430 lines
62 KiB
XML
430 lines
62 KiB
XML
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<mods:title id="A859A43F653E71D737568324CB0ECCD9">Revision of the Aspistomella group of genera (Diptera: Ulidiidae: Pterocallinae: Lipsanini)</mods:title>
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<mods:namePart id="93DC3738EE54A8EB2FED8502F28BC20E">Kovac, Damir</mods:namePart>
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<mods:affiliation id="C053C659DA4C39553CFD5D08BFA39B3C">Forschungsinstitut Senckenberg, Senckenberganlage 25, Frankfurt am Main, 60325 Germany https: // orcid. org / 0009 - 0000 - 2202 - 4052 https: // orcid. org / 0000 - 0001 - 8451 - 5500</mods:affiliation>
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<mods:namePart id="5EEC8D0DEE3702A3B5C0F18B90C0E282">Kameneva, Elena P.</mods:namePart>
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<mods:affiliation id="0384C62F9A40D1AF5BDCA26128DF98C9">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 & Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
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<mods:nameIdentifier id="803F8029DA5D38D86887AD830C4F721E" type="email">valery.korneyev@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A17D52C2C1917A0C57394B3230043C5E">Korneyev, Severyn V.</mods:namePart>
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<mods:affiliation id="F9E94FBAEE655A55C6182E2F4CC2CB5C">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 & University of California, Davis, Department of Entomology and Nematology, College of Agricultural and Environmental Sciences, One Shields Avenue, Davis, California, 95616 USA https: // orcid. org / 0000 - 0001 - 8599 - 7695 & California Department of Food and Agriculture, Plant Pest Diagnostics Branch, 3294 Meadowview Road, Sacramento, California, 95832 - 1448 USA</mods:affiliation>
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<mods:namePart id="6F684FC9A531BEF60C4CE184FDCE3B0C">Araújo, Alexandre Santos</mods:namePart>
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<mods:affiliation id="A46F5DCFA69AFAA8B9E7A3914F6303C0">University of São Paulo, “ Luiz de Queiroz ” College of Agriculture, Av. Pádua Dias, 11, Piracicaba, São Paulo, Brazil https: // orcid. org / 0000 - 0003 - 0245 - 763 X https: // orcid. org / 0000 - 0002 - 9145 - 6059</mods:affiliation>
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<mods:namePart id="B0A8797F17169EC72E8C41AFFEBFD21D">Savaris, Marcoandre</mods:namePart>
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<mods:affiliation id="0BC3A39D82A2B6F88EEA1B004AADBDFB">University of São Paulo, “ Luiz de Queiroz ” College of Agriculture, Av. Pádua Dias, 11, Piracicaba, São Paulo, Brazil https: // orcid. org / 0000 - 0003 - 0245 - 763 X https: // orcid. org / 0000 - 0002 - 9145 - 6059</mods:affiliation>
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<mods:namePart id="28A9A69B73783C6470BF9662B3B6906E">Smit, John T.</mods:namePart>
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<mods:affiliation id="992846639EA926D8F6B43B0FF097BED3">EIS Kenniscentrum Insecten en andere ongewervelden p / a Naturalis Biodiversity Center Postbus 9517 2300 RA Leiden, The Netherlands https: // orcid. org / 0000 - 0002 - 1568 - 5183</mods:affiliation>
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<mods:affiliation id="52CB5DEEDB90972E986950585DA50987">Forschungsinstitut Senckenberg, Senckenberganlage 25, Frankfurt am Main, 60325 Germany https: // orcid. org / 0009 - 0000 - 2202 - 4052 https: // orcid. org / 0000 - 0001 - 8451 - 5500</mods:affiliation>
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<mods:namePart id="272CE85D7638653341E8DABBC2D73002">Schreiber, Robert</mods:namePart>
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<mods:affiliation id="63223698EA9DD11916E341B7CDED5E9A">Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
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<mods:nameIdentifier id="0061F7B0D870C140562D5FC680711190" type="email">valery.korneyev@gmail.com</mods:nameIdentifier>
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<mods:namePart id="251E548CD68A173ED92BF5753D044412">Korneyev, Valery A.</mods:namePart>
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<mods:affiliation id="58DE6D366A1B185EDD2EE79C6E176172">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 & Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
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<mods:nameIdentifier id="151ECDBE9D0AC68CE1892F9D02E58DA0" type="email">valery.korneyev@gmail.com</mods:nameIdentifier>
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<mods:title id="F6EBF08F1DACB4F075D709A8CB5A9D69">Zootaxa</mods:title>
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<mods:date id="49729B202C44165B1ACACF3FD509C256">2024</mods:date>
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<mods:number id="60AF6216BCFD0BE364DC107C2284AA0C">5530</mods:number>
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<treatment id="03F029281E46FFDDFF21536FFA43FD09" ID-DOI="http://doi.org/10.5281/zenodo.14023180" ID-Zenodo-Dep="14023180" LSID="urn:lsid:zoobank.org:act:95BFDD08-B74D-4CA1-93AF-021E86C2E16D" httpUri="http://treatment.plazi.org/id/03F029281E46FFDDFF21536FFA43FD09" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39">
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<subSubSection id="C343CBB51E46FFD2FF21536FFC0CFDA1" pageId="38" pageNumber="39" type="nomenclature">
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<taxonomicName id="4C59E3BD1E46FFD2FF21536FFCDBFDFD" authority="Kameneva & V. Korneyev" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[151,769,546,573]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="enderleini" status="sp. nov.">
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<emphasis id="B92D442C1E46FFD2FF21536FFE69FDFD" bold="true" box="[151,435,546,573]" italics="true" pageId="38" pageNumber="39">Aspistomella enderleini</emphasis>
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Kameneva & V. Korneyev
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21EF9571E46FFD2FCB8536EFCB6FDFD" box="[782,876,547,573]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8BE6983E1E46FFD2FF21530BFC0CFDA1" blockId="38.[151,982,546,645]" box="[151,982,582,609]" pageId="38" pageNumber="39">
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<subSubSection id="C343CBB51E46FFD2FF215326FE96FD45" box="[151,332,618,645]" pageId="38" pageNumber="39" type="description">
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<paragraph id="8BE6983E1E46FFD2FF215326FE96FD45" blockId="38.[151,982,546,645]" box="[151,332,618,645]" pageId="38" pageNumber="39">
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<figureCitation id="136284BB1E46FFD2FF215326FF2DFD45" box="[151,247,618,645]" captionStart="FIGURE 5" captionStartId="12.[151,250,1617,1641]" captionTargetBox="[175,1418,206,1566]" captionTargetId="figure-30@12.[151,1436,181,1593]" captionTargetPageId="12" captionText="FIGURE 5. Aspistomella, wing. A, A. pachitea sp. nov.; B, A. heteroptera Hendel; C, A. duo sp. nov.; D, A. tres sp. nov.; E, A. lobioptera Hendel; F, A. steyskali sp. nov.; G, A. lunata (Hendel); H, A. enderleini sp. nov. Abbreviations: C—costal vein; c—costal cell; dm—discal medial cell; h—humeral crossvein; M1—first branch of media; M4—fourth branch of media; m1— first medial cell; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023161" httpUri="https://zenodo.org/record/14023161/files/figure.png" pageId="38" pageNumber="39">Figs 5 H</figureCitation>
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,
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<figureCitation id="136284BB1E46FFD2FEB05326FE96FD45" box="[262,332,619,645]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStartId-0="39.[151,250,1458,1482]" captionStartId-1="40.[151,250,1264,1288]" captionTargetBox-0="[151,1436,187,1432]" captionTargetBox-1="[158,1434,187,1236]" captionTargetId-0="figure-20@39.[151,1436,181,1433]" captionTargetId-1="figure-30@40.[151,1436,181,1240]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionText-0="FIGURE 22. Aspistomella enderleini sp. nov. holotype ♂ (A, C–G) and paratype ♀ (B). A, B, habitus, left; C, head and thorax, dorsal; D, head left; E, same, anterior; F, wing, anterobasal part; G, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r 1—first radial cell; r 2+3—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for C, D, and E)." captionText-1="FIGURE 23. Aspistomella enderleini sp. nov. ♂ genitalia. A, postabdomen and part of abdomen, ventral; B, postabdomen, posterior; C, same, lateral; D, same, ventral; E, surstylus and hypandrium, posterior; F, cerci, surstylus, and hypandrium, ventral.Abbreviations: bph—basiphallus; cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; pgt—postgonite; ph g—phallic guide; ph—phallus; phapod—phallapodeme; pregt—pregonite; s—seta; st 4— fourth sternite; st 5— fifth sternite; st 6— sixth sternite; st 7— seventh sternite; st 8—eighth sternite; sur—surstylus; tg5—fifth tergite." figureDoi-0="http://doi.org/10.5281/zenodo.14023216" figureDoi-1="http://doi.org/10.5281/zenodo.14023218" httpUri-0="https://zenodo.org/record/14023216/files/figure.png" httpUri-1="https://zenodo.org/record/14023218/files/figure.png" pageId="38" pageNumber="39">22–23</figureCitation>
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</paragraph>
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<emphasis id="B92D442C1E46FFD2FF2153FFFE16FD0D" bold="true" box="[151,460,690,717]" pageId="38" pageNumber="39">Material examined. Type.</emphasis>
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<materialsCitation id="3B3192631E46FFD2FE6253FEFEF1FD31" collectionCode="MNKB" country="Colombia" location="Colombia" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54E2269C1E46FFD2FE6253FEFDE6FD0D" box="[468,572,691,717]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D5F53B71E46FFD2FDF353FEFD80FD0D" box="[581,602,691,717]" count="1" pageId="38" pageNumber="39" type="male">♂</specimenCount>
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:
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<collectingCountry id="F34ED8AE1E46FFD2FDDC53FFFD05FD0C" box="[618,735,690,716]" name="Colombia" pageId="38" pageNumber="39">Colombia</collectingCountry>
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</emphasis>
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: “Cordill.[eren] v.[on] Columbien / terra temporada / Thieme S.” (
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<collectionCode id="ED4800FB1E46FFD2FF7F539AFEF9FD31" box="[201,291,727,753]" pageId="38" pageNumber="39">MNKB</collectionCode>
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)
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.
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<materialsCitation id="3B3192631E46FFD2FE80539AFAFBFD31" box="[310,1313,727,753]" collectionCode="MNKB" country="Colombia" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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|
||
<typeStatus id="54E2269C1E46FFD2FE80539AFE79FD31" box="[310,419,727,753]" pageId="38" pageNumber="39" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D5F53B71E46FFD2FE07539AFE0CFD31" box="[433,470,727,753]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
|
||
: same label as in
|
||
<typeStatus id="54E2269C1E46FFD2FD2A539AFCDBFD31" box="[668,769,727,753]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
, “69”, “
|
||
<taxonomicName id="4C59E3BD1E46FFD2FCEF539AFBAEFD31" authority="Enderlein" authorityName="Enderlein" box="[857,1140,727,753]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lunata">Euxesta lunata Enderlein</taxonomicName>
|
||
det.” (
|
||
<collectionCode id="ED4800FB1E46FFD2FB09539AFAC3FD31" box="[1215,1305,727,753]" pageId="38" pageNumber="39">MNKB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343CBB51E46FFD2FF7153B7FB2FFAB9" pageId="38" pageNumber="39" type="diagnosis">
|
||
<paragraph id="8BE6983E1E46FFD2FF7153B7FB2FFAB9" blockId="38.[151,1437,690,1905]" pageId="38" pageNumber="39">
|
||
<emphasis id="B92D442C1E46FFD2FF7153B7FE98FCD4" bold="true" box="[199,322,762,788]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C59E3BD1E46FFD2FEFE53B6FD88FCD5" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[328,594,763,789]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="enderleini">
|
||
<emphasis id="B92D442C1E46FFD2FEFE53B6FD88FCD5" box="[328,594,763,789]" italics="true" pageId="38" pageNumber="39">Aspistomella enderleini</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C59E3BD1E46FFD2FD5253B6FCA2FCD5" authorityName="Hendel" authorityYear="1909" box="[740,888,763,789]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lobioptera">
|
||
<emphasis id="B92D442C1E46FFD2FD5253B6FCA2FCD5" box="[740,888,763,789]" italics="true" pageId="38" pageNumber="39">A. lobioptera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C59E3BD1E46FFD2FC3053B6FC37FCD5" box="[902,1005,763,789]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lunata">
|
||
<emphasis id="B92D442C1E46FFD2FC3053B6FC37FCD5" box="[902,1005,763,789]" italics="true" pageId="38" pageNumber="39">A. lunata</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C59E3BD1E46FFD2FB9C53B6FB70FCD5" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[1066,1194,763,789]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="steyskali">
|
||
<emphasis id="B92D442C1E46FFD2FB9C53B6FB70FCD5" box="[1066,1194,763,789]" italics="true" pageId="38" pageNumber="39">A. steyskali</emphasis>
|
||
</taxonomicName>
|
||
in that the wing apex has a long, narrow and oblique preapical hyaline incision from the apical half of cell r
|
||
<subScript id="17DD9A7B1E46FFD2FBE7527CFB83FCFF" attach="left" box="[1105,1113,817,831]" fontSize="6" pageId="38" pageNumber="39">1</subScript>
|
||
into cell m, usually reaching the anterior margin of the wing and separating a narrow crescent-shaped brown apical band from the rest of the dark pattern, with the base of cell m completely dark (
|
||
<figureCitation id="136284BB1E46FFD2FD40522AFC89FC41" box="[758,851,870,897]" captionStart="FIGURE 5" captionStartId="12.[151,250,1617,1641]" captionTargetBox="[175,1418,206,1566]" captionTargetId="figure-30@12.[151,1436,181,1593]" captionTargetPageId="12" captionText="FIGURE 5. Aspistomella, wing. A, A. pachitea sp. nov.; B, A. heteroptera Hendel; C, A. duo sp. nov.; D, A. tres sp. nov.; E, A. lobioptera Hendel; F, A. steyskali sp. nov.; G, A. lunata (Hendel); H, A. enderleini sp. nov. Abbreviations: C—costal vein; c—costal cell; dm—discal medial cell; h—humeral crossvein; M1—first branch of media; M4—fourth branch of media; m1— first medial cell; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023161" httpUri="https://zenodo.org/record/14023161/files/figure.png" pageId="38" pageNumber="39">Fig. 5 H</figureCitation>
|
||
), vein R
|
||
<subScript id="17DD9A7B1E46FFD2FC035234FC67FC47" attach="left" box="[949,957,889,903]" fontSize="6" pageId="38" pageNumber="39">1</subScript>
|
||
bare, prescutellar acrostichal setae present and halter yellow; it is similar to
|
||
<taxonomicName id="4C59E3BD1E46FFD2FD9652C6FD6DFC65" authorityName="Hendel" authorityYear="1909" box="[544,695,907,933]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lobioptera">
|
||
<emphasis id="B92D442C1E46FFD2FD9652C6FD6DFC65" box="[544,695,907,933]" italics="true" pageId="38" pageNumber="39">A. lobioptera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C59E3BD1E46FFD2FD4252C6FCACFC65" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[756,886,907,933]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="steyskali">
|
||
<emphasis id="B92D442C1E46FFD2FD4252C6FCACFC65" box="[756,886,907,933]" italics="true" pageId="38" pageNumber="39">A. steyskali</emphasis>
|
||
</taxonomicName>
|
||
in that the costal vein is bent anteriorly before the apex of vein Sc, the costal cell is lobed (though less than in the compared species), the costal vein is slightly thickened before the apex of the subcostal vein, forming a more or less conspicuous cleft, and differs from them in that the pterostigma is only moderately short, almost as long as the width of the costal cell (
|
||
<figureCitation id="136284BB1E46FFD2FB7D52BAFAF5FBD1" box="[1227,1327,1014,1041]" captionStart="FIGURE 5" captionStartId="12.[151,250,1617,1641]" captionTargetBox="[175,1418,206,1566]" captionTargetId="figure-30@12.[151,1436,181,1593]" captionTargetPageId="12" captionText="FIGURE 5. Aspistomella, wing. A, A. pachitea sp. nov.; B, A. heteroptera Hendel; C, A. duo sp. nov.; D, A. tres sp. nov.; E, A. lobioptera Hendel; F, A. steyskali sp. nov.; G, A. lunata (Hendel); H, A. enderleini sp. nov. Abbreviations: C—costal vein; c—costal cell; dm—discal medial cell; h—humeral crossvein; M1—first branch of media; M4—fourth branch of media; m1— first medial cell; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023161" httpUri="https://zenodo.org/record/14023161/files/figure.png" pageId="38" pageNumber="39">Figs 5 H</figureCitation>
|
||
,
|
||
<figureCitation id="136284BB1E46FFD2FA8952BAFAACFBD1" box="[1343,1398,1015,1041]" captionStart="FIGURE 22" captionStartId="39.[151,250,1458,1482]" captionTargetBox="[151,1436,187,1432]" captionTargetId="figure-20@39.[151,1436,181,1433]" captionTargetPageId="39" captionText="FIGURE 22. Aspistomella enderleini sp. nov. holotype ♂ (A, C–G) and paratype ♀ (B). A, B, habitus, left; C, head and thorax, dorsal; D, head left; E, same, anterior; F, wing, anterobasal part; G, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r 1—first radial cell; r 2+3—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for C, D, and E)." figureDoi="http://doi.org/10.5281/zenodo.14023216" httpUri="https://zenodo.org/record/14023216/files/figure.png" pageId="38" pageNumber="39">22 F</figureCitation>
|
||
). It shares the wing pattern very similar to
|
||
<taxonomicName id="4C59E3BD1E46FFD2FDE15556FD02FBF5" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[599,728,1051,1077]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="steyskali">
|
||
<emphasis id="B92D442C1E46FFD2FDE15556FD02FBF5" box="[599,728,1051,1077]" italics="true" pageId="38" pageNumber="39">A. steyskali</emphasis>
|
||
</taxonomicName>
|
||
: both differ from
|
||
<taxonomicName id="4C59E3BD1E46FFD2FC1E5556FBE4FBF5" authorityName="Hendel" authorityYear="1909" box="[936,1086,1051,1077]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lobioptera">
|
||
<emphasis id="B92D442C1E46FFD2FC1E5556FBE4FBF5" box="[936,1086,1051,1077]" italics="true" pageId="38" pageNumber="39">A. lobioptera</emphasis>
|
||
</taxonomicName>
|
||
by the cell dm apically of r-m with hyaline spot connected to the hyaline spot in cell m
|
||
<subScript id="17DD9A7B1E46FFD2FCAC551CFCF8FB9F" attach="left" box="[794,802,1105,1119]" fontSize="6" pageId="38" pageNumber="39">4</subScript>
|
||
and by the phallus not modified, moderately developed (in
|
||
<taxonomicName id="4C59E3BD1E46FFD2FF0B552EFE8BFBBD" authorityName="Hendel" authorityYear="1909" box="[189,337,1123,1149]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lobioptera">
|
||
<emphasis id="B92D442C1E46FFD2FF0B552EFE8BFBBD" box="[189,337,1123,1149]" italics="true" pageId="38" pageNumber="39">A. lobioptera</emphasis>
|
||
</taxonomicName>
|
||
cell dm entirely dark and phallus very long and trichose). It can be distinguished from
|
||
<taxonomicName id="4C59E3BD1E46FFD2FAAB552EFA46FBBD" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[1309,1436,1123,1149]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="steyskali">
|
||
<emphasis id="B92D442C1E46FFD2FAAB552EFA46FBBD" box="[1309,1436,1123,1149]" italics="true" pageId="38" pageNumber="39">A. steyskali</emphasis>
|
||
</taxonomicName>
|
||
only by the structure of surstyli, which are ventrally blunt, with short denticles on the posterior margin, but without expressed lobes and prensisetae, only with slightly thickened setae (in
|
||
<taxonomicName id="4C59E3BD1E46FFD2FC0955E6FB9AFB05" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[959,1088,1195,1221]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="steyskali">
|
||
<emphasis id="B92D442C1E46FFD2FC0955E6FB9AFB05" box="[959,1088,1195,1221]" italics="true" pageId="38" pageNumber="39">A. steyskali</emphasis>
|
||
</taxonomicName>
|
||
, outer surstylus with two well developed lobes, inner surstylus with 3–4 thick prensisetae); it also differs from
|
||
<taxonomicName id="4C59E3BD1E46FFD2FBB85582FB57FB29" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[1038,1165,1231,1257]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="steyskali">
|
||
<emphasis id="B92D442C1E46FFD2FBB85582FB57FB29" box="[1038,1165,1231,1257]" italics="true" pageId="38" pageNumber="39">A. steyskali</emphasis>
|
||
</taxonomicName>
|
||
by a pterostigma almost as long as wide (in
|
||
<taxonomicName id="4C59E3BD1E46FFD2FEC655BEFE35FACD" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[368,495,1267,1293]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="steyskali">
|
||
<emphasis id="B92D442C1E46FFD2FEC655BEFE35FACD" box="[368,495,1267,1293]" italics="true" pageId="38" pageNumber="39">A. steyskali</emphasis>
|
||
</taxonomicName>
|
||
, pterostigma very narrow); but the variability of this character has not been studied.
|
||
<taxonomicName id="4C59E3BD1E46FFD2FF21545AFE78FAF1" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[151,418,1303,1329]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="enderleini">
|
||
<emphasis id="B92D442C1E46FFD2FF21545AFE78FAF1" box="[151,418,1303,1329]" italics="true" pageId="38" pageNumber="39">Aspistomella enderleini</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C59E3BD1E46FFD2FD81545AFD45FAF1" box="[567,671,1303,1329]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lunata">
|
||
<emphasis id="B92D442C1E46FFD2FD81545AFD45FAF1" box="[567,671,1303,1329]" italics="true" pageId="38" pageNumber="39">A. lunata</emphasis>
|
||
</taxonomicName>
|
||
in having pterostigma almost as long as wide, but differs from it in that the hyaline incision distal to the pterostigma is short, extending at most into cell r
|
||
<subScript id="17DD9A7B1E46FFD2FBFD5400FBBEFA9B" attach="left" box="[1099,1124,1357,1371]" fontSize="6" pageId="38" pageNumber="39">2+3</subScript>
|
||
and isolated from the round hyaline spot at the base of cell r
|
||
<subScript id="17DD9A7B1E46FFD2FE4C543CFDCEFABF" attach="left" box="[506,532,1393,1407]" fontSize="6" pageId="38" pageNumber="39">4+5</subScript>
|
||
(in
|
||
<taxonomicName id="4C59E3BD1E46FFD2FDF75412FD72FAB9" box="[577,680,1375,1401]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lunata">
|
||
<emphasis id="B92D442C1E46FFD2FDF75412FD72FAB9" box="[577,680,1375,1401]" italics="true" pageId="38" pageNumber="39">A. lunata</emphasis>
|
||
</taxonomicName>
|
||
, it extends into cell r
|
||
<subScript id="17DD9A7B1E46FFD2FC22543CFC74FABF" attach="left" box="[916,942,1393,1407]" fontSize="6" pageId="38" pageNumber="39">4+5</subScript>
|
||
distally from crossvein r-m).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343CBB51E46FFDDFF7154CFFEDCFD9D" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39" type="description">
|
||
<paragraph id="8BE6983E1E46FFD2FF7154CFFEE3F8B1" blockId="38.[151,1437,690,1905]" pageId="38" pageNumber="39">
|
||
<emphasis id="B92D442C1E46FFD2FF7154CFFE8BFA5C" bold="true" box="[199,337,1410,1436]" pageId="38" pageNumber="39">Description</emphasis>
|
||
. Male.
|
||
<emphasis id="B92D442C1E46FFD2FE1254CEFE3EFA5D" bold="true" box="[420,484,1411,1437]" pageId="38" pageNumber="39">Head</emphasis>
|
||
(
|
||
<figureCitation id="136284BB1E46FFD2FE4754CEFDA5FA5D" box="[497,639,1411,1437]" captionStart="FIGURE 22" captionStartId="39.[151,250,1458,1482]" captionTargetBox="[151,1436,187,1432]" captionTargetId="figure-20@39.[151,1436,181,1433]" captionTargetPageId="39" captionText="FIGURE 22. Aspistomella enderleini sp. nov. holotype ♂ (A, C–G) and paratype ♀ (B). A, B, habitus, left; C, head and thorax, dorsal; D, head left; E, same, anterior; F, wing, anterobasal part; G, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r 1—first radial cell; r 2+3—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for C, D, and E)." figureDoi="http://doi.org/10.5281/zenodo.14023216" httpUri="https://zenodo.org/record/14023216/files/figure.png" pageId="38" pageNumber="39">Figs 22 D–E</figureCitation>
|
||
) ratio (length: height: width) = 1: 1.5: 1.9; frons, face above transverse fold, parafacial, and gena reddish or yellowish brown, epistome and occiput mostly black. Frons moderately wide, slightly narrowed posteriorly (
|
||
<figureCitation id="136284BB1E46FFD2FE445486FDB8FA25" box="[498,610,1483,1509]" captionStart="FIGURE 22" captionStartId="39.[151,250,1458,1482]" captionTargetBox="[151,1436,187,1432]" captionTargetId="figure-20@39.[151,1436,181,1433]" captionTargetPageId="39" captionText="FIGURE 22. Aspistomella enderleini sp. nov. holotype ♂ (A, C–G) and paratype ♀ (B). A, B, habitus, left; C, head and thorax, dorsal; D, head left; E, same, anterior; F, wing, anterobasal part; G, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r 1—first radial cell; r 2+3—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for C, D, and E)." figureDoi="http://doi.org/10.5281/zenodo.14023216" httpUri="https://zenodo.org/record/14023216/files/figure.png" pageId="38" pageNumber="39">Fig. 22 E</figureCitation>
|
||
), as long (from lunule to anterior ocellus) or 1.4× (from lunule to inner vertical seta) as wide at lunule; vertex black, steel shining, sparsely microtrichose; orbits, vertex, gena and occiput silver-white microtrichose; parafacial narrow, 0.2–0.3× broader than postpedicel, reddish-yellow, sparsely white microtrichose. Frontal plates with 8–9 pairs of parafrontal setulae, 3–5 pairs of inclinate frontal setae slightly shorter than outer vertical seta, frontal vitta whitish microtrichose, with 2–4 shorter, proclinate or inclinate interfrontal setulae (
|
||
<figureCitation id="136284BB1E46FFD2FF415732FEB2F959" box="[247,360,1663,1689]" captionStart="FIGURE 22" captionStartId="39.[151,250,1458,1482]" captionTargetBox="[151,1436,187,1432]" captionTargetId="figure-20@39.[151,1436,181,1433]" captionTargetPageId="39" captionText="FIGURE 22. Aspistomella enderleini sp. nov. holotype ♂ (A, C–G) and paratype ♀ (B). A, B, habitus, left; C, head and thorax, dorsal; D, head left; E, same, anterior; F, wing, anterobasal part; G, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r 1—first radial cell; r 2+3—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for C, D, and E)." figureDoi="http://doi.org/10.5281/zenodo.14023216" httpUri="https://zenodo.org/record/14023216/files/figure.png" pageId="38" pageNumber="39">Fig. 22 E</figureCitation>
|
||
). Lunule very narrow, indistinct, reddish-yellow. Eye 1.4× higher than long. Epistome widely black medially, reddish brown laterally and anteroventrally, sparsely white microtrichose, with metallic bluish sheen, strongly produced anteriorly and separated from upper part of face by deep fold; face above epistome brownish yellow, white microtrichose. Clypeus yellow to brown, sparsely white microtrichose, subshiny, 0.38× higher than epistome. Gena brownish yellow, with long genal seta and 5–7 additional long peristomal setae anterior of it, 0.3–0.8× longer than genal seta. Occiput black, except postgena partly brown, with cyan metallic sheen, greyish microtrichose.
|
||
</paragraph>
|
||
<caption id="DF26C8B61E47FFD3FF2154FFFAA9F9D1" ID-DOI="http://doi.org/10.5281/zenodo.14023216" ID-Zenodo-Dep="14023216" httpUri="https://zenodo.org/record/14023216/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1458,1482]" targetBox="[151,1436,187,1432]" targetPageId="39" targetType="figure">
|
||
<paragraph id="8BE6983E1E47FFD3FF2154FFFAA9F9D1" blockId="39.[151,1437,1458,1561]" pageId="39" pageNumber="40">
|
||
<emphasis id="B92D442C1E47FFD3FF2154FFFEC5FA0A" bold="true" box="[151,287,1458,1482]" pageId="39" pageNumber="40">FIGURE 22.</emphasis>
|
||
<taxonomicName id="4C59E3BD1E47FFD3FE9254FFFDCEFA0A" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[292,532,1458,1482]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="enderleini" status="sp. nov.">
|
||
<emphasis id="B92D442C1E47FFD3FE9254FFFDCEFA0A" box="[292,532,1458,1482]" italics="true" pageId="39" pageNumber="40">Aspistomella enderleini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92D442C1E47FFD3FDAF54FFFDB3FA0A" bold="true" box="[537,617,1458,1482]" pageId="39" pageNumber="40">
|
||
<taxonomicNameLabel id="A21EF9571E47FFD3FDAF54FFFDB3FA0A" box="[537,617,1458,1482]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype ♂ (
|
||
<emphasis id="B92D442C1E47FFD3FD5854FFFCDAFA0A" bold="true" box="[750,768,1458,1482]" pageId="39" pageNumber="40">A</emphasis>
|
||
,
|
||
<emphasis id="B92D442C1E47FFD3FCBA54FFFCE5FA0A" bold="true" box="[780,831,1458,1482]" pageId="39" pageNumber="40">C–G</emphasis>
|
||
) and paratype ♀ (
|
||
<emphasis id="B92D442C1E47FFD3FC4254FFFBDFFA0A" bold="true" box="[1012,1029,1458,1482]" pageId="39" pageNumber="40">B</emphasis>
|
||
).
|
||
<emphasis id="B92D442C1E47FFD3FBAF54FFFBF1FA0A" bold="true" box="[1049,1067,1458,1482]" pageId="39" pageNumber="40">A</emphasis>
|
||
,
|
||
<emphasis id="B92D442C1E47FFD3FB8E54FFFB93FA0A" bold="true" box="[1080,1097,1458,1482]" pageId="39" pageNumber="40">B</emphasis>
|
||
, habitus, left;
|
||
<emphasis id="B92D442C1E47FFD3FB6E54FFFB30FA0A" bold="true" box="[1240,1258,1458,1482]" pageId="39" pageNumber="40">C</emphasis>
|
||
, head and thorax, dorsal;
|
||
<emphasis id="B92D442C1E47FFD3FF55549BFF2FFA2E" bold="true" box="[227,245,1494,1518]" pageId="39" pageNumber="40">D</emphasis>
|
||
, head left;
|
||
<emphasis id="B92D442C1E47FFD3FEDC549BFEA1FA2E" bold="true" box="[362,379,1494,1518]" pageId="39" pageNumber="40">E</emphasis>
|
||
, same, anterior;
|
||
<emphasis id="B92D442C1E47FFD3FD91549BFDECFA2E" bold="true" box="[551,566,1494,1518]" pageId="39" pageNumber="40">F</emphasis>
|
||
, wing, anterobasal part;
|
||
<emphasis id="B92D442C1E47FFD3FC82549BFC92FA2E" bold="true" box="[820,840,1494,1518]" pageId="39" pageNumber="40">G</emphasis>
|
||
, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r
|
||
<subScript id="17DD9A7B1E47FFD3FF165747FF7DF9D8" attach="right" box="[160,167,1546,1560]" fontSize="6" pageId="39" pageNumber="40">1</subScript>
|
||
—first radial cell; r
|
||
<subScript id="17DD9A7B1E47FFD3FEDC5747FE5BF9D8" attach="right" box="[362,385,1546,1560]" fontSize="6" pageId="39" pageNumber="40">2+3</subScript>
|
||
—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for
|
||
<emphasis id="B92D442C1E47FFD3FB5D54B7FB27F9D2" bold="true" box="[1259,1277,1530,1554]" pageId="39" pageNumber="40">C</emphasis>
|
||
,
|
||
<emphasis id="B92D442C1E47FFD3FABC54B7FAC6F9D2" bold="true" box="[1290,1308,1530,1554]" pageId="39" pageNumber="40">D</emphasis>
|
||
, and
|
||
<emphasis id="B92D442C1E47FFD3FAE254B4FAB7F9D1" bold="true" box="[1364,1389,1529,1553]" pageId="39" pageNumber="40">E)</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE6983E1E47FFD3FD1A5746FD19F9D9" blockId="39.[151,1437,1458,1561]" box="[684,707,1547,1561]" pageId="39" pageNumber="40">
|
||
<subScript id="17DD9A7B1E47FFD3FD1A5746FD19F9D9" attach="left" box="[684,707,1547,1561]" fontSize="6" pageId="39" pageNumber="40">4+5</subScript>
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E47FFD3FF71571AFF37F8C0" blockId="39.[151,1437,1623,2042]" pageId="39" pageNumber="40">Antenna brown; scape and pedicel with black setulae; postpedicel yellowish brown, 1.6× longer than wide, darker apico-dorsally, whitish microtrichose, twice as long as wide, apically rounded; arista black except yellow at base, almost bare. Mouthparts black, prementum black, shiny. Palp yellow or yellowish brown, narrowly crescent-shaped, apically rounded, with 10–12 strong black setulae, of which 3–4 subapical setulae 1.6–2× longer than other setulae.</paragraph>
|
||
<paragraph id="8BE6983E1E47FFD3FF715644FEBAF83A" blockId="39.[151,1437,1623,2042]" pageId="39" pageNumber="40">
|
||
<emphasis id="B92D442C1E47FFD3FF715644FEFAF8E3" bold="true" box="[199,288,1801,1827]" pageId="39" pageNumber="40">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="136284BB1E47FFD3FE865644FE46F8E3" box="[304,412,1801,1827]" captionStart="FIGURE 22" captionStartId="39.[151,250,1458,1482]" captionTargetBox="[151,1436,187,1432]" captionTargetId="figure-20@39.[151,1436,181,1433]" captionTargetPageId="39" captionText="FIGURE 22. Aspistomella enderleini sp. nov. holotype ♂ (A, C–G) and paratype ♀ (B). A, B, habitus, left; C, head and thorax, dorsal; D, head left; E, same, anterior; F, wing, anterobasal part; G, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r 1—first radial cell; r 2+3—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for C, D, and E)." figureDoi="http://doi.org/10.5281/zenodo.14023216" httpUri="https://zenodo.org/record/14023216/files/figure.png" pageId="39" pageNumber="40">Fig. 22 C</figureCitation>
|
||
) brown to black, with bluish sheen and sparse white and brown microtrichia not obscuring underlying cuticle. Mesonotal scutum 1.2× longer than wide; black setulose, with 6–12 irregular rows of setulae between rows of dorsocentral setulae; acrostichal prescutellar seta present; prescutellar area with 4–6 rows of setulae between posterior dorsocentral setae. Scutellum dorsally very slightly convex, brown to black, subshining, sparsely microtrichose, without setulae, with golden, silver or bluish sheen. Subscutellum shiny, dark brown to black. Mediotergite shiny black, medially not microtrichose. Other setae as described for the genus. All the setae and setulae black.
|
||
</paragraph>
|
||
<caption id="DF26C8B61E48FFDCFF2155BDFAE4FA58" ID-DOI="http://doi.org/10.5281/zenodo.14023218" ID-Zenodo-Dep="14023218" httpUri="https://zenodo.org/record/14023218/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1264,1288]" targetBox="[158,1434,187,1236]" targetPageId="40" targetType="figure">
|
||
<paragraph id="8BE6983E1E48FFDCFF2155BDFAE4FA58" blockId="40.[151,1437,1264,1432]" pageId="40" pageNumber="41">
|
||
<emphasis id="B92D442C1E48FFDCFF2155BDFEF8FAC8" bold="true" box="[151,290,1264,1288]" pageId="40" pageNumber="41">FIGURE 23.</emphasis>
|
||
<taxonomicName id="4C59E3BD1E48FFDCFE9C55BDFDC7FAC8" authorityName="Kovac & Kameneva & Korneyev & Araújo & Savaris & Smit & Schneider & Schreiber & Korneyev" authorityYear="2024" box="[298,541,1264,1288]" class="Insecta" family="Ulidiidae" genus="Aspistomella" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="enderleini" status="sp. nov.">
|
||
<emphasis id="B92D442C1E48FFDCFE9C55BDFDC7FAC8" box="[298,541,1264,1288]" italics="true" pageId="40" pageNumber="41">Aspistomella enderleini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92D442C1E48FFDCFD9355BCFDA2FAC9" bold="true" box="[549,632,1265,1289]" pageId="40" pageNumber="41">
|
||
<taxonomicNameLabel id="A21EF9571E48FFDCFD9355BCFDA2FAC9" box="[549,632,1265,1289]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂ genitalia.
|
||
<emphasis id="B92D442C1E48FFDCFCB455BDFCCEFAC8" bold="true" box="[770,788,1264,1288]" pageId="40" pageNumber="41">A</emphasis>
|
||
, postabdomen and part of abdomen, ventral;
|
||
<emphasis id="B92D442C1E48FFDCFB5955BCFADAFAC9" bold="true" box="[1263,1280,1265,1289]" pageId="40" pageNumber="41">B</emphasis>
|
||
, postabdomen, posterior;
|
||
<emphasis id="B92D442C1E48FFDCFEB45459FECEFAEC" bold="true" box="[258,276,1300,1324]" pageId="40" pageNumber="41">C</emphasis>
|
||
, same, lateral;
|
||
<emphasis id="B92D442C1E48FFDCFE0F5458FE11FAED" bold="true" box="[441,459,1301,1325]" pageId="40" pageNumber="41">D</emphasis>
|
||
, same, ventral;
|
||
<emphasis id="B92D442C1E48FFDCFDCE5458FD53FAED" bold="true" box="[632,649,1301,1325]" pageId="40" pageNumber="41">E</emphasis>
|
||
, surstylus and hypandrium, posterior;
|
||
<emphasis id="B92D442C1E48FFDCFB975458FBEAFAED" bold="true" box="[1057,1072,1301,1325]" pageId="40" pageNumber="41">F</emphasis>
|
||
, cerci, surstylus, and hypandrium, ventral. Abbreviations: bph—basiphallus; cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; pgt—postgonite; ph g—phallic guide; ph—phallus; phapod—phallapodeme; pregt—pregonite; s—seta; st 4— fourth sternite; st 5— fifth sternite; st 6— sixth sternite; st 7— seventh sternite; st 8—eighth sternite; sur—surstylus; tg5—fifth tergite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE6983E1E48FFDCFF7154A7FB15F815" blockId="40.[151,1437,1514,2006]" pageId="40" pageNumber="41">
|
||
<emphasis id="B92D442C1E48FFDCFF7154A7FED2F9C4" bold="true" box="[199,264,1514,1540]" pageId="40" pageNumber="41">Wing</emphasis>
|
||
(
|
||
<figureCitation id="136284BB1E48FFDCFEAE54A7FEA5F9C5" box="[280,383,1514,1541]" captionStart="FIGURE 5" captionStartId="12.[151,250,1617,1641]" captionTargetBox="[175,1418,206,1566]" captionTargetId="figure-30@12.[151,1436,181,1593]" captionTargetPageId="12" captionText="FIGURE 5. Aspistomella, wing. A, A. pachitea sp. nov.; B, A. heteroptera Hendel; C, A. duo sp. nov.; D, A. tres sp. nov.; E, A. lobioptera Hendel; F, A. steyskali sp. nov.; G, A. lunata (Hendel); H, A. enderleini sp. nov. Abbreviations: C—costal vein; c—costal cell; dm—discal medial cell; h—humeral crossvein; M1—first branch of media; M4—fourth branch of media; m1— first medial cell; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023161" httpUri="https://zenodo.org/record/14023161/files/figure.png" pageId="40" pageNumber="41">Figs 5 H</figureCitation>
|
||
,
|
||
<figureCitation id="136284BB1E48FFDCFE3A54A6FDDFF9C5" box="[396,517,1515,1541]" captionStart="FIGURE 22" captionStartId="39.[151,250,1458,1482]" captionTargetBox="[151,1436,187,1432]" captionTargetId="figure-20@39.[151,1436,181,1433]" captionTargetPageId="39" captionText="FIGURE 22. Aspistomella enderleini sp. nov. holotype ♂ (A, C–G) and paratype ♀ (B). A, B, habitus, left; C, head and thorax, dorsal; D, head left; E, same, anterior; F, wing, anterobasal part; G, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r 1—first radial cell; r 2+3—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for C, D, and E)." figureDoi="http://doi.org/10.5281/zenodo.14023216" httpUri="https://zenodo.org/record/14023216/files/figure.png" pageId="40" pageNumber="41">22 A, B, F</figureCitation>
|
||
)
|
||
<quantity id="4CA135DB1E48FFDCFDA354A7FD47F9C5" box="[533,669,1514,1541]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.2" metricValueMin="3.5" pageId="40" pageNumber="41" unit="mm" value="3.85" valueMax="4.2" valueMin="3.5">3.5–4.2 mm</quantity>
|
||
(
|
||
<specimenCount id="9D5F53B71E48FFDCFD0654A6FD18F9C5" box="[688,706,1515,1541]" count="1" pageId="40" pageNumber="41" type="female">♀</specimenCount>
|
||
) long, 2.4–2.5× longer than wide; basicostal cell hyaline; costal cell lobed, 5× longer than wide, brown in basal and apical quarter of length, with costa conspicuously curved, posteriorly covered with longer setulae and forming conspicuous cleft before apex of vein Sc; pterostigma entirely brown, narrow triangular, 1.3–1.4× longer than wide and as long as costal cell width, vein R
|
||
<subScript id="17DD9A7B1E48FFDCFB305725FB54F9B6" attach="left" box="[1158,1166,1640,1654]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
|
||
bare, ending proximally of crossvein r-m level in both sexes; vein R
|
||
<subScript id="17DD9A7B1E48FFDCFD3757C1FD41F95A" attach="left" box="[641,667,1676,1690]" fontSize="6" pageId="40" pageNumber="41">2+3</subScript>
|
||
slightly sinuate. Crossvein r-m distally of middle (at 0.6) of cell dm width. Cell r
|
||
<subScript id="17DD9A7B1E48FFDCFE9157E2FE9BF97D" attach="left" box="[295,321,1711,1725]" fontSize="6" pageId="40" pageNumber="41">4+5</subScript>
|
||
5× longer than wide, apically narrowed. Cell m
|
||
<subScript id="17DD9A7B1E48FFDCFCD057FDFCB4F97E" attach="left" box="[870,878,1712,1726]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
|
||
subtriangular; ultimate section of M
|
||
<subScript id="17DD9A7B1E48FFDCFAB957FDFACDF97E" attach="left" box="[1295,1303,1712,1726]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
|
||
= 2.4–2.8× longer than crossvein dm-m and 1.5× longer than penultimate section. Vein CuA Z-shaped, forming moderately short posteroapical lobe of cell cua along vein CuP, 1.7–1.9× longer than its anterior shoulder. Wing pattern with short subbasal crossband from humeral vein through cell cua into alula, and large dark brown mark from apical part of costal cell to wing apex, with 4 hyaline marks: incision from cell r
|
||
<subScript id="17DD9A7B1E48FFDCFC115672FC75F88D" attach="left" box="[935,943,1855,1869]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
|
||
into cell r
|
||
<subScript id="17DD9A7B1E48FFDCFB925672FBE7F88D" attach="left" box="[1060,1085,1855,1869]" fontSize="6" pageId="40" pageNumber="41">2+3</subScript>
|
||
basal to of crossvein r-m, oval spot in base of cell r
|
||
<subScript id="17DD9A7B1E48FFDCFEC9562FFE43F8B0" attach="left" box="[383,409,1890,1904]" fontSize="6" pageId="40" pageNumber="41">4+5</subScript>
|
||
and hyaline incision in apex of cell m
|
||
<subScript id="17DD9A7B1E48FFDCFCFB562FFC8FF8B0" attach="left" box="[845,853,1890,1904]" fontSize="6" pageId="40" pageNumber="41">4</subScript>
|
||
extending into apical half of cell dm, and long and narrow crescent-shaped or arcuate incision from apex of cell m
|
||
<subScript id="17DD9A7B1E48FFDCFCD056CBFCB4F854" attach="left" box="[870,878,1926,1940]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
|
||
into cell r
|
||
<subScript id="17DD9A7B1E48FFDCFC5256CBFC36F854" attach="left" box="[996,1004,1926,1940]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
|
||
parallel to wing apex (
|
||
<figureCitation id="136284BB1E48FFDCFB43563EFA8FF84E" box="[1269,1365,1907,1934]" captionStart="FIGURE 5" captionStartId="12.[151,250,1617,1641]" captionTargetBox="[175,1418,206,1566]" captionTargetId="figure-30@12.[151,1436,181,1593]" captionTargetPageId="12" captionText="FIGURE 5. Aspistomella, wing. A, A. pachitea sp. nov.; B, A. heteroptera Hendel; C, A. duo sp. nov.; D, A. tres sp. nov.; E, A. lobioptera Hendel; F, A. steyskali sp. nov.; G, A. lunata (Hendel); H, A. enderleini sp. nov. Abbreviations: C—costal vein; c—costal cell; dm—discal medial cell; h—humeral crossvein; M1—first branch of media; M4—fourth branch of media; m1— first medial cell; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023161" httpUri="https://zenodo.org/record/14023161/files/figure.png" pageId="40" pageNumber="41">Fig. 5 H</figureCitation>
|
||
). Cell cup (anal cell sensu
|
||
<bibRefCitation id="EFC8E5CF1E48FFDCFEC856D5FD16F871" author="Kameneva, E. & Korneyev, V." box="[382,716,1943,1970]" pageId="40" pageNumber="41" refId="ref75381" refString="Kameneva, E. & Korneyev, V. (2010) 66. Ulidiidae (picture-winged flies). In: Brown, B. V., Borkent, A., Wood, D. M. & Zumbado, M. (Eds.), Manual of Central American Diptera. Vol. 2. NRC, Ottawa, pp. 883 - 904." type="book" year="2010">Kameneva & Korneyev 2010</bibRefCitation>
|
||
) and anal lobe as wide as cell cua. Alula twice as long as wide, uniformly pale grey or darkened in apical half. Calypters white, with white ciliae. Halter yellow.
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E49FFDDFF7151DAFF28FEDC" blockId="41.[151,1437,151,713]" pageId="41" pageNumber="42">
|
||
<emphasis id="B92D442C1E49FFDDFF7151DAFF25FF71" bold="true" box="[199,255,151,177]" pageId="41" pageNumber="42">Legs</emphasis>
|
||
(
|
||
<figureCitation id="136284BB1E49FFDDFEB951DAFE7CFF71" box="[271,422,151,177]" captionStart="FIGURE 22" captionStartId="39.[151,250,1458,1482]" captionTargetBox="[151,1436,187,1432]" captionTargetId="figure-20@39.[151,1436,181,1433]" captionTargetPageId="39" captionText="FIGURE 22. Aspistomella enderleini sp. nov. holotype ♂ (A, C–G) and paratype ♀ (B). A, B, habitus, left; C, head and thorax, dorsal; D, head left; E, same, anterior; F, wing, anterobasal part; G, labels. Abbreviations: c—costal cell; ptstg—pterostigma; r 1—first radial cell; r 2+3—second + third radial cell; r —fourth + fifth radial cell. Scale bar: 1 mm (same for C, D, and E)." figureDoi="http://doi.org/10.5281/zenodo.14023216" httpUri="https://zenodo.org/record/14023216/files/figure.png" pageId="41" pageNumber="42">Figs 22 B, C</figureCitation>
|
||
): coxae and trochanters, apices of femora and tibiae brown, fore femur yellowish brown, otherwise black; tibiae mostly black; tarsi brown, black setose and setulose. Fore femur with row of 6–7 posterodorsal and 4–5 posteroventral setae. Mid femur anteriorly with short suberect setulae; mid tibia apically not visible, hidden by glue.
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E49FFDDFF715068FAFDFEFF" blockId="41.[151,1437,151,713]" box="[199,1319,293,320]" pageId="41" pageNumber="42">
|
||
<emphasis id="B92D442C1E49FFDDFF715068FEE6FEFF" bold="true" box="[199,316,293,319]" pageId="41" pageNumber="42">Abdomen</emphasis>
|
||
brown to black, sparsely black setulose, metallic sheen inconspicuous in
|
||
<typeStatus id="54E2269C1E49FFDDFBC5506BFB79FE80" box="[1139,1187,294,320]" pageId="41" pageNumber="42">type</typeStatus>
|
||
specimens.
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E49FFDDFF715004FA83FDD6" blockId="41.[151,1437,151,713]" pageId="41" pageNumber="42">
|
||
Male genitalia (
|
||
<figureCitation id="136284BB1E49FFDDFEC85004FDC9FEA3" box="[382,531,329,355]" captionStart="FIGURE 23" captionStartId="40.[151,250,1264,1288]" captionTargetBox="[158,1434,187,1236]" captionTargetId="figure-30@40.[151,1436,181,1240]" captionTargetPageId="40" captionText="FIGURE 23. Aspistomella enderleini sp. nov. ♂ genitalia. A, postabdomen and part of abdomen, ventral; B, postabdomen, posterior; C, same, lateral; D, same, ventral; E, surstylus and hypandrium, posterior; F, cerci, surstylus, and hypandrium, ventral.Abbreviations: bph—basiphallus; cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; pgt—postgonite; ph g—phallic guide; ph—phallus; phapod—phallapodeme; pregt—pregonite; s—seta; st 4— fourth sternite; st 5— fifth sternite; st 6— sixth sternite; st 7— seventh sternite; st 8—eighth sternite; sur—surstylus; tg5—fifth tergite." figureDoi="http://doi.org/10.5281/zenodo.14023218" httpUri="https://zenodo.org/record/14023218/files/figure.png" pageId="41" pageNumber="42">Figs 23 A–F</figureCitation>
|
||
) with short non-projecting cerci, wide ventrally blunt surstyli, with two short denticles on posterior margin, without expressed lobes and prensisetae, 3–5 unmodified setulae (one on inner surstylus, of which one twice as long and thick as the others; phallus moderately large, at most twice as long as epandrium height, short and sparsely trichose (
|
||
<figureCitation id="136284BB1E49FFDDFD1D50F9FC99FE0F" box="[683,835,436,463]" captionStart="FIGURE 23" captionStartId="40.[151,250,1264,1288]" captionTargetBox="[158,1434,187,1236]" captionTargetId="figure-30@40.[151,1436,181,1240]" captionTargetPageId="40" captionText="FIGURE 23. Aspistomella enderleini sp. nov. ♂ genitalia. A, postabdomen and part of abdomen, ventral; B, postabdomen, posterior; C, same, lateral; D, same, ventral; E, surstylus and hypandrium, posterior; F, cerci, surstylus, and hypandrium, ventral.Abbreviations: bph—basiphallus; cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; pgt—postgonite; ph g—phallic guide; ph—phallus; phapod—phallapodeme; pregt—pregonite; s—seta; st 4— fourth sternite; st 5— fifth sternite; st 6— sixth sternite; st 7— seventh sternite; st 8—eighth sternite; sur—surstylus; tg5—fifth tergite." figureDoi="http://doi.org/10.5281/zenodo.14023218" httpUri="https://zenodo.org/record/14023218/files/figure.png" pageId="41" pageNumber="42">Figs 23 B–D</figureCitation>
|
||
). Hypandrium: phallic guide with laterally triangular and moderately sclerotised lobes (
|
||
<figureCitation id="136284BB1E49FFDDFDA95095FD6EFE32" box="[543,692,472,498]" captionStart="FIGURE 23" captionStartId="40.[151,250,1264,1288]" captionTargetBox="[158,1434,187,1236]" captionTargetId="figure-30@40.[151,1436,181,1240]" captionTargetPageId="40" captionText="FIGURE 23. Aspistomella enderleini sp. nov. ♂ genitalia. A, postabdomen and part of abdomen, ventral; B, postabdomen, posterior; C, same, lateral; D, same, ventral; E, surstylus and hypandrium, posterior; F, cerci, surstylus, and hypandrium, ventral.Abbreviations: bph—basiphallus; cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; pgt—postgonite; ph g—phallic guide; ph—phallus; phapod—phallapodeme; pregt—pregonite; s—seta; st 4— fourth sternite; st 5— fifth sternite; st 6— sixth sternite; st 7— seventh sternite; st 8—eighth sternite; sur—surstylus; tg5—fifth tergite." figureDoi="http://doi.org/10.5281/zenodo.14023218" httpUri="https://zenodo.org/record/14023218/files/figure.png" pageId="41" pageNumber="42">Figs 23 D–F</figureCitation>
|
||
) forming conspicuous posterior projections, bearing pair of oval postgonites just anterior to basiphallus ring, and symmetrical pregonites anterolateral to them (
|
||
<figureCitation id="136284BB1E49FFDDFB0D50B1FA96FDD6" box="[1211,1356,508,535]" captionStart="FIGURE 23" captionStartId="40.[151,250,1264,1288]" captionTargetBox="[158,1434,187,1236]" captionTargetId="figure-30@40.[151,1436,181,1240]" captionTargetPageId="40" captionText="FIGURE 23. Aspistomella enderleini sp. nov. ♂ genitalia. A, postabdomen and part of abdomen, ventral; B, postabdomen, posterior; C, same, lateral; D, same, ventral; E, surstylus and hypandrium, posterior; F, cerci, surstylus, and hypandrium, ventral.Abbreviations: bph—basiphallus; cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; pgt—postgonite; ph g—phallic guide; ph—phallus; phapod—phallapodeme; pregt—pregonite; s—seta; st 4— fourth sternite; st 5— fifth sternite; st 6— sixth sternite; st 7— seventh sternite; st 8—eighth sternite; sur—surstylus; tg5—fifth tergite." figureDoi="http://doi.org/10.5281/zenodo.14023218" httpUri="https://zenodo.org/record/14023218/files/figure.png" pageId="41" pageNumber="42">Figs 23 D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE6983E1E49FFDDFF71536DFEDCFD9D" blockId="41.[151,1437,151,713]" pageId="41" pageNumber="42">
|
||
<emphasis id="B92D442C1E49FFDDFF71536DFE7FFDF9" bold="true" box="[199,421,543,570]" pageId="41" pageNumber="42">Female terminalia</emphasis>
|
||
: oviscape 1.3× longer than tergite 5; aculeus with ovoid, long-setulose cercal unit; not dissected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343CBB51E49FFDDFF71532AFB8AFD65" pageId="41" pageNumber="42" type="etymology">
|
||
<paragraph id="8BE6983E1E49FFDDFF71532AFB8AFD65" blockId="41.[151,1437,151,713]" pageId="41" pageNumber="42">
|
||
<emphasis id="B92D442C1E49FFDDFF71532AFE92FD41" bold="true" box="[199,328,615,641]" pageId="41" pageNumber="42">Etymology</emphasis>
|
||
. The species is named after the German entomologist and microbiologist Prof. Günter Enderlein, curator of the
|
||
<taxonomicName id="4C59E3BD1E49FFDDFE8353C6FE53FD65" box="[309,393,651,677]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
|
||
collection of the Humboldt University of
|
||
<collectingRegion id="499D56DC1E49FFDDFCD753C6FC7DFD65" box="[865,935,651,677]" country="Germany" name="Berlin" pageId="41" pageNumber="42">Berlin</collectingRegion>
|
||
in 1907–1942.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343CBB51E49FFDDFF7153E2FA43FD09" box="[199,1433,687,713]" pageId="41" pageNumber="42" type="discussion">
|
||
<paragraph id="8BE6983E1E49FFDDFF7153E2FA43FD09" blockId="41.[151,1437,151,713]" box="[199,1433,687,713]" pageId="41" pageNumber="42">
|
||
<emphasis id="B92D442C1E49FFDDFF7153E2FEEEFD09" bold="true" box="[199,308,687,713]" pageId="41" pageNumber="42">Remarks</emphasis>
|
||
. The
|
||
<typeStatus id="54E2269C1E49FFDDFEC253E2FE7EFD09" box="[372,420,687,713]" pageId="41" pageNumber="42">type</typeStatus>
|
||
specimens were misidentified by Enderlein as
|
||
<emphasis id="B92D442C1E49FFDDFC0253E2FBA0FD09" box="[948,1146,687,713]" italics="true" pageId="41" pageNumber="42">
|
||
“
|
||
<taxonomicName id="4C59E3BD1E49FFDDFC0853E2FBAAFD09" box="[958,1136,687,713]" class="Insecta" family="Ulidiidae" genus="Euxesta" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="lunata">Euxesta lunata</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
in the MNKB collection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |