C552EDAB2FDCB617077B12">Forschungsinstitut Senckenberg, Senckenberganlage 25, Frankfurt am Main, 60325 Germany https: // orcid. org / 0009 - 0000 - 2202 - 4052 https: // orcid. org / 0000 - 0001 - 8451 - 5500</mods:affiliation>
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<mods:namePart id="59A220925CDD23E73507EE6E270CB28E">Schreiber, Robert</mods:namePart>
<mods:affiliation id="BABB210626258266AD71AB3CBE489B36">Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
<mods:nameIdentifier id="8B8071B00433A786EF9C1EC3EEBDC081" type="email">valery.korneyev@gmail.com</mods:nameIdentifier>
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<mods:namePart id="76FE602B9DDCA269C5780B2CB92D3697">Korneyev, Valery A.</mods:namePart>
<mods:affiliation id="DAD8AEE359E1B2BE3254DEE717EBEAB7">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 &amp; Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
<mods:nameIdentifier id="24767F26C65F94421E466ED3349EC004" type="email">valery.korneyev@gmail.com</mods:nameIdentifier>
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<mods:title id="DE09B8E51501F8DF4A91AB65FD4F8B93">Zootaxa</mods:title>
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<mods:date id="0B26B6E7E967479257E036F77C2961B7">2024</mods:date>
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<mods:number id="4BCD5ACC5DFA74D4BE616E9FD375095B">2024-10-30</mods:number>
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<mods:number id="E664B1B7A30C3F7A99F7BCFB0DE268A2">5530</mods:number>
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<mods:url id="0F3FC95790ABD6B021A60918BE1AAE30">http://dx.doi.org/10.11646/zootaxa.5530.1.1</mods:url>
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<mods:classification id="AA78935ED45E044ADDB7D9EE06006AB3">journal article</mods:classification>
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<treatment id="03F029281E49FFDFFF215257FD77FBF5" LSID="urn:lsid:zoobank.org:act:539952F5-EBD0-4671-828B-92BB1BF92327" httpUri="http://treatment.plazi.org/id/03F029281E49FFDFFF215257FD77FBF5" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
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<emphasis id="B92D442C1E49FFDDFF215257FC8DFCF5" bold="true" box="[151,855,794,821]" pageId="41" pageNumber="42">
<taxonomicName id="4C59E3BD1E49FFDDFF215257FD36FCF5" authority="V. Korneyev &amp; Kameneva" authorityName="V. Korneyev &amp; Kameneva" authorityYear="2024" box="[151,748,794,821]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="garleppi" status="sp. nov.">
<emphasis id="B92D442C1E49FFDDFF215257FE46FCF5" bold="true" box="[151,412,794,821]" italics="true" pageId="41" pageNumber="42">Aspistomella garleppi</emphasis>
V. Korneyev &amp; Kameneva
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,
<taxonomicNameLabel id="A21EF9571E49FFDDFD4C5256FC8DFCF5" box="[762,855,795,821]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C343CBB51E49FFDDFF21522EFEF8FCBD" box="[151,290,867,893]" pageId="41" pageNumber="42" type="description">
<paragraph id="8BE6983E1E49FFDDFF21522EFEF8FCBD" blockId="41.[151,952,794,893]" box="[151,290,867,893]" pageId="41" pageNumber="42">
<figureCitation id="136284BB1E49FFDDFF21522EFF20FCBD" box="[151,250,867,893]" captionStart="FIGURE 6" captionStartId="14.[151,250,1521,1545]" captionTargetBox="[177,1387,245,1460]" captionTargetId="figure-30@14.[151,1436,211,1497]" captionTargetPageId="14" captionText="FIGURE 6. Aspistomella, wing. A, A. sachavaca sp. nov.; B, A. schnusei sp. nov.; C, A. angustifrons (Hendel); D, A. garleppi sp. nov.; E, A. crucifera (Hendel); F, A. quinquincisa sp. nov.; G, A. teresensis sp. nov.; H, A. obliqua sp. nov. Abbreviations: C—costal vein; c—costal cell; M1—first branch of media; m1—first medial cell; ptstg—pterostigma; r1—first radial cell; rm—radial-medial crossvein. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023163" httpUri="https://zenodo.org/record/14023163/files/figure.png" pageId="41" pageNumber="42">Figs 6 D</figureCitation>
,
<figureCitation id="136284BB1E49FFDDFEB0522EFEF8FCBD" box="[262,290,867,893]" captionStart="FIGURE 24" captionStartId="42.[151,250,1698,1722]" captionTargetBox="[151,1436,191,1675]" captionTargetId="figure-30@42.[151,1436,181,1675]" captionTargetPageId="42" captionText="FIGURE 24. Aspistomella garleppi sp. nov. holotype ♂ (A–E, H) and paratype (sex unknown) (F–H). A, habitus, left; B, head and thorax, dorsal; C, head, antero-dorsal; D, head left; E, thorax and fore femur, left; F, thorax and base of abdomen, dorsal; G, abdominal syntergite 1+2, setae and microtrichia; H, labels." figureDoi="http://doi.org/10.5281/zenodo.14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="41" pageNumber="42">24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E49FFDDFF2152E6FDF6FBB9" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8BE6983E1E49FFDDFF2152E6FDF6FBB9" blockId="41.[151,1437,939,2010]" pageId="41" pageNumber="42">
<emphasis id="B92D442C1E49FFDDFF2152E6FE17FC05" bold="true" box="[151,461,939,965]" pageId="41" pageNumber="42">Material examined. Type.</emphasis>
<materialsCitation id="3B3192631E49FFDDFE6052E6FA43FBCD" collectionCode="SDEI" collectorName="O. Garlepp &amp; Coll. W. Schnuse &amp; W. Hennig" country="Peru" location="Madre De Dios" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="B92D442C1E49FFDDFE6052E6FD99FC05" bold="true" box="[470,579,939,965]" pageId="41" pageNumber="42">
<typeStatus id="54E2269C1E49FFDDFE6052E6FD99FC05" box="[470,579,939,965]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D5F53B71E49FFDDFDFA52E6FDBBFC05" box="[588,609,939,965]" count="1" pageId="41" pageNumber="42" type="male">♂</specimenCount>
:
<emphasis id="B92D442C1E49FFDDFDC452E6FD5AFC05" bold="true" box="[626,640,939,965]" pageId="41" pageNumber="42">“</emphasis>
<collectingCountry id="F34ED8AE1E49FFDDFD3652E6FD69FC05" box="[640,691,939,965]" name="Peru" pageId="41" pageNumber="42">Peru</collectingCountry>
/
<collectingRegion id="499D56DC1E49FFDDFD7B52E6FCA4FC05" box="[717,894,939,965]" country="Peru" name="Madre de Dios" pageId="41" pageNumber="42">Madre De Dios</collectingRegion>
/
<collectorName id="26ACFDE81E49FFDDFC2E52E6FBCCFC05" box="[920,1046,939,965]" pageId="41" pageNumber="42">O. Garlepp</collectorName>
c.”, “
<collectorName id="26ACFDE81E49FFDDFBEC52E6FAFAFC05" box="[1114,1312,939,965]" pageId="41" pageNumber="42">Coll. W. Schnuse</collectorName>
/ 1911-3”, “
<taxonomicName id="4C59E3BD1E49FFDDFF175282FF26FC29" authorityName="Loew" authorityYear="1868" box="[161,252,975,1001]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">Euxesta</taxonomicName>
s. (nova)” [pencil hadwritten label of
<collectorName id="26ACFDE81E49FFDDFD095282FC9BFC29" box="[703,833,975,1001]" pageId="41" pageNumber="42">W. Hennig</collectorName>
]; (right antenna, left wing and left mid leg missing; abdominal segments 4–5 missing, apparently removed by
<collectorName id="26ACFDE81E49FFDDFCAE52BEFC55FBCD" box="[792,911,1011,1037]" pageId="41" pageNumber="42">W. Hennig</collectorName>
for preparation in 1938–1939 and lost) (
<collectionCode id="ED4800FB1E49FFDDFAFA52BEFA4AFBCD" box="[1356,1424,1011,1037]" pageId="41" pageNumber="42">SDEI</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54E2269C1E49FFDDFF21555AFED2FBF1" box="[151,264,1047,1073]" pageId="41" pageNumber="42" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5F53B71E49FFDDFEA0555AFE42FBF1" box="[278,408,1047,1073]" count="1" pageId="41" pageNumber="42" type="generic">1 specimen</specimenCount>
[sex unknown] (both antennae, hind legs, mouthparts and abdomen missing),
<materialsCitation id="3B3192631E49FFDDFA9F555AFDF2FBB9" collectionCode="SMTD" country="Peru" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="9D5F53B71E49FFDDFA9F555AFA8DFBF1" box="[1321,1367,1047,1073]" count="1" pageId="41" pageNumber="42" type="male">1 ♂</specimenCount>
(both antennae, mouthparts and ventral part of abdomen including all sternites, pleural membrane and genitalia missing), same labels as in
<typeStatus id="54E2269C1E49FFDDFEED5512FE67FBB9" box="[347,445,1119,1145]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED4800FB1E49FFDDFE7A5512FDFAFBB9" box="[460,544,1119,1145]" pageId="41" pageNumber="42">SMTD</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E49FFDDFF7155CEFDD2FA59" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="8BE6983E1E49FFDDFF7155CEFDD2FA59" blockId="41.[151,1437,939,2010]" pageId="41" pageNumber="42">
<emphasis id="B92D442C1E49FFDDFF7155CEFEE0FB5D" bold="true" box="[199,314,1155,1181]" pageId="41" pageNumber="42">Diagnosis</emphasis>
.
<taxonomicName id="4C59E3BD1E49FFDDFEF155CEFDE6FB5D" authorityName="V. Korneyev &amp; Kameneva" authorityYear="2024" box="[327,572,1155,1181]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="garleppi">
<emphasis id="B92D442C1E49FFDDFEF155CEFDE6FB5D" box="[327,572,1155,1181]" italics="true" pageId="41" pageNumber="42">Aspistomella garleppi</emphasis>
</taxonomicName>
are moderately large flies (WL&gt; 6.0–
<quantity id="4CA135DB1E49FFDDFC6955CEFBEDFB5D" box="[991,1079,1155,1181]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="41" pageNumber="42" unit="mm" value="6.4">6.4 mm</quantity>
), differing from all other species of the group of genera by the combination of the very long pterostigma, straight costal cell, vein R
<subScript id="17DD9A7B1E49FFDDFB4355F4FB27FB07" attach="left" box="[1269,1277,1209,1223]" fontSize="6" pageId="41" pageNumber="42">1</subScript>
dorsoapically setulose, crossvein r-m slightly proximal to the level of R
<subScript id="17DD9A7B1E49FFDDFCA35590FCC7FB2B" attach="left" box="[789,797,1245,1259]" fontSize="6" pageId="41" pageNumber="42">1</subScript>
apex but far distal from Sc apex, head with narrow frons, face uniformly grey or silvery microtrichose, epistome strongly developed, ocellar and outer vertical seta short, 0.2–0.3× longer than orbital and inner vertical seta, prescutellar acrostichal seta absent, wing extremely elongated, 3.8–4× longer than wide; with pattern of narrow subbasal and wide discal crossband separated from the entirely dark apex by hyaline interspace covering the entire area of cell dm between crossveins r-m and dm-m; and mid tibia ventroapically with 1 long spine.
</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E49FFDFFF7154EEFD95FC9D" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42" type="description">
<paragraph id="8BE6983E1E49FFDDFF7154EEFAF6F889" blockId="41.[151,1437,939,2010]" pageId="41" pageNumber="42">
<emphasis id="B92D442C1E49FFDDFF7154EEFE8BFA7D" bold="true" box="[199,337,1443,1469]" pageId="41" pageNumber="42">Description</emphasis>
. Male.
<emphasis id="B92D442C1E49FFDDFE1054EEFE3CFA7D" bold="true" box="[422,486,1443,1469]" pageId="41" pageNumber="42">Head</emphasis>
(
<figureCitation id="136284BB1E49FFDDFE4254EEFD8FFA7D" box="[500,597,1443,1469]" captionStart="FIGURE 3" captionStartId="10.[151,250,1454,1478]" captionTargetBox="[165,1431,193,1414]" captionTargetId="figure-30@10.[151,1435,181,1430]" captionTargetPageId="10" captionText="FIGURE 3. Lipsanini, head (A–B—dorsal, C–D—left lateral view). A, Ulivellia amnoni sp. nov.; B, Euxesta quaternaria Loew; C, Aspistomella garleppi sp. nov.; D, “Euxesta” insolita Hendel. Abbreviations: a orb s—anterior orbital seta; ar— arista; clyp—clypeus; epst—epistome; fr s—frontal seta; fr stl—frontal setulae; fr vit—frontal vitta; frorb plt—fronto-orbital plate; gn s—genal seta; i vt s—inner vertical seta; ifr s—interfrontal seta; o vt s—outer vertical seta; oc s—ocellar seta; p orb s—posterior orbital seta; pafc—parafacial; ped—pedicel; pfr stl—parafrontal setula; pgn s—postgenal seta; poc s—postocellar seta; pocl s—postocular setae; pped—postpedicel; pst s—perisomal setae; vt plt—vertical plate." figureDoi="http://doi.org/10.5281/zenodo.14023157" httpUri="https://zenodo.org/record/14023157/files/figure.png" pageId="41" pageNumber="42">Figs 3 C</figureCitation>
,
<figureCitation id="136284BB1E49FFDDFDD654EEFD63FA7D" box="[608,697,1443,1469]" captionStart="FIGURE 24" captionStartId="42.[151,250,1698,1722]" captionTargetBox="[151,1436,191,1675]" captionTargetId="figure-30@42.[151,1436,181,1675]" captionTargetPageId="42" captionText="FIGURE 24. Aspistomella garleppi sp. nov. holotype ♂ (A–E, H) and paratype (sex unknown) (F–H). A, habitus, left; B, head and thorax, dorsal; C, head, antero-dorsal; D, head left; E, thorax and fore femur, left; F, thorax and base of abdomen, dorsal; G, abdominal syntergite 1+2, setae and microtrichia; H, labels." figureDoi="http://doi.org/10.5281/zenodo.14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="41" pageNumber="42">24 B–D</figureCitation>
) ratio (length: height: width) = 1: 1.1–1.2: 1.6; frons, parafacial, and gena reddish brown, most of epistome and occiput dark brown to black; face below lunule white microtrichose. Frons moderately wide, slightly narrowed posteriorly (
<figureCitation id="136284BB1E49FFDDFCB654A6FCB4F9C5" box="[768,878,1515,1541]" captionStart="FIGURE 24" captionStartId="42.[151,250,1698,1722]" captionTargetBox="[151,1436,191,1675]" captionTargetId="figure-30@42.[151,1436,181,1675]" captionTargetPageId="42" captionText="FIGURE 24. Aspistomella garleppi sp. nov. holotype ♂ (A–E, H) and paratype (sex unknown) (F–H). A, habitus, left; B, head and thorax, dorsal; C, head, antero-dorsal; D, head left; E, thorax and fore femur, left; F, thorax and base of abdomen, dorsal; G, abdominal syntergite 1+2, setae and microtrichia; H, labels." figureDoi="http://doi.org/10.5281/zenodo.14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="41" pageNumber="42">Fig. 24 C</figureCitation>
), 1–1.1× as long (from lunule to anterior ocellus) or 1.6–2× (from lunule to inner vertical seta) as wide (at lunule), with black, steel shining, sparsely microtrichose vertex; orbits in posterior half, parafacial, narrow transverse band on face above fold, gena and occiput silver-white microtrichose; parafacial less than half as wide as postpedicel. Vertical plates with 1 strong and 1 setula-like orbital setae. Frontal plate with 10–12 short lateroclinate parafrontal setulae, frontal vitta sparsely whitish microtrichose, with 6–7 moderately developed frontal and 3–5 stronger interfrontal setae on each side (
<figureCitation id="136284BB1E49FFDDFBDC57D2FB0CF979" box="[1130,1238,1695,1721]" captionStart="FIGURE 24" captionStartId="42.[151,250,1698,1722]" captionTargetBox="[151,1436,191,1675]" captionTargetId="figure-30@42.[151,1436,181,1675]" captionTargetPageId="42" captionText="FIGURE 24. Aspistomella garleppi sp. nov. holotype ♂ (A–E, H) and paratype (sex unknown) (F–H). A, habitus, left; B, head and thorax, dorsal; C, head, antero-dorsal; D, head left; E, thorax and fore femur, left; F, thorax and base of abdomen, dorsal; G, abdominal syntergite 1+2, setae and microtrichia; H, labels." figureDoi="http://doi.org/10.5281/zenodo.14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="41" pageNumber="42">Fig. 24 C</figureCitation>
). Eye 1.3× higher than long. Face white microtrichose above epistome; epistome black to brown, sparsely white microtrichose, with metallic bluish sheen. Clypeus black, sparsely white microtrichose, with metallic sheen, 0.3× higher than epistome. Gena brownish yellow, narrowly white microtrichose along orbit, with 1–2 long genal setae and 4–6 additional long peristomal setae anterior to it, shorter than genal seta. Occiput brown to black, silver-grey microtrichose.
</paragraph>
<paragraph id="8BE6983E1E49FFDDFF71561EFB0FF819" blockId="41.[151,1437,939,2010]" pageId="41" pageNumber="42">Antenna yellow to reddish brown, inserted below the middle of the eye; scape with black setulae; postpedicel moderately long, rounded, 1.8–2.0× longer than wide; arista brown apically, yellow in basal 1/4, 2-segmented, bare. Mouthparts brown to black, prementum black, sparsely microtrichose. Palp yellow to reddish brown, apically rounded, with 10–15 black setulae, of which 5–6 subapical setulae twice as long as other setulae.</paragraph>
<caption id="DF26C8B61E4AFFDEFF2157EFFCCDF8C3" ID-DOI="http://doi.org/10.5281/zenodo.14023222" ID-Zenodo-Dep="14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1698,1722]" targetBox="[151,1436,191,1675]" targetPageId="42" targetType="figure">
<paragraph id="8BE6983E1E4AFFDEFF2157EFFCCDF8C3" blockId="42.[151,1436,1698,1795]" pageId="42" pageNumber="43">
<emphasis id="B92D442C1E4AFFDEFF2157EFFEC5F97B" bold="true" box="[151,287,1698,1723]" pageId="42" pageNumber="43">FIGURE 24.</emphasis>
<taxonomicName id="4C59E3BD1E4AFFDEFE9357EEFDDEF97B" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[293,516,1699,1723]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="garleppi" status="sp. nov.">
<emphasis id="B92D442C1E4AFFDEFE9357EEFDDEF97B" box="[293,516,1699,1723]" italics="true" pageId="42" pageNumber="43">Aspistomella garleppi</emphasis>
</taxonomicName>
<emphasis id="B92D442C1E4AFFDEFDBF57EEFD83F97B" bold="true" box="[521,601,1699,1723]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="A21EF9571E4AFFDEFDBF57EEFD83F97B" box="[521,601,1699,1723]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype ♂ (
<emphasis id="B92D442C1E4AFFDEFD6957EFFCD5F97A" bold="true" box="[735,783,1698,1722]" pageId="42" pageNumber="43">A–E</emphasis>
,
<emphasis id="B92D442C1E4AFFDEFCAD57EEFCF5F97B" bold="true" box="[795,815,1699,1723]" pageId="42" pageNumber="43">H</emphasis>
) and paratype (sex unknown) (
<emphasis id="B92D442C1E4AFFDEFBDF57EEFB43F97B" bold="true" box="[1129,1177,1699,1723]" pageId="42" pageNumber="43">F–H</emphasis>
).
<emphasis id="B92D442C1E4AFFDEFB1B57EFFB65F97A" bold="true" box="[1197,1215,1698,1722]" pageId="42" pageNumber="43">A</emphasis>
, habitus, left;
<emphasis id="B92D442C1E4AFFDEFAF957EEFABAF97B" bold="true" box="[1359,1376,1699,1723]" pageId="42" pageNumber="43">B</emphasis>
, head and thorax, dorsal;
<emphasis id="B92D442C1E4AFFDEFEEB578BFEB5F91E" bold="true" box="[349,367,1734,1758]" pageId="42" pageNumber="43">C</emphasis>
, head, antero-dorsal;
<emphasis id="B92D442C1E4AFFDEFDFB578AFD85F91F" bold="true" box="[589,607,1735,1759]" pageId="42" pageNumber="43">D</emphasis>
, head left;
<emphasis id="B92D442C1E4AFFDEFD65578AFD3EF91F" bold="true" box="[723,740,1735,1759]" pageId="42" pageNumber="43">E</emphasis>
, thorax and fore femur, left;
<emphasis id="B92D442C1E4AFFDEFBBB578AFBC6F91F" bold="true" box="[1037,1052,1735,1759]" pageId="42" pageNumber="43">F</emphasis>
, thorax and base of abdomen, dorsal;
<emphasis id="B92D442C1E4AFFDEFF2157A7FF71F8C2" bold="true" box="[151,171,1770,1794]" pageId="42" pageNumber="43">G</emphasis>
, abdominal syntergite 1+2, setae and microtrichia;
<emphasis id="B92D442C1E4AFFDEFD0257A6FD12F8C3" bold="true" box="[692,712,1771,1795]" pageId="42" pageNumber="43">H</emphasis>
, labels.
</paragraph>
</caption>
<paragraph id="8BE6983E1E4AFFDFFF715618FC22FF15" blockId="42.[151,1437,1877,2011]" lastBlockId="43.[151,1437,151,1077]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="B92D442C1E4AFFDEFF715618FEFAF8AF" bold="true" box="[199,288,1877,1903]" pageId="42" pageNumber="43">Thorax</emphasis>
(
<figureCitation id="136284BB1E4AFFDEFE995618FE64F8AF" box="[303,446,1877,1903]" captionStart="FIGURE 24" captionStartId="42.[151,250,1698,1722]" captionTargetBox="[151,1436,191,1675]" captionTargetId="figure-30@42.[151,1436,181,1675]" captionTargetPageId="42" captionText="FIGURE 24. Aspistomella garleppi sp. nov. holotype ♂ (A–E, H) and paratype (sex unknown) (F–H). A, habitus, left; B, head and thorax, dorsal; C, head, antero-dorsal; D, head left; E, thorax and fore femur, left; F, thorax and base of abdomen, dorsal; G, abdominal syntergite 1+2, setae and microtrichia; H, labels." figureDoi="http://doi.org/10.5281/zenodo.14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="42" pageNumber="43">Figs 24 E, F</figureCitation>
) black, on pleura with silvery sheen and sparse grey and brown (posterior of transverse suture and in supraalar area) microtrichia not hiding underlying cuticle. Mesonotal scutum (
<figureCitation id="136284BB1E4AFFDEFB0C5634FAFDF853" box="[1210,1319,1913,1939]" captionStart="FIGURE 24" captionStartId="42.[151,250,1698,1722]" captionTargetBox="[151,1436,191,1675]" captionTargetId="figure-30@42.[151,1436,181,1675]" captionTargetPageId="42" captionText="FIGURE 24. Aspistomella garleppi sp. nov. holotype ♂ (A–E, H) and paratype (sex unknown) (F–H). A, habitus, left; B, head and thorax, dorsal; C, head, antero-dorsal; D, head left; E, thorax and fore femur, left; F, thorax and base of abdomen, dorsal; G, abdominal syntergite 1+2, setae and microtrichia; H, labels." figureDoi="http://doi.org/10.5281/zenodo.14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="42" pageNumber="43">Fig. 24 F</figureCitation>
) 1.6–1.7× longer than wide; black setulose, with 12–14 rows of setulae between rows of dorsocentral setulae; acrostichal prescutellar seta absent; prescutellar area with 2–4 rows of setulae between posterior dorsocentral setae. Scutellum dorsally slightly convex, black, subshining, very sparsely white microtrichose, apically shagreened, without setulae. Subscutellum and mediotergite shining black. All the setae and setulae black.
</paragraph>
<paragraph id="8BE6983E1E4BFFDFFF715193FCB6FD45" blockId="43.[151,1437,151,1077]" pageId="43" pageNumber="44">
<emphasis id="B92D442C1E4BFFDFFF715193FED2FF38" bold="true" box="[199,264,222,248]" pageId="43" pageNumber="44">Wing</emphasis>
(
<figureCitation id="136284BB1E4BFFDFFEAE5192FEA2FF39" box="[280,376,223,249]" captionStart="FIGURE 6" captionStartId="14.[151,250,1521,1545]" captionTargetBox="[177,1387,245,1460]" captionTargetId="figure-30@14.[151,1436,211,1497]" captionTargetPageId="14" captionText="FIGURE 6. Aspistomella, wing. A, A. sachavaca sp. nov.; B, A. schnusei sp. nov.; C, A. angustifrons (Hendel); D, A. garleppi sp. nov.; E, A. crucifera (Hendel); F, A. quinquincisa sp. nov.; G, A. teresensis sp. nov.; H, A. obliqua sp. nov. Abbreviations: C—costal vein; c—costal cell; M1—first branch of media; m1—first medial cell; ptstg—pterostigma; r1—first radial cell; rm—radial-medial crossvein. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023163" httpUri="https://zenodo.org/record/14023163/files/figure.png" pageId="43" pageNumber="44">Fig. 6 D</figureCitation>
)
<quantity id="4CA135DB1E4BFFDFFE3F5193FDCBFF39" box="[393,529,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="6.7" metricValueMin="6.5" pageId="43" pageNumber="44" unit="mm" value="6.6" valueMax="6.7" valueMin="6.5">6.5–6.7 mm</quantity>
long, 3.8–4× longer than wide; basicostal cell hyaline; costal cell straight, 5–6× longer than wide, pale brownish or yellowish at base and apex; pterostigma entirely brown, triangular, 6–8× longer than wide, vein R
<subScript id="17DD9A7B1E4BFFDFFED55074FEB1FE87" attach="left" box="[355,363,313,327]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
with 12–14 setulae dorsally, ending slightly distal to level of crossvein r-m level; costal vein between apices of R
<subScript id="17DD9A7B1E4BFFDFFECA5010FE5EFEAB" attach="left" box="[380,388,349,363]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
and R
<subScript id="17DD9A7B1E4BFFDFFE665010FE33FEAB" attach="left" box="[464,489,349,363]" fontSize="6" pageId="43" pageNumber="44">2+3</subScript>
slightly arcuate; vein R
<subScript id="17DD9A7B1E4BFFDFFD4F5010FCC8FEAB" attach="left" box="[761,786,349,363]" fontSize="6" pageId="43" pageNumber="44">2+3</subScript>
inconspicuously undulate. Crossvein r-m at proximal 2/5 of cell dm. Cell r
<subScript id="17DD9A7B1E4BFFDFFEEB50CCFEADFE4F" attach="left" box="[349,375,385,399]" fontSize="6" pageId="43" pageNumber="44">4+5</subScript>
4.0–4.5× longer than wide, apically narrowed. Cell m
<subScript id="17DD9A7B1E4BFFDFFC5250CCFC36FE4F" attach="left" box="[996,1004,385,399]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
narrow triangular, with pointed apex; ultimate section of M
<subScript id="17DD9A7B1E4BFFDFFE3A50E8FE4EFE73" attach="left" box="[396,404,421,435]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
3.6× longer than crossvein dm-m and 1.2–1.4× longer than penultimate section. Vein CuA Z-shaped, forming moderately long posteroapical lobe of cell cua along vein CuP, 1.8× longer than its anterior shoulder. Wing pattern with short pale brown subbasal crossband from humeral vein through cell cua into alula, wide discal crossband from apex of costal cell and pterostigma into basal half of cell dm, with wide hyaline interspace covering the entire area of cell dm between crossveins r-m and dm-m separating entirely dark apex by hyaline; cell m
<subScript id="17DD9A7B1E4BFFDFFF1B5314FF6FFDA7" attach="left" box="[173,181,601,615]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
entirely pale brown (
<figureCitation id="136284BB1E4BFFDFFE1C530AFDD4FDA1" box="[426,526,583,609]" captionStart="FIGURE 6" captionStartId="14.[151,250,1521,1545]" captionTargetBox="[177,1387,245,1460]" captionTargetId="figure-30@14.[151,1436,211,1497]" captionTargetPageId="14" captionText="FIGURE 6. Aspistomella, wing. A, A. sachavaca sp. nov.; B, A. schnusei sp. nov.; C, A. angustifrons (Hendel); D, A. garleppi sp. nov.; E, A. crucifera (Hendel); F, A. quinquincisa sp. nov.; G, A. teresensis sp. nov.; H, A. obliqua sp. nov. Abbreviations: C—costal vein; c—costal cell; M1—first branch of media; m1—first medial cell; ptstg—pterostigma; r1—first radial cell; rm—radial-medial crossvein. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023163" httpUri="https://zenodo.org/record/14023163/files/figure.png" pageId="43" pageNumber="44">Fig. 6 D</figureCitation>
). Cell cup and anal lobe 0.7–0.8× broader than cell cua
<subScript id="17DD9A7B1E4BFFDFFB335314FB57FDA7" attach="left" box="[1157,1165,601,615]" fontSize="6" pageId="43" pageNumber="44">1</subScript>
. Alula 2.4× longer than wide, grey. Calypters white, with long white cilia. Halter yellow.
</paragraph>
<paragraph id="8BE6983E1E4BFFDFFF7153C2FBACFCD5" blockId="43.[151,1437,151,1077]" pageId="43" pageNumber="44">
<emphasis id="B92D442C1E4BFFDFFF7153C2FF25FD69" bold="true" box="[199,255,655,681]" pageId="43" pageNumber="44">Legs</emphasis>
(
<figureCitation id="136284BB1E4BFFDFFEB953C2FE78FD69" box="[271,418,655,681]" captionStart="FIGURE 24" captionStartId="42.[151,250,1698,1722]" captionTargetBox="[151,1436,191,1675]" captionTargetId="figure-30@42.[151,1436,181,1675]" captionTargetPageId="42" captionText="FIGURE 24. Aspistomella garleppi sp. nov. holotype ♂ (A–E, H) and paratype (sex unknown) (F–H). A, habitus, left; B, head and thorax, dorsal; C, head, antero-dorsal; D, head left; E, thorax and fore femur, left; F, thorax and base of abdomen, dorsal; G, abdominal syntergite 1+2, setae and microtrichia; H, labels." figureDoi="http://doi.org/10.5281/zenodo.14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="43" pageNumber="44">Figs 24 A, E</figureCitation>
) long and narrow, mostly black, except tarsi brown; femora with metallic sheen, sparsely grey microtrichose. Fore femur with 2 rows of posterodorsal and posterior setae 0.6× longer than femur width, and 2–4 moderately strong posteroventral setulae in apical half. Mid femur anteriorly with short suberect setulae; mid tibia moderately thickened, ventrally with one spur-like setae 2× longer than tibia width.
</paragraph>
<paragraph id="8BE6983E1E4BFFDFFF715253FB66FCF9" blockId="43.[151,1437,151,1077]" box="[199,1212,798,825]" pageId="43" pageNumber="44">
<emphasis id="B92D442C1E4BFFDFFF715253FEE6FCF8" bold="true" box="[199,316,798,824]" pageId="43" pageNumber="44">Abdomen</emphasis>
black, short black setulose, with rows of whitish microtrichia (as on
<figureCitation id="136284BB1E4BFFDFFBF75252FB71FCF9" box="[1089,1195,799,825]" captionStart="FIGURE 24" captionStartId="42.[151,250,1698,1722]" captionTargetBox="[151,1436,191,1675]" captionTargetId="figure-30@42.[151,1436,181,1675]" captionTargetPageId="42" captionText="FIGURE 24. Aspistomella garleppi sp. nov. holotype ♂ (A–E, H) and paratype (sex unknown) (F–H). A, habitus, left; B, head and thorax, dorsal; C, head, antero-dorsal; D, head left; E, thorax and fore femur, left; F, thorax and base of abdomen, dorsal; G, abdominal syntergite 1+2, setae and microtrichia; H, labels." figureDoi="http://doi.org/10.5281/zenodo.14023222" httpUri="https://zenodo.org/record/14023222/files/figure.png" pageId="43" pageNumber="44">Fig. 24 G</figureCitation>
).
</paragraph>
<paragraph id="8BE6983E1E4BFFDFFF71520EFD95FC9D" blockId="43.[151,1437,151,1077]" box="[199,591,835,861]" pageId="43" pageNumber="44">Genitalia missing in all specimens.</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E4BFFDFFF71522AFB92FC41" box="[199,1096,870,897]" pageId="43" pageNumber="44" type="etymology">
<paragraph id="8BE6983E1E4BFFDFFF71522AFB92FC41" blockId="43.[151,1437,151,1077]" box="[199,1096,870,897]" pageId="43" pageNumber="44">
<emphasis id="B92D442C1E4BFFDFFF71522AFE92FC41" bold="true" box="[199,328,871,897]" pageId="43" pageNumber="44">Etymology</emphasis>
. The species is named after its collector, Otto Garlepp (1864–1959).
</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E4BFFDFFF7152C6FD77FBF5" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BE6983E1E4BFFDFFF7152C6FD77FBF5" blockId="43.[151,1437,151,1077]" pageId="43" pageNumber="44">
<emphasis id="B92D442C1E4BFFDFFF7152C6FEEEFC65" bold="true" box="[199,308,907,933]" pageId="43" pageNumber="44">Remarks</emphasis>
. The
<typeStatus id="54E2269C1E4BFFDFFEC152C6FE03FC65" box="[375,473,907,933]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E2269C1E4BFFDFFDA452C6FDA7FC65" box="[530,637,907,933]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
are damaged: abdominal segments 4–5 of the
<typeStatus id="54E2269C1E4BFFDFFB2652C6FB28FC65" box="[1168,1266,907,933]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
were detached by Willi Hennig in 1938–1939, but no slides can be found either in SDEI, or in MNKB. The
<typeStatus id="54E2269C1E4BFFDFFB0952E2FAC5FC09" box="[1215,1311,943,969]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
specimens are partially eaten by psocids and the abdomens are partly postabdomens completely or completely missing. The generic position of the species is provisional, as it requires more detailed analysis of the male genitalia, morphology- and / or molecular-based phylogenetic analysis.
</paragraph>
</subSubSection>
</treatment>
</document>
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