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<mods:title id="50E17889A9FD374819E6530803EA99AE">Revision of the Aspistomella group of genera (Diptera: Ulidiidae: Pterocallinae: Lipsanini)</mods:title>
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<mods:affiliation id="3C6BBAABF3AFE0B306EF1089DC8929E3">I. I. Schmalhausen Institute of Zoology National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15, 01054 Kyiv, Ukraine https: // orcid. org / 0000 - 0002 - 7986 - 9942 &amp; Museum für Naturkunde, Invalidenstr. 43, Berlin, 10115 Germany Corresponding author: valery. korneyev @ gmail. com; https: // orcid. org / 0000 - 0001 - 9631 - 1038</mods:affiliation>
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<figureCitation id="136284BB1E3EFFAAFF215748FEC9F9E0" box="[151,275,1541,1568]" captionStart="FIGURE 2" captionStartId="7.[151,250,1936,1960]" captionTargetBox="[164,1423,181,1904]" captionTargetId="figure-20@7.[164,1423,181,1912]" captionTargetPageId="7" captionText="FIGURE 2.Adults of bamboo-inhabiting Ulidiidae from Peru.AC, Aspistomella lobioptera on a bamboo culm; D, Aspistomella sachavaca sp. nov.; EF, Ulivellia laetitiae sp. nov. on a bamboo culm showing wings during courtship or territorial behaviour. (All photographs by John T. Smit.)" figureDoi="http://doi.org/10.5281/zenodo.14023153" httpUri="https://zenodo.org/record/14023153/files/figure.png" pageId="94" pageNumber="95">Figs 2 E, F</figureCitation>
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<figureCitation id="136284BB1E3EFFAAFEA85748FE92F9DF" box="[286,328,1541,1567]" captionStart="FIGURE 4" captionStartId="11.[151,250,1686,1710]" captionTargetBox="[247,1341,187,1657]" captionTargetId="figure-20@11.[241,1346,181,1662]" captionTargetPageId="11" captionText="FIGURE 4. Ulivellia, Polyteloptera and similar genera, wing. A, U. amnoni sp. nov.; B, U. inversa Speiser; C, U. laetitiae sp. nov.; DE, U. arcuata sp. nov. (D—male, E—female); F, U. tenoris sp. nov.; G, U. pseudinsolita sp. nov.; H, P. apotropa Hendel; I, “Euxesta” insolita Hendel; J, Gen. sp. near insolita; K, Axiologina ferrumequinum Hendel. Abbreviations. Veins and cells: bm—basal medial cell; br—basal radial cell; c—costal cell; C—costal vein; cua—anterior cubital cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; h—humeral crossvein; M1—first branch of media; m1—first medial cell; M4—fourth branch of media; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; r2+3—second + third radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell; r-m—radial-medial crossvein; Sc—subcostal vein. Wing pattern: DB = discal band; HB = humeral band; AAB = anterior apical band; SAB = subapical band; SCB = subcostal band. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023159" httpUri="https://zenodo.org/record/14023159/files/figure.png" pageId="94" pageNumber="95">4 C</figureCitation>
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<figureCitation id="136284BB1E3EFFAAFEE25748FE43F9E0" box="[340,409,1541,1568]" captionStart-0="FIGURE 53" captionStart-1="FIGURE 54" captionStartId-0="95.[151,250,1514,1538]" captionStartId-1="97.[151,250,1457,1481]" captionTargetBox-0="[151,1436,181,1490]" captionTargetBox-1="[151,1432,182,1433]" captionTargetId-0="figure-20@95.[151,1436,181,1490]" captionTargetId-1="figure-20@97.[151,1435,181,1433]" captionTargetPageId-0="95" captionTargetPageId-1="97" captionText-0="FIGURE 53. Ulivellia laetitiae sp. nov. holotype ♀ (A, CE) and paratype ♂ (B, F, G). A, habitus, left; B, same, dorsal; C, head, mesonotum and fore leg, lateral; D, head, anterior; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. (A— photograph by John T. Smit)" captionText-1="FIGURE 54. Ulivellia laetitiae sp. nov. paratype ♂ (SIZK) A, postabdomen, lateral; B, same, anteroventral; C, same, ventral; D, epandrium, with cerci and surstyli, posterolateral; E, surstyli and hypandrium, ventral. Abbreviations: cerc—cerci; epand— epandrium; ph g—phallic guide; ph—phallus; phapd—phallapodeme; pi p—pimple-like process; pregt—pregonite; sur— surstylus; v l sur—ventral lobe of surstylus." figureDoi-0="http://doi.org/10.5281/zenodo.14023306" figureDoi-1="http://doi.org/10.5281/zenodo.14023308" httpUri-0="https://zenodo.org/record/14023306/files/figure.png" httpUri-1="https://zenodo.org/record/14023308/files/figure.png" pageId="94" pageNumber="95">5354</figureCitation>
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/
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<collectingDate id="EFA347161E3EFFAAFEAA573CFE3BF94B" box="[284,481,1649,1675]" pageId="94" pageNumber="95" value="2008-10-29">29 October 2008</collectingDate>
</date>
leg.
<collectorName id="26ACFDE81E3EFFAAFD97573FFD56F94B" box="[545,652,1649,1676]" pageId="94" pageNumber="95">J.T. Smit</collectorName>
” “RMNH.INS.555078 [including QR-code]”, “on bamboo
<taxonomicName id="4C59E3BD1E3EFFAAFAF4573CFE7AF96F" authority="&quot; (RMNH)" baseAuthorityName="RMNH" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B92D442C1E3EFFAAFAF4573CFEFFF96F" italics="true" pageId="94" pageNumber="95">Guadua weberbaueri</emphasis>
” (RMNH)
</taxonomicName>
.
<collectorName id="26ACFDE81E3EFFAAFE1A57D8FE02F96F" box="[428,472,1685,1711]" pageId="94" pageNumber="95">The</collectorName>
<typeStatus id="54E2269C1E3EFFAAFE6957D8FD9BF96F" box="[479,577,1685,1711]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
is in good condition.
</materialsCitation>
</paragraph>
<paragraph id="8BE6983E1E3EFFAAFF7157F4FDA5F833" blockId="94.[151,1436,1613,2036]" pageId="94" pageNumber="95">
<materialsCitation id="3B3192631E3EFFAAFF7157F4FAD4F914" box="[199,1294,1721,1748]" country="Peru" elevation="350" latitude="-12.656388" location="Pantiacolla" longLatPrecision="21" longitude="-71.23111" municipality="Manu" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Madre de Dios" typeStatus="paratype">
<emphasis id="B92D442C1E3EFFAAFF7157F4FEE5F913" bold="true" box="[199,319,1721,1747]" pageId="94" pageNumber="95">
<typeStatus id="54E2269C1E3EFFAAFF7157F4FEE5F913" box="[199,319,1721,1747]" pageId="94" pageNumber="95" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D5F53B71E3EFFAAFEF857F4FEABF913" box="[334,369,1721,1747]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
,
<emphasis id="B92D442C1E3EFFAAFE3657F4FE60F913" bold="true" box="[384,442,1721,1747]" pageId="94" pageNumber="95">
<collectingCountry id="F34ED8AE1E3EFFAAFE3657F4FE60F913" box="[384,442,1721,1747]" name="Peru" pageId="94" pageNumber="95">Peru</collectingCountry>
</emphasis>
:
<collectingRegion id="499D56DC1E3EFFAAFE7F57F4FDAEF913" box="[457,628,1721,1747]" country="Peru" name="Madre de Dios" pageId="94" pageNumber="95">Madre de Dios</collectingRegion>
,
<collectingMunicipality id="6B8202441E3EFFAAFDC957F7FD19F914" box="[639,707,1722,1748]" pageId="94" pageNumber="95">Manu</collectingMunicipality>
,
<location id="8E86CEE51E3EFFAAFD7957F4FC91F913" LSID="urn:lsid:plazi:treatment:03F029281E3EFF95FF2154F0FE94F8C1:8E86CEE51E3EFFAAFD7957F4FC91F913" box="[719,843,1721,1747]" country="Peru" latitude="-12.656388" longLatPrecision="21" longitude="-71.23111" municipality="Manu" name="Pantiacolla" pageId="94" pageNumber="95" stateProvince="Madre de Dios">Pantiacolla</location>
lodge,
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,
<geoCoordinate id="EE6DFEF91E3EFFAAFB9A57F4FB6FF914" box="[1068,1205,1721,1748]" degrees="71" direction="west" minutes="13" orientation="longitude" pageId="94" pageNumber="95" precision="15" seconds="52" value="-71.23111">71°13'52''W</geoCoordinate>
,
<quantity id="4CA135DB1E3EFFAAFB7757F4FAD4F914" box="[1217,1294,1721,1748]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="94" pageNumber="95" unit="m" value="350.0">
<specimenCount id="9D5F53B71E3EFFAAFB7757F4FAD4F914" box="[1217,1294,1721,1748]" count="350" pageId="94" pageNumber="95" type="generic">
<elevation id="00747F0D1E3EFFAAFB7757F4FAD4F914" box="[1217,1294,1721,1748]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="94" pageNumber="95" unit="m" value="350.0">350 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
., on bamboo
<taxonomicName id="4C59E3BD1E3EFFAAFF215790FE5CF937" box="[151,390,1757,1783]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B92D442C1E3EFFAAFF215790FE5CF937" box="[151,390,1757,1783]" italics="true" pageId="94" pageNumber="95">Guadua weberbaueri</emphasis>
</taxonomicName>
,
<date id="FFE7BEFE1E3EFFAAFE255790FDF1F937" box="[403,555,1757,1784]" pageId="94" pageNumber="95" value="2008-05-22">22 May 2008</date>
, leg. J.T. Smit, RMNH.INS.555079;
<materialsCitation id="3B3192631E3EFFAAFC7E5790FB4BF8DC" country="Peru" county="Rio Tambopata" elevation="184" latitude="-12.85714" location="Bridge Condonado Trail" longLatPrecision="1" longitude="-69.373085" municipality="Sachavacayoc" pageId="94" pageNumber="95" specimenCount="19" specimenCount-female="9" specimenCount-male="10" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="9D5F53B71E3EFFAAFC7E5790FC21F937" box="[968,1019,1757,1783]" count="10" pageId="94" pageNumber="95" type="male">10♂</specimenCount>
,
<specimenCount id="9D5F53B71E3EFFAAFBB15790FBF1F937" box="[1031,1067,1757,1783]" count="9" pageId="94" pageNumber="95" type="female">9♀</specimenCount>
,
<collectingRegion id="499D56DC1E3EFFAAFB815790FB3AF937" box="[1079,1248,1757,1783]" country="Peru" name="Madre de Dios" pageId="94" pageNumber="95">Madre de Dios</collectingRegion>
,
<collectingCounty id="6287E0B21E3EFFAAFB5D5790FA42F937" box="[1259,1432,1757,1783]" pageId="94" pageNumber="95">Rio Tambopata</collectingCounty>
,
<collectingMunicipality id="6B8202441E3EFFAAFF21564CFEECF8DB" box="[151,310,1793,1819]" pageId="94" pageNumber="95">Sachavacayoc</collectingMunicipality>
centre,
<location id="8E86CEE51E3EFFAAFE14564CFD13F8DB" LSID="urn:lsid:plazi:treatment:03F029281E3EFF95FF2154F0FE94F8C1:8E86CEE51E3EFFAAFE14564CFD13F8DB" box="[418,713,1793,1819]" country="Peru" county="Rio Tambopata" latitude="-12.85714" longLatPrecision="1" longitude="-69.373085" municipality="Sachavacayoc" name="Bridge Condonado Trail" pageId="94" pageNumber="95" stateProvince="Madre de Dios">Bridge Condonado Trail</location>
,
<geoCoordinate id="EE6DFEF91E3EFFAAFD68564CFCA9F8DC" box="[734,883,1793,1820]" degrees="12" direction="south" minutes="51" orientation="latitude" pageId="94" pageNumber="95" precision="1" seconds="25.7" value="-12.85714">12°51'25.7''S</geoCoordinate>
,
<geoCoordinate id="EE6DFEF91E3EFFAAFC3E564CFBFCF8DB" box="[904,1062,1793,1819]" degrees="69" direction="west" minutes="22" orientation="longitude" pageId="94" pageNumber="95" precision="1" seconds="23.1" value="-69.373085">69°22'23.1''W</geoCoordinate>
,
<quantity id="4CA135DB1E3EFFAAFB8D564CFB4BF8DC" box="[1083,1169,1793,1820]" metricMagnitude="2" metricUnit="m" metricValue="1.8399999999999999" pageId="94" pageNumber="95" unit="m" value="184.0">
<specimenCount id="9D5F53B71E3EFFAAFB8D564CFB4BF8DC" box="[1083,1169,1793,1820]" count="184" pageId="94" pageNumber="95" type="generic">
<elevation id="00747F0D1E3EFFAAFB8D564CFB4BF8DC" box="[1083,1169,1793,1820]" metricMagnitude="2" metricUnit="m" metricValue="1.8399999999999999" pageId="94" pageNumber="95" unit="m" value="184.0">184 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
., on bamboo
<taxonomicName id="4C59E3BD1E3EFFAAFAF4564CFEFFF8FF" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B92D442C1E3EFFAAFAF4564CFEFFF8FF" italics="true" pageId="94" pageNumber="95">Guadua weberbaueri</emphasis>
</taxonomicName>
,
<date id="FFE7BEFE1E3EFFAAFE855668FE03F8FF" box="[307,473,1829,1856]" pageId="94" pageNumber="95" value="2010-06-05" valueMax="2010-06-09" valueMin="2010-06-05">59 June 2010</date>
, leg. J.T. Smit;
<materialsCitation id="3B3192631E3EFFAAFD305668FC8BF8A4" country="Peru" county="Rio Tambopata" elevation="182" latitude="-12.849972" location="Chacra" longLatPrecision="1" longitude="-69.36897" municipality="Sachavacayoc" pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="9D5F53B71E3EFFAAFD305668FD71F8FF" box="[646,683,1829,1855]" count="1" pageId="94" pageNumber="95" type="male">1♂</specimenCount>
,
<specimenCount id="9D5F53B71E3EFFAAFD015668FD00F8FF" box="[695,730,1829,1855]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
,
<collectingRegion id="499D56DC1E3EFFAAFD5E5668FC4BF8FF" box="[744,913,1829,1855]" country="Peru" name="Madre de Dios" pageId="94" pageNumber="95">Madre de Dios</collectingRegion>
,
<collectingCounty id="6287E0B21E3EFFAAFC2A5668FB90F8FF" box="[924,1098,1829,1855]" pageId="94" pageNumber="95">Rio Tambopata</collectingCounty>
,
<collectingMunicipality id="6B8202441E3EFFAAFBE35668FB2EF8FF" box="[1109,1268,1829,1855]" pageId="94" pageNumber="95">Sachavacayoc</collectingMunicipality>
centre,
<location id="8E86CEE51E3EFFAAFAFB5668FA46F8FF" LSID="urn:lsid:plazi:treatment:03F029281E3EFF95FF2154F0FE94F8C1:8E86CEE51E3EFFAAFAFB5668FA46F8FF" box="[1357,1436,1829,1855]" country="Peru" county="Rio Tambopata" latitude="-12.849972" longLatPrecision="1" longitude="-69.36897" municipality="Sachavacayoc" name="Chacra" pageId="94" pageNumber="95" stateProvince="Madre de Dios">Chacra</location>
other side of
<location id="8E86CEE51E3EFFAAFE825604FE7EF8A3" LSID="urn:lsid:plazi:treatment:03F029281E3EFF95FF2154F0FE94F8C1:8E86CEE51E3EFFAAFE825604FE7EF8A3" box="[308,420,1865,1891]" country="Peru" county="Rio Tambopata" latitude="-12.849972" longLatPrecision="1" longitude="-69.36897" municipality="Sachavacayoc" name="Quebrada" pageId="94" pageNumber="95" stateProvince="Madre de Dios">Quebrada</location>
,
<geoCoordinate id="EE6DFEF91E3EFFAAFE045604FD9DF8A4" box="[434,583,1865,1892]" degrees="12" direction="south" minutes="50" orientation="latitude" pageId="94" pageNumber="95" precision="1" seconds="59.9" value="-12.849972">12°50'59.9''S</geoCoordinate>
,
<geoCoordinate id="EE6DFEF91E3EFFAAFDE05604FD2EF8A3" box="[598,756,1865,1891]" degrees="69" direction="west" minutes="22" orientation="longitude" pageId="94" pageNumber="95" precision="1" seconds="08.3" value="-69.36897">69°22'08.3''W</geoCoordinate>
,
<quantity id="4CA135DB1E3EFFAAFCB45604FC8BF8A4" box="[770,849,1865,1892]" metricMagnitude="2" metricUnit="m" metricValue="1.8199999999999998" pageId="94" pageNumber="95" unit="m" value="182.0">
<specimenCount id="9D5F53B71E3EFFAAFCB45604FC8BF8A4" box="[770,849,1865,1892]" count="182" pageId="94" pageNumber="95" type="generic">
<elevation id="00747F0D1E3EFFAAFCB45604FC8BF8A4" box="[770,849,1865,1892]" metricMagnitude="2" metricUnit="m" metricValue="1.8199999999999998" pageId="94" pageNumber="95" unit="m" value="182.0">182 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
., on bamboo
<taxonomicName id="4C59E3BD1E3EFFAAFC595604FB3BF8A3" box="[1007,1249,1865,1891]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B92D442C1E3EFFAAFC595604FB3BF8A3" box="[1007,1249,1865,1891]" italics="true" pageId="94" pageNumber="95">Guadua weberbaueri</emphasis>
</taxonomicName>
,
<date id="FFE7BEFE1E3EFFAAFB445604FF08F847" pageId="94" pageNumber="95" value="2010-10-14" valueMax="2010-10-20" valueMin="2010-10-14">1420 October 2010</date>
, Malaise trap, leg. J.T. Smit;
<materialsCitation id="3B3192631E3EFFAAFD865620FB2FF86B" collectingDate="2010-11-22" collectorName="J. T. Smit" country="Peru" county="Rio Tambopata" latitude="-12.862889" location="Main Trail" longLatPrecision="1" longitude="-69.362946" municipality="Sachavacayoc" pageId="94" pageNumber="95" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="9D5F53B71E3EFFAAFD865620FD8CF847" box="[560,598,1901,1927]" count="2" pageId="94" pageNumber="95" type="male">2♂</specimenCount>
,
<specimenCount id="9D5F53B71E3EFFAAFDD35620FD52F847" box="[613,648,1901,1927]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
,
<collectingRegion id="499D56DC1E3EFFAAFD2C5620FC91F847" box="[666,843,1901,1927]" country="Peru" name="Madre de Dios" pageId="94" pageNumber="95">Madre de Dios</collectingRegion>
,
<collectingCounty id="6287E0B21E3EFFAAFCEC5620FBD6F847" box="[858,1036,1901,1927]" pageId="94" pageNumber="95">Rio Tambopata</collectingCounty>
,
<collectingMunicipality id="6B8202441E3EFFAAFBAD5620FB60F847" box="[1051,1210,1901,1927]" pageId="94" pageNumber="95">Sachavacayoc</collectingMunicipality>
centre,
<location id="8E86CEE51E3EFFAAFAAC5620FA42F847" LSID="urn:lsid:plazi:treatment:03F029281E3EFF95FF2154F0FE94F8C1:8E86CEE51E3EFFAAFAAC5620FA42F847" box="[1306,1432,1901,1927]" country="Peru" county="Rio Tambopata" latitude="-12.862889" longLatPrecision="1" longitude="-69.362946" municipality="Sachavacayoc" name="Main Trail" pageId="94" pageNumber="95" stateProvince="Madre de Dios">Main Trail</location>
,
<geoCoordinate id="EE6DFEF91E3EFFAAFF2156DCFEEDF86C" box="[151,311,1937,1964]" degrees="12" direction="south" minutes="51" orientation="latitude" pageId="94" pageNumber="95" precision="1" seconds="46.4" value="-12.862889">12°5146.4S</geoCoordinate>
,
<geoCoordinate id="EE6DFEF91E3EFFAAFEF556DCFE31F86B" box="[323,491,1937,1963]" degrees="69" direction="west" minutes="21" orientation="longitude" pageId="94" pageNumber="95" precision="1" seconds="46.6" value="-69.362946">69°2146.6W</geoCoordinate>
,
<date id="FFE7BEFE1E3EFFAAFE4156DCFD08F86B" box="[503,722,1937,1963]" pageId="94" pageNumber="95" value="2010-11-22">
<collectingDate id="EFA347161E3EFFAAFE4156DCFD08F86B" box="[503,722,1937,1963]" pageId="94" pageNumber="95" value="2010-11-22">22 November 2010</collectingDate>
</date>
, on bamboo
<taxonomicName id="4C59E3BD1E3EFFAAFCD756DCFB8AF86B" box="[865,1104,1937,1963]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B92D442C1E3EFFAAFCD756DCFB8AF86B" box="[865,1104,1937,1963]" italics="true" pageId="94" pageNumber="95">Guadua weberbaueri</emphasis>
</taxonomicName>
, leg.
<collectorName id="26ACFDE81E3EFFAAFB3856DFFB2FF86B" box="[1166,1269,1937,1964]" pageId="94" pageNumber="95">J.T. Smit</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3192631E3EFFAAFAB756DCFB1DF810" country="Peru" county="Rio Tambopata" elevation="200" location="Chacra" longitude="-12.854306" municipality="Sachavacayoc" pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="2" stateProvince="Madre de Dios" typeStatus="paratype">
<specimenCount id="9D5F53B71E3EFFAAFAB756DCFAFFF86B" box="[1281,1317,1937,1963]" count="2" pageId="94" pageNumber="95" type="female">2♀</specimenCount>
,
<collectingRegion id="499D56DC1E3EFFAAFA8456DCFF15F80F" country="Peru" name="Madre de Dios" pageId="94" pageNumber="95">Madre de Dios</collectingRegion>
,
<collectingCounty id="6287E0B21E3EFFAAFF6D56F8FE50F80F" box="[219,394,1973,1999]" pageId="94" pageNumber="95">Rio Tambopata</collectingCounty>
,
<collectingMunicipality id="6B8202441E3EFFAAFE2056F8FDEFF80F" box="[406,565,1973,1999]" pageId="94" pageNumber="95">Sachavacayoc</collectingMunicipality>
centre, old
<location id="8E86CEE51E3EFFAAFD0D56F8FCD7F80F" LSID="urn:lsid:plazi:treatment:03F029281E3EFF95FF2154F0FE94F8C1:8E86CEE51E3EFFAAFD0D56F8FCD7F80F" box="[699,781,1973,1999]" country="Peru" county="Rio Tambopata" longitude="-12.854306" municipality="Sachavacayoc" name="Chacra" pageId="94" pageNumber="95" stateProvince="Madre de Dios">Chacra</location>
, S
<geoCoordinate id="EE6DFEF91E3EFFAAFC8756F8FC05F810" box="[817,991,1973,2000]" degrees="12" direction="west" minutes="51" orientation="longitude" pageId="94" pageNumber="95" precision="1" seconds="15.5" value="-12.854306">12°5115.5 W</geoCoordinate>
69°2156.6
<quantity id="4CA135DB1E3EFFAAFBCC56F8FB1DF810" box="[1146,1223,1973,2000]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="94" pageNumber="95" unit="m" value="200.0">
<specimenCount id="9D5F53B71E3EFFAAFBCC56F8FB1DF810" box="[1146,1223,1973,2000]" count="200" pageId="94" pageNumber="95" type="generic">
<elevation id="00747F0D1E3EFFAAFBCC56F8FB1DF810" box="[1146,1223,1973,2000]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="94" pageNumber="95" unit="m" value="200.0">200 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation id="3B3192631E3EFFAAFB7156FBFDA0F833" collectingDate="2010-11-20" collectingDateMax="2010-11-29" collectingDateMin="2010-11-20" collectionCode="RMNH" collectorName="J. T. Smit" country="Peru" pageId="94" pageNumber="95" specimenCount="1" typeStatus="paratype">
.,
<date id="FFE7BEFE1E3EFFAAFB6E56F8FF08F833" pageId="94" pageNumber="95" value="2010-11-20" valueMax="2010-11-29" valueMin="2010-11-20">
<collectingDate id="EFA347161E3EFFAAFB6E56F8FF08F833" pageId="94" pageNumber="95" value="2010-11-20" valueMax="2010-11-29" valueMin="2010-11-20">2029 November 2010</collectingDate>
</date>
,
<collectingMethod id="5218E0291E3EFFAAFF6B5694FEB1F834" box="[221,363,2009,2036]" pageId="94" pageNumber="95">Malaise trap</collectingMethod>
, leg.
<collectorName id="26ACFDE81E3EFFAAFE115697FDD0F833" box="[423,522,2009,2036]" pageId="94" pageNumber="95">J.T. Smit</collectorName>
(
<collectionCode id="ED4800FB1E3EFFAAFDAD5697FDB5F834" box="[539,623,2010,2036]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="94" pageNumber="95" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF26C8B61E3FFFABFF2154A7FE63F98A" ID-DOI="http://doi.org/10.5281/zenodo.14023306" ID-Zenodo-Dep="14023306" httpUri="https://zenodo.org/record/14023306/files/figure.png" pageId="95" pageNumber="96" startId="95.[151,250,1514,1538]" targetBox="[151,1436,181,1490]" targetPageId="95" targetType="figure">
<paragraph id="8BE6983E1E3FFFABFF2154A7FE63F98A" blockId="95.[151,1436,1514,1610]" pageId="95" pageNumber="96">
<emphasis id="B92D442C1E3FFFABFF2154A7FEFBF9C2" bold="true" box="[151,289,1514,1538]" pageId="95" pageNumber="96">FIGURE 53.</emphasis>
<taxonomicName id="4C59E3BD1E3FFFABFE9E54A7FE01F9C2" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[296,475,1514,1538]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="laetitiae" status="sp. nov.">
<emphasis id="B92D442C1E3FFFABFE9E54A7FE01F9C2" box="[296,475,1514,1538]" italics="true" pageId="95" pageNumber="96">Ulivellia laetitiae</emphasis>
</taxonomicName>
<emphasis id="B92D442C1E3FFFABFE5554A7FDEEF9C2" bold="true" box="[483,564,1514,1538]" pageId="95" pageNumber="96">
<taxonomicNameLabel id="A21EF9571E3FFFABFE5554A7FDEEF9C2" box="[483,564,1514,1538]" pageId="95" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype ♀ (
<emphasis id="B92D442C1E3FFFABFD0954A7FD0BF9C2" bold="true" box="[703,721,1514,1538]" pageId="95" pageNumber="96">A</emphasis>
,
<emphasis id="B92D442C1E3FFFABFD6954A7FCD5F9C2" bold="true" box="[735,783,1514,1538]" pageId="95" pageNumber="96">CE</emphasis>
) and paratype ♂ (
<emphasis id="B92D442C1E3FFFABFC7A54A7FC07F9C2" bold="true" box="[972,989,1514,1538]" pageId="95" pageNumber="96">B</emphasis>
,
<emphasis id="B92D442C1E3FFFABFC5D54A7FBC0F9C2" bold="true" box="[1003,1050,1514,1538]" pageId="95" pageNumber="96">F, G</emphasis>
).
<emphasis id="B92D442C1E3FFFABFB8654A7FB98F9C2" bold="true" box="[1072,1090,1514,1538]" pageId="95" pageNumber="96">A</emphasis>
, habitus, left;
<emphasis id="B92D442C1E3FFFABFB6154A7FB32F9C2" bold="true" box="[1239,1256,1514,1538]" pageId="95" pageNumber="96">B</emphasis>
, same, dorsal;
<emphasis id="B92D442C1E3FFFABFA3254A7FA4CF9C2" bold="true" box="[1412,1430,1514,1538]" pageId="95" pageNumber="96">C</emphasis>
, head, mesonotum and fore leg, lateral;
<emphasis id="B92D442C1E3FFFABFD855743FD9FF9E6" bold="true" box="[563,581,1550,1574]" pageId="95" pageNumber="96">D</emphasis>
, head, anterior;
<emphasis id="B92D442C1E3FFFABFD445743FCD9F9E6" bold="true" box="[754,771,1550,1574]" pageId="95" pageNumber="96">E</emphasis>
, mesonotum, dorsal;
<emphasis id="B92D442C1E3FFFABFC505743FC2FF9E6" bold="true" box="[998,1013,1550,1574]" pageId="95" pageNumber="96">F</emphasis>
, wing, anterobasal part;
<emphasis id="B92D442C1E3FFFABFB4D5743FAD5F9E6" bold="true" box="[1275,1295,1550,1574]" pageId="95" pageNumber="96">G</emphasis>
, labels. (
<emphasis id="B92D442C1E3FFFABFAC75743FA59F9E6" bold="true" box="[1393,1411,1550,1574]" pageId="95" pageNumber="96">A</emphasis>
— photograph by John T. Smit)
</paragraph>
</caption>
<subSubSection id="C343CBB51E3FFFABFF7157D1FDFDF93F" pageId="95" pageNumber="96" type="diagnosis">
<paragraph id="8BE6983E1E3FFFABFF7157D1FDFDF93F" blockId="95.[151,1437,1692,2043]" pageId="95" pageNumber="96">
<emphasis id="B92D442C1E3FFFABFF7157D1FEE0F976" bold="true" box="[199,314,1692,1718]" pageId="95" pageNumber="96">Diagnosis</emphasis>
.
<taxonomicName id="4C59E3BD1E3FFFABFEFF57D1FDD5F976" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[329,527,1692,1718]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="laetitiae">
<emphasis id="B92D442C1E3FFFABFEFF57D1FDD5F976" box="[329,527,1692,1718]" italics="true" pageId="95" pageNumber="96">Ulivellia laetitiae</emphasis>
</taxonomicName>
differs from all the other species of the genus in that the subbasal band does not continue on the alula, the discal band is more arched than in
<taxonomicName id="4C59E3BD1E3FFFABFCFE578CFC64F91A" authorityName="Speiser" authorityYear="1929" box="[840,958,1729,1754]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="inversa">
<emphasis id="B92D442C1E3FFFABFCFE578CFC64F91A" box="[840,958,1729,1754]" italics="true" pageId="95" pageNumber="96">U. inversa</emphasis>
</taxonomicName>
and crossvein dm-m is basally bent in the apical fourth (
<figureCitation id="136284BB1E3FFFABFE8257A9FE4DF93F" box="[308,407,1764,1791]" captionStart="FIGURE 4" captionStartId="11.[151,250,1686,1710]" captionTargetBox="[247,1341,187,1657]" captionTargetId="figure-20@11.[241,1346,181,1662]" captionTargetPageId="11" captionText="FIGURE 4. Ulivellia, Polyteloptera and similar genera, wing. A, U. amnoni sp. nov.; B, U. inversa Speiser; C, U. laetitiae sp. nov.; DE, U. arcuata sp. nov. (D—male, E—female); F, U. tenoris sp. nov.; G, U. pseudinsolita sp. nov.; H, P. apotropa Hendel; I, “Euxesta” insolita Hendel; J, Gen. sp. near insolita; K, Axiologina ferrumequinum Hendel. Abbreviations. Veins and cells: bm—basal medial cell; br—basal radial cell; c—costal cell; C—costal vein; cua—anterior cubital cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; h—humeral crossvein; M1—first branch of media; m1—first medial cell; M4—fourth branch of media; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; r2+3—second + third radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell; r-m—radial-medial crossvein; Sc—subcostal vein. Wing pattern: DB = discal band; HB = humeral band; AAB = anterior apical band; SAB = subapical band; SCB = subcostal band. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023159" httpUri="https://zenodo.org/record/14023159/files/figure.png" pageId="95" pageNumber="96">Figs 4 C</figureCitation>
,
<figureCitation id="136284BB1E3FFFABFE1557A9FDC0F93F" box="[419,538,1764,1791]" captionStart="FIGURE 53" captionStartId="95.[151,250,1514,1538]" captionTargetBox="[151,1436,181,1490]" captionTargetId="figure-20@95.[151,1436,181,1490]" captionTargetPageId="95" captionText="FIGURE 53. Ulivellia laetitiae sp. nov. holotype ♀ (A, CE) and paratype ♂ (B, F, G). A, habitus, left; B, same, dorsal; C, head, mesonotum and fore leg, lateral; D, head, anterior; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. (A— photograph by John T. Smit)" figureDoi="http://doi.org/10.5281/zenodo.14023306" httpUri="https://zenodo.org/record/14023306/files/figure.png" pageId="95" pageNumber="96">53 A, B, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E3FFF94FF715645FAA2F88D" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96" type="description">
<paragraph id="8BE6983E1E3FFF94FF715645FCD2FDA1" blockId="95.[151,1437,1692,2043]" lastBlockId="96.[151,1437,150,1941]" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
<emphasis id="B92D442C1E3FFFABFF715645FE8BF8E2" bold="true" box="[199,337,1800,1826]" pageId="95" pageNumber="96">Description</emphasis>
. Female. Head (
<figureCitation id="136284BB1E3FFFABFDBE5645FD4BF8E3" box="[520,657,1800,1827]" captionStart="FIGURE 53" captionStartId="95.[151,250,1514,1538]" captionTargetBox="[151,1436,181,1490]" captionTargetId="figure-20@95.[151,1436,181,1490]" captionTargetPageId="95" captionText="FIGURE 53. Ulivellia laetitiae sp. nov. holotype ♀ (A, CE) and paratype ♂ (B, F, G). A, habitus, left; B, same, dorsal; C, head, mesonotum and fore leg, lateral; D, head, anterior; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. (A— photograph by John T. Smit)" figureDoi="http://doi.org/10.5281/zenodo.14023306" httpUri="https://zenodo.org/record/14023306/files/figure.png" pageId="95" pageNumber="96">Figs 53 E, F</figureCitation>
) ratio (length: height: width) 1: 1.4: 1.5, mostly black, except lower half of frons, gena and parafacial brownish brown. Frons 0.7× longer than wide (measured from ptilinale fissure to fore ocellus), orange brown with black ocellar triangle and vertical plates; orbits silver-white microtrichose, brown. Parafacial narrow, 0.20.3× broader than postpedicel, slightly white microtrichose. Frontal setulae relatively short, black, inclinate and partly proclinate, forming two irregular rows of 36 setulae on each side, all more than ⅓ and up to ½ the length of ocellar setae; 2 orbital setae, posterior longer than anterior, about 0.50.6× longer than inner vertical seta and 34× longer than outer vertical seta; postocellar 0.3× longer than inner vertical setae; outer vertical seta short, 0.20.25× longer than inner vertical seta and barely distinguishable from postoculair setae forming one row allied by 912 dorsal and 1015 ventral occipital setae. Lunule very narrow, indisctinct, orange-yellow. Eye 1.6× higher than long. Face (above suture) as high as wide at transverse fold, brown, obscured ventrally, densely white microtrichose; epistome brown to black, sparsely white microtrichose, metallic shiny, often with greenish or cyan shine, strongly produced anteriorly and separated from upper part of face by deep suture. Clypeus brown to black, sparsely white microtrichose, metallic shining, often with cyan sheen, 0.30.4× higher than epistome. Gena brownish yellow, with long genal seta and 34 additional setae anterior of it, 0.50.8× longer than genal seta. Occiput black, except postgena brown, with silvery to cyan metallic sheen, moderately densely covered with white microtrichia partly hiding underlying cuticle; ventral part of occiput (postgena) with 45 setae almost as long as genal seta. Antenna brownish yellow; scape and pedicel with black setulae; postpedicel brown, blackish antero-dorsally, whitish microtrichose, 1.9× longer than wide, narrowed in apical half and rounded; arista black except yellow at base, almost bare. Mouthparts black, prementum black, shiny. Palp yellowish-brown, with black setulae, ventrally and subapically with setulae 23 times longer.
</paragraph>
<paragraph id="8BE6983E1E00FF94FF715326FB83FC65" blockId="96.[151,1437,150,1941]" pageId="96" pageNumber="97">
<emphasis id="B92D442C1E00FF94FF715326FEFAFD45" bold="true" box="[199,288,619,645]" pageId="96" pageNumber="97">Thorax</emphasis>
(
<figureCitation id="136284BB1E00FF94FE985326FE4EFD45" box="[302,404,618,645]" captionStart="FIGURE 53" captionStartId="95.[151,250,1514,1538]" captionTargetBox="[151,1436,181,1490]" captionTargetId="figure-20@95.[151,1436,181,1490]" captionTargetPageId="95" captionText="FIGURE 53. Ulivellia laetitiae sp. nov. holotype ♀ (A, CE) and paratype ♂ (B, F, G). A, habitus, left; B, same, dorsal; C, head, mesonotum and fore leg, lateral; D, head, anterior; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. (A— photograph by John T. Smit)" figureDoi="http://doi.org/10.5281/zenodo.14023306" httpUri="https://zenodo.org/record/14023306/files/figure.png" pageId="96" pageNumber="97">Fig. 53 F</figureCitation>
) black, metallically shiny with yellowish-green or cyan sheen and sparse yellow microtrichia not obscuring underlying cuticle. Mesonotal scutum 1.4× longer than wide; with 3 almost inconspicuous yellowish-green vittae in otherwise silvery cyan sheen in the presutural area, postsutural area with yellowish-green shine, black setulose, with 810 rows of setulae between dorsocentral setulae; 1 pair of acrostichal setae; prescutellar area without setulae. Scutellum dorsally flattened, posteriorly black or brownish, finely shagreened, sparsely microtrichose, without setulae. Subscutellum subshining, black with yellowish -green metallic sheen. Mediotergite shining black with metallic yellowish-green sheen, sparsely microtrichose. Set of setae normal for the genus. All the setae and setulae black. Anepisternum apicoventrally with yellowish-green sheen, as well as anepimeron ventrally and katepisternum nearly entirely, pleurae otherwise with metallic silvery cyan sheen.
</paragraph>
<paragraph id="8BE6983E1E00FF94FF7152E3FBBDFAB9" blockId="96.[151,1437,150,1941]" pageId="96" pageNumber="97">
<emphasis id="B92D442C1E00FF94FF7152E3FED2FC08" bold="true" box="[199,264,942,968]" pageId="96" pageNumber="97">Wing</emphasis>
(
<figureCitation id="136284BB1E00FF94FEA052E2FEACFC09" box="[278,374,943,969]" captionStart="FIGURE 4" captionStartId="11.[151,250,1686,1710]" captionTargetBox="[247,1341,187,1657]" captionTargetId="figure-20@11.[241,1346,181,1662]" captionTargetPageId="11" captionText="FIGURE 4. Ulivellia, Polyteloptera and similar genera, wing. A, U. amnoni sp. nov.; B, U. inversa Speiser; C, U. laetitiae sp. nov.; DE, U. arcuata sp. nov. (D—male, E—female); F, U. tenoris sp. nov.; G, U. pseudinsolita sp. nov.; H, P. apotropa Hendel; I, “Euxesta” insolita Hendel; J, Gen. sp. near insolita; K, Axiologina ferrumequinum Hendel. Abbreviations. Veins and cells: bm—basal medial cell; br—basal radial cell; c—costal cell; C—costal vein; cua—anterior cubital cell; CuA+CuP— anterior branch of cubital vein + posterior branch of cubital vein; dm—discal medial cell; dm-m—discal medial crossvein; h—humeral crossvein; M1—first branch of media; m1—first medial cell; M4—fourth branch of media; m4—fourth medial cell; ptstg—pterostigma; R1—anterior branch of radius; r1—first radial cell; r2+3—second + third radial cell; R2+3—second branch of radius; R 4+5 —third branch of radius; r 4+5 —fourth + fifth radial cell; r-m—radial-medial crossvein; Sc—subcostal vein. Wing pattern: DB = discal band; HB = humeral band; AAB = anterior apical band; SAB = subapical band; SCB = subcostal band. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14023159" httpUri="https://zenodo.org/record/14023159/files/figure.png" pageId="96" pageNumber="97">Figs 4 C</figureCitation>
,
<figureCitation id="136284BB1E00FF94FE3652E3FE69FC09" box="[384,435,942,969]" captionStart="FIGURE 53" captionStartId="95.[151,250,1514,1538]" captionTargetBox="[151,1436,181,1490]" captionTargetId="figure-20@95.[151,1436,181,1490]" captionTargetPageId="95" captionText="FIGURE 53. Ulivellia laetitiae sp. nov. holotype ♀ (A, CE) and paratype ♂ (B, F, G). A, habitus, left; B, same, dorsal; C, head, mesonotum and fore leg, lateral; D, head, anterior; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. (A— photograph by John T. Smit)" figureDoi="http://doi.org/10.5281/zenodo.14023306" httpUri="https://zenodo.org/record/14023306/files/figure.png" pageId="96" pageNumber="97">53 F</figureCitation>
)
<quantity id="4CA135DB1E00FF94FE7652E3FD9CFC09" box="[448,582,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="6.6" metricValueMin="5.8" pageId="96" pageNumber="97" unit="mm" value="6.199999999999999" valueMax="6.6" valueMin="5.8">5.86.6 mm</quantity>
(
<specimenCount id="9D5F53B71E00FF94FDE252E2FDB0FC09" box="[596,618,943,969]" count="1" pageId="96" pageNumber="97" type="male"></specimenCount>
)
<quantity id="4CA135DB1E00FF94FDCF52E2FD24FC09" box="[633,766,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="6.8" metricValueMin="6.2" pageId="96" pageNumber="97" unit="mm" value="6.5" valueMax="6.8" valueMin="6.2">6.26.8 mm</quantity>
(
<specimenCount id="9D5F53B71E00FF94FCB852E2FCFAFC09" box="[782,800,943,969]" count="1" pageId="96" pageNumber="97" type="female"></specimenCount>
) long: 2.32.5× longer than wide, basicostal cell hyaline; costal cell long, 4.8× longer than wide, brown in basal third, with costa almost straight, forming neither lobe nor cleft; pterostigma entirely brown, narrow triangular, 5.1× longer than wide, vein R
<subScript id="17DD9A7B1E00FF94FB9B5544FBEFFBD7" attach="left" box="[1069,1077,1033,1047]" fontSize="6" pageId="96" pageNumber="97">1</subScript>
setulose dorsally at pterostigma level, ending at level of crossvein dm-m; vein R
<subScript id="17DD9A7B1E00FF94FD735560FD04FBFB" attach="left" box="[709,734,1069,1083]" fontSize="6" pageId="96" pageNumber="97">2+3</subScript>
at apex bent anteriorly and then subparallel to costa, turning towards it before very apex; vein R
<subScript id="17DD9A7B1E00FF94FD95551CFDE7FB9F" attach="left" box="[547,573,1105,1119]" fontSize="6" pageId="96" pageNumber="97">4+5</subScript>
arcuate, first upward from crossvein r-m and downward distally, terminating just shortof wing apex; vein M
<subScript id="17DD9A7B1E00FF94FE4E5538FDCBFB43" attach="left" box="[504,529,1141,1155]" fontSize="6" pageId="96" pageNumber="97">1+2</subScript>
similarly curved but in opposite direction, terminating well beyond wing apex; crossvein r-m at basal ¼ of cell dm; crossvein dm-m bent basally in the apical fourth. Cell r
<subScript id="17DD9A7B1E00FF94FB2555D4FB77FB67" attach="left" box="[1171,1197,1177,1191]" fontSize="6" pageId="96" pageNumber="97">4+5</subScript>
4.55.5× longer than wide, broadest just beyond the middle and narrowed apically; Cell m1 narrowly triangular; ultimate section of M
<subScript id="17DD9A7B1E00FF94FA2255F0FA46FB0B" attach="left" box="[1428,1436,1213,1227]" fontSize="6" pageId="96" pageNumber="97">1</subScript>
2.4×. as long as crossvein dm-m and 1.9× longer than penultimate section. Vein CuA slightly convex in upper part, cell cua with short posteroapical lobe along vein CuP. Wing pattern similar to
<taxonomicName id="4C59E3BD1E00FF94FBBF55BEFBA4FACC" authorityName="Speiser" authorityYear="1929" box="[1033,1150,1267,1292]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="inversa">
<emphasis id="B92D442C1E00FF94FBBF55BEFBA4FACC" box="[1033,1150,1267,1292]" italics="true" pageId="96" pageNumber="97">U. inversa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C59E3BD1E00FF94FB0055BEFAF7FACC" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[1206,1325,1267,1292]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="amnoni">
<emphasis id="B92D442C1E00FF94FB0055BEFAF7FACC" box="[1206,1325,1267,1292]" italics="true" pageId="96" pageNumber="97">U. amnoni</emphasis>
</taxonomicName>
, subbasal band broader and more or less oblique, not continued on alula, discal band strongly arched, stronger than in
<taxonomicName id="4C59E3BD1E00FF94FA37545AFF33FA94" authorityName="Speiser" authorityYear="1929" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="inversa">
<emphasis id="B92D442C1E00FF94FA37545AFF33FA94" italics="true" pageId="96" pageNumber="97">U. inversa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C59E3BD1E00FF94FE965476FE4CFA94" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[288,406,1339,1364]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="amnoni">
<emphasis id="B92D442C1E00FF94FE965476FE4CFA94" box="[288,406,1339,1364]" italics="true" pageId="96" pageNumber="97">U. amnoni</emphasis>
</taxonomicName>
, broader at cell br and clearly narrowed at posterior end, preapical band slightly less arched than in
<taxonomicName id="4C59E3BD1E00FF94FF5D5412FE85FAB8" authorityName="Speiser" authorityYear="1929" box="[235,351,1375,1400]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="inversa">
<emphasis id="B92D442C1E00FF94FF5D5412FE85FAB8" box="[235,351,1375,1400]" italics="true" pageId="96" pageNumber="97">U. inversa</emphasis>
</taxonomicName>
. Calypters white, with white ciliae. Halter creamy with yellow knob.
</paragraph>
<paragraph id="8BE6983E1E00FF94FF7154CEFB04FA25" blockId="96.[151,1437,150,1941]" pageId="96" pageNumber="97">
<emphasis id="B92D442C1E00FF94FF7154CEFF25FA5D" bold="true" box="[199,255,1411,1437]" pageId="96" pageNumber="97">Legs</emphasis>
(
<figureCitation id="136284BB1E00FF94FEB854CEFE44FA5D" box="[270,414,1410,1437]" captionStart="FIGURE 53" captionStartId="95.[151,250,1514,1538]" captionTargetBox="[151,1436,181,1490]" captionTargetId="figure-20@95.[151,1436,181,1490]" captionTargetPageId="95" captionText="FIGURE 53. Ulivellia laetitiae sp. nov. holotype ♀ (A, CE) and paratype ♂ (B, F, G). A, habitus, left; B, same, dorsal; C, head, mesonotum and fore leg, lateral; D, head, anterior; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. (A— photograph by John T. Smit)" figureDoi="http://doi.org/10.5281/zenodo.14023306" httpUri="https://zenodo.org/record/14023306/files/figure.png" pageId="96" pageNumber="97">Figs 53 AC</figureCitation>
) all entirely black bluish cyan sheen, femora partially with yellowish-green metallic sheen, except coxae ventrally and trochanters entirely yellow-brown and knees narrowly brown, all sparse with microtrichia except for coxae of prolegs. Mid tibia ventrally with one spur-like seta 3× longer than tibia width.
</paragraph>
<paragraph id="8BE6983E1E00FF94FF7154A3FBADF9ED" blockId="96.[151,1437,150,1941]" pageId="96" pageNumber="97">
<emphasis id="B92D442C1E00FF94FF7154A3FEE6F9C8" bold="true" box="[199,316,1518,1544]" pageId="96" pageNumber="97">Abdomen</emphasis>
(
<figureCitation id="136284BB1E00FF94FEE654A2FE30F9C9" box="[336,490,1518,1545]" captionStart="FIGURE 53" captionStartId="95.[151,250,1514,1538]" captionTargetBox="[151,1436,181,1490]" captionTargetId="figure-20@95.[151,1436,181,1490]" captionTargetPageId="95" captionText="FIGURE 53. Ulivellia laetitiae sp. nov. holotype ♀ (A, CE) and paratype ♂ (B, F, G). A, habitus, left; B, same, dorsal; C, head, mesonotum and fore leg, lateral; D, head, anterior; E, mesonotum, dorsal; F, wing, anterobasal part; G, labels. (A— photograph by John T. Smit)" figureDoi="http://doi.org/10.5281/zenodo.14023306" httpUri="https://zenodo.org/record/14023306/files/figure.png" pageId="96" pageNumber="97">Figs 53 AB</figureCitation>
) entirely black, tergites very finely shagreened and sparsely microtrichose, with metallic yellowish-green metallic sheen, less shiny than pleurae; setulae and setae black.
</paragraph>
<paragraph id="8BE6983E1E00FF94FF71577AFAA2F88D" blockId="96.[151,1437,150,1941]" pageId="96" pageNumber="97">
<emphasis id="B92D442C1E00FF94FF71577AFE6BF991" bold="true" box="[199,433,1591,1617]" pageId="96" pageNumber="97">Male postabdomen</emphasis>
. Sternite 8 setulose, with 2 longer setae. Cerci short oval, setulose (
<figureCitation id="136284BB1E00FF94FB59577AFA4AF991" box="[1263,1424,1590,1617]" captionStart="FIGURE 54" captionStartId="97.[151,250,1457,1481]" captionTargetBox="[151,1432,182,1433]" captionTargetId="figure-20@97.[151,1435,181,1433]" captionTargetPageId="97" captionText="FIGURE 54. Ulivellia laetitiae sp. nov. paratype ♂ (SIZK) A, postabdomen, lateral; B, same, anteroventral; C, same, ventral; D, epandrium, with cerci and surstyli, posterolateral; E, surstyli and hypandrium, ventral. Abbreviations: cerc—cerci; epand— epandrium; ph g—phallic guide; ph—phallus; phapd—phallapodeme; pi p—pimple-like process; pregt—pregonite; sur— surstylus; v l sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.14023308" httpUri="https://zenodo.org/record/14023308/files/figure.png" pageId="96" pageNumber="97">Figs 54 AD</figureCitation>
). Epandrium ovoid, dorsally short, moderately setulose. Surstyli basally wide, inner surstylus with 1011 prensisetae, outer surstylus simple with narrow, elongated, mesally directed ventral lobe apically bearing dorsally directed pimple-like process, but without distinct antero-ventral and postero-ventral lobes; inner surstylus with 1215 thickened prensisetae (
<figureCitation id="136284BB1E00FF94FE20578AFDDFF921" box="[406,517,1734,1761]" captionStart="FIGURE 54" captionStartId="97.[151,250,1457,1481]" captionTargetBox="[151,1432,182,1433]" captionTargetId="figure-20@97.[151,1435,181,1433]" captionTargetPageId="97" captionText="FIGURE 54. Ulivellia laetitiae sp. nov. paratype ♂ (SIZK) A, postabdomen, lateral; B, same, anteroventral; C, same, ventral; D, epandrium, with cerci and surstyli, posterolateral; E, surstyli and hypandrium, ventral. Abbreviations: cerc—cerci; epand— epandrium; ph g—phallic guide; ph—phallus; phapd—phallapodeme; pi p—pimple-like process; pregt—pregonite; sur— surstylus; v l sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.14023308" httpUri="https://zenodo.org/record/14023308/files/figure.png" pageId="96" pageNumber="97">Fig. 54 D</figureCitation>
). Pregonites symmetrical, with 89 setulae ventro-mesally, conspicuously anterior of postgonites. Postgonites on laterobasal part of the phallic guide lobes, with 46 small trichoid sensilla. Phallus moderately short and narrow, about 34 times as long as epandrium high, almost bare, in rest coiled at right side of abdomen (
<figureCitation id="136284BB1E00FF94FEBB567EFEA0F88D" box="[269,378,1842,1869]" captionStart="FIGURE 54" captionStartId="97.[151,250,1457,1481]" captionTargetBox="[151,1432,182,1433]" captionTargetId="figure-20@97.[151,1435,181,1433]" captionTargetPageId="97" captionText="FIGURE 54. Ulivellia laetitiae sp. nov. paratype ♂ (SIZK) A, postabdomen, lateral; B, same, anteroventral; C, same, ventral; D, epandrium, with cerci and surstyli, posterolateral; E, surstyli and hypandrium, ventral. Abbreviations: cerc—cerci; epand— epandrium; ph g—phallic guide; ph—phallus; phapd—phallapodeme; pi p—pimple-like process; pregt—pregonite; sur— surstylus; v l sur—ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.14023308" httpUri="https://zenodo.org/record/14023308/files/figure.png" pageId="96" pageNumber="97">Fig. 54 B</figureCitation>
). Ejaculatory apodeme short, bar-like, with relatively small sperm pump (
<figureCitation id="136284BB1E00FF94FB02567EFAFDF88D" box="[1204,1319,1842,1869]" captionStart="FIGURE 52" captionStartId="91.[151,250,1586,1610]" captionTargetBox="[154,1434,394,1550]" captionTargetId="figure-138@91.[153,1434,381,1562]" captionTargetPageId="91" captionText="FIGURE 52. Ulivellia arcuata sp. nov. paratype ♂ (SIZK) A, preabdomen; B, postabdomen, ventral; C, same, posterior; D, same, lateral E, same, dorsal F, hypandrium, ventral; G, surstyli, posterior; H, phallus. Abbreviations: cerc—cerci; epand— epandrium; ej apod—ejaculatory apodeme; hypd—hypandrium; ph g—phallic guide; ph—phallus; phapd—phallapodeme; pstg—postgonite." figureDoi="http://doi.org/10.5281/zenodo.14023304" httpUri="https://zenodo.org/record/14023304/files/figure.png" pageId="96" pageNumber="97">Figs 52 D</figureCitation>
,
<figureCitation id="136284BB1E00FF94FA84567FFAB0F88D" box="[1330,1386,1842,1869]" captionStart="FIGURE 56" captionStartId="100.[151,250,1759,1783]" captionTargetBox="[158,1425,188,1728]" captionTargetId="figure-30@100.[151,1436,181,1735]" captionTargetPageId="100" captionText="FIGURE 56. Ulivellia pseudinsolita sp. nov. holotype ♂. A, postabdomen, posterior; B C, same, ventral; D, same, ventrolateral; E, surstyli and hypandrium, ventral; F, preabdomen, ventral. Abbreviations: cerc—cerci; ej apod—ejaculatory apodeme; epand—epandrium; ph g—phallic guide; ph—phallus; pregt—pregonite; p l sur—posterior lobe of surstylus; v l sur—ventral lobe of surstylus; st 2— second sternite; st 3—third sternite; st 4—fourth sternite; st 5—fifth sternite." figureDoi="http://doi.org/10.5281/zenodo.14023312" httpUri="https://zenodo.org/record/14023312/files/figure.png" pageId="96" pageNumber="97">56 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C343CBB51E00FF94FF71561AFC20F855" pageId="96" pageNumber="97" type="etymology">
<paragraph id="8BE6983E1E00FF94FF71561AFC20F855" blockId="96.[151,1437,150,1941]" pageId="96" pageNumber="97">
<emphasis id="B92D442C1E00FF94FF71561AFE92F8B1" bold="true" box="[199,328,1879,1905]" pageId="96" pageNumber="97">Etymology</emphasis>
. This species is named in honor of Laetitia Martina, the authors girlfriend and mother of his children. The specific epithet should be treated as a noun in the genitive case.
</paragraph>
</subSubSection>
<caption id="DF26C8B61E01FF95FF2154FCFD89F9F5" ID-DOI="http://doi.org/10.5281/zenodo.14023308" ID-Zenodo-Dep="14023308" httpUri="https://zenodo.org/record/14023308/files/figure.png" pageId="97" pageNumber="98" startId="97.[151,250,1457,1481]" targetBox="[151,1432,182,1433]" targetPageId="97" targetType="figure">
<paragraph id="8BE6983E1E01FF95FF2154FCFD89F9F5" blockId="97.[151,1436,1457,1589]" pageId="97" pageNumber="98">
<emphasis id="B92D442C1E01FF95FF2154FCFEFAFA09" bold="true" box="[151,288,1457,1481]" pageId="97" pageNumber="98">FIGURE 54.</emphasis>
<taxonomicName id="4C59E3BD1E01FF95FE9054FCFE02FA09" authorityName="Kovac &amp; Kameneva &amp; Korneyev &amp; Araújo &amp; Savaris &amp; Smit &amp; Schneider &amp; Schreiber &amp; Korneyev" authorityYear="2024" box="[294,472,1457,1481]" class="Insecta" family="Ulidiidae" genus="Ulivellia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="laetitiae" status="sp. nov.">
<emphasis id="B92D442C1E01FF95FE9054FCFE02FA09" box="[294,472,1457,1481]" italics="true" pageId="97" pageNumber="98">Ulivellia laetitiae</emphasis>
</taxonomicName>
<emphasis id="B92D442C1E01FF95FE6954FFFDEAFA0A" bold="true" box="[479,560,1458,1482]" pageId="97" pageNumber="98">
<taxonomicNameLabel id="A21EF9571E01FF95FE6954FFFDEAFA0A" box="[479,560,1458,1482]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype ♂ (SIZK)
<emphasis id="B92D442C1E01FF95FD4B54FCFCD5FA09" bold="true" box="[765,783,1457,1481]" pageId="97" pageNumber="98">A</emphasis>
, postabdomen, lateral;
<emphasis id="B92D442C1E01FF95FC4B54FFFBD4FA0A" bold="true" box="[1021,1038,1458,1482]" pageId="97" pageNumber="98">B</emphasis>
, same, anteroventral;
<emphasis id="B92D442C1E01FF95FB5954FCFADBFA09" bold="true" box="[1263,1281,1457,1481]" pageId="97" pageNumber="98">C</emphasis>
, same, ventral;
<emphasis id="B92D442C1E01FF95FF21549BFF73FA2E" bold="true" box="[151,169,1494,1518]" pageId="97" pageNumber="98">D</emphasis>
, epandrium, with cerci and surstyli, posterolateral;
<emphasis id="B92D442C1E01FF95FD02549BFD1FFA2E" bold="true" box="[692,709,1494,1518]" pageId="97" pageNumber="98">E</emphasis>
, surstyli and hypandrium, ventral. Abbreviations: cerc—cerci; epand— epandrium; ph g—phallic guide; ph—phallus; phapd—phallapodeme; pi p—pimple-like process; pregt—pregonite; sur— surstylus; v l sur—ventral lobe of surstylus.
</paragraph>
</caption>
<subSubSection id="C343CBB51E01FF95FF715737FE94F8C1" pageId="97" pageNumber="98" type="biology_ecology">
<paragraph id="8BE6983E1E01FF95FF715737FE94F8C1" blockId="97.[151,1436,1658,1793]" pageId="97" pageNumber="98">
<emphasis id="B92D442C1E01FF95FF715737FEFBF954" bold="true" box="[199,289,1658,1684]" pageId="97" pageNumber="98">Biology</emphasis>
. All specimens were collected and many additional specimens were observed on mature stems of the bamboo
<taxonomicName id="4C59E3BD1E01FF95FF4157D3FE3CF978" box="[247,486,1694,1720]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="97" pageNumber="98" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B92D442C1E01FF95FF4157D3FE3CF978" box="[247,486,1694,1720]" italics="true" pageId="97" pageNumber="98">Guadua weberbaueri</emphasis>
</taxonomicName>
, in contrast to many other species that were specifically attracted to freshly cut young shoots of the same bamboo species. This species appears to develop in the internodes of damaged mature stems of
<taxonomicName id="4C59E3BD1E01FF95FF2157AAFE9DF8C0" box="[151,327,1766,1793]" class="Liliopsida" family="Poaceae" genus="Guadua" kingdom="Plantae" order="Poales" pageId="97" pageNumber="98" phylum="Tracheophyta" rank="species" species="weberbaueri">
<emphasis id="B92D442C1E01FF95FF2157AAFE9DF8C0" box="[151,327,1766,1793]" italics="true" pageId="97" pageNumber="98">G. weberbaueri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>