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<document id="92DD25C6FF306E208ECA4A5B04FFF827" ID-CLB-Dataset="20529" ID-DOI="10.11646/zootaxa.5100.4.1" ID-GBIF-Dataset="38ddf0a4-c35b-47a4-bf6a-5851ea91b0b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="6224598" ID-ZooBank="74EEFDED-EEB9-46DF-83D5-2FB5693F920E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645518222118" checkinUser="plazi" docAuthor="Mateos, Eduardo &amp; Álvarez-Presas, Marta" docDate="2022" docId="03E387D5FF97FF9164C1FD48BC3482A8" docLanguage="en" docName="zootaxa.5100.4.1.pdf" docOrigin="Zootaxa 5100 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidocyrtus fuscocephalus Mateos &amp; Álvarez-Presas 2022, sp. nov." docType="treatment" docUuid="63B8C59B-4F13-45A9-A26B-3A71E784ADA4" docUuidSource="ZooBank" docVersion="7" lastPageNumber="463" masterDocId="FFDAFFADFF91FF9C6456FFFABD09827D" masterDocTitle="Integrative taxonomy reveals three new species of European Lepidocyrtus lignorum-group (Collembola, Entomobryidae)" masterLastPageNumber="481" masterPageNumber="451" pageNumber="457" updateTime="1699315952817" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FAA23EA964D2A86ED30921E71D91444C">Integrative taxonomy reveals three new species of European Lepidocyrtus lignorum-group (Collembola, Entomobryidae)</mods:title>
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<mods:namePart id="CB5A82B857B08580A5BDC0515E92B395">Mateos, Eduardo</mods:namePart>
<mods:affiliation id="9B60952C55473E81833D256ACDA76BC8">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals; Facultat de Biologia; Universitat de Barcelona; Avinguda Diagonal 643, 08028 - Barcelona; Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona, Barcelona, Spain.</mods:affiliation>
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<mods:namePart id="B42A62BCF686C404986D66DFFF8DE68E">Álvarez-Presas, Marta</mods:namePart>
<mods:affiliation id="32347EA56921FE0BE26AE1D2EF96D897">University of Bristol, School of Life Sciences, Bristol, United Kingdom.</mods:affiliation>
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<mods:title id="10C9B955CA901A968B34C8EED9599C31">Zootaxa</mods:title>
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<mods:date id="061043694DCF6CD9C503760A9B17CB28">2022</mods:date>
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<mods:number id="5E03E342D390C7C1F532330E09CBCF49">2022-02-22</mods:number>
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<treatment id="03E387D5FF97FF9164C1FD48BC3482A8" ID-DOI="http://doi.org/10.5281/zenodo.6315108" ID-GBIF-Taxon="193556205" ID-Zenodo-Dep="6315108" LSID="urn:lsid:zoobank.org:act:63B8C59B-4F13-45A9-A26B-3A71E784ADA4" httpUri="http://treatment.plazi.org/id/03E387D5FF97FF9164C1FD48BC3482A8" lastPageId="13" lastPageNumber="463" pageId="6" pageNumber="457">
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<heading id="D0BD81AFFF97FF9A64C1FD48BFA680B0" bold="true" box="[151,687,690,717]" fontSize="11" level="1" pageId="6" pageNumber="457" reason="1">
<emphasis id="B93EEAD1FF97FF9A64C1FD48BFA680B0" bold="true" box="[151,687,690,717]" pageId="6" pageNumber="457">
<taxonomicName id="4C4A4D40FF97FF9A64C1FD48BF4380B0" ID-CoL="8ZNC8" authority="Mateos" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[151,586,690,717]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF97FF9A64C1FD48BCE180B0" bold="true" box="[151,488,690,717]" italics="true" pageId="6" pageNumber="457">Lepidocyrtus fuscocephalus</emphasis>
Mateos
</taxonomicName>
<taxonomicNameLabel id="A20D57AAFF97FF9A6604FD49BFA680B0" box="[594,687,691,717]" pageId="6" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3506548FF97FF9A64C1FD2DB94A8168" pageId="6" pageNumber="457" type="description">
<paragraph id="8BF536C3FF97FF9A64C1FD2DBC70808C" blockId="6.[151,1091,690,789]" box="[151,377,727,753]" pageId="6" pageNumber="457">
<figureCitation id="13712A46FF97FF9A64C1FD2DBC01808C" box="[151,264,727,753]" captionStart-0="FIGURE 8" captionStart-1="FIGURES 912" captionStart-2="FIGURE 13" captionStart-3="FIGURES 1416" captionStart-4="FIGURES 1718" captionStart-5="FIGURE 19" captionStart-6="FIGURES 2022" captionStart-7="FIGURE 23" captionStartId-0="6.[151,250,1914,1938]" captionStartId-1="7.[151,264,1929,1953]" captionStartId-2="8.[151,250,1874,1898]" captionStartId-3="9.[151,264,1943,1967]" captionStartId-4="10.[151,264,1426,1450]" captionStartId-5="11.[151,250,1943,1967]" captionStartId-6="12.[151,264,633,657]" captionStartId-7="13.[151,250,1892,1916]" captionTargetBox-0="[152,1436,1209,1891]" captionTargetBox-1="[151,1436,812,1906]" captionTargetBox-2="[151,1436,181,1850]" captionTargetBox-3="[169,1414,183,1919]" captionTargetBox-4="[156,1429,190,1402]" captionTargetBox-5="[243,1320,181,1919]" captionTargetBox-6="[151,1436,189,609]" captionTargetBox-7="[151,1436,237,1855]" captionTargetId-0="figure-385@6.[152,1436,1209,1891]" captionTargetId-1="figure-313@7.[151,1436,812,1906]" captionTargetId-2="figure-34@8.[151,1436,181,1850]" captionTargetId-3="figure-17@9.[169,1418,181,1919]" captionTargetId-4="figure-34@10.[151,1436,181,1402]" captionTargetId-5="figure-17@11.[243,1343,181,1919]" captionTargetId-6="figure-34@12.[151,1436,181,609]" captionTargetId-7="figure-40@13.[151,1436,237,1869]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionText-0="FIGURE 8. Lepidocyrtus fuscocephalus sp. nov.: Habitus lateral (specimen in alcohol, without scales)." captionText-1="FIGURES 912. Lepidocyrtus fuscocephalus sp. nov.: 9, clypeal and labral chaetotaxy; 10, maxillary palp (right side); 11, outer labial papilla (right side); 12, labial and postlabial chaetotaxy (the arrow points to a supernumerary M chaeta present in one paratype). Black dotsciliated chaetae, crossscales, p.c.—proximal chaetae of labial palp." captionText-2="FIGURE 13. Lepidocyrtus fuscocephalus sp. nov.: Dorsal head chaetotaxy (left side). Broad circleslong ciliated macrochaetae, small circlesshort ciliated macrochaetae." captionText-3="FIGURES 1416. Lepidocyrtus fuscocephalus sp. nov. dorsal chaetotaxy (left side): 14, Th.II; 15, Th.III; 16 Abd.I. Circlesciliated chaetae." captionText-4="FIGURES 1718. Lepidocyrtus fuscocephalus sp. nov. dorsal chaetotaxy (left side): 17, Abd.II; 18, Abd.III. Broad circlesbroad ciliated macrochaetae, small circlesthin ciliated macrochaetae." captionText-5="FIGURE 19. Lepidocyrtus fuscocephalus sp. nov.: Abd.IV chaetotaxy (left side). Broad black circlesbroad ciliated macrochaetae, small black circlesthin ciliated macrochaetae." captionText-6="FIGURES 2022. Lepidocyrtus fuscocephalus sp. nov.: 20, Abd.V chaetotaxy (left side), broad circleslong ciliated chaetae, small circlesshort ciliated or smooth chaetae; 21, trochanteral organ; 22, third leg unguis and unguiculus." captionText-7="FIGURE 23. Lepidocyrtus fuscocephalus sp. nov, L. milagrosae sp. nov. and L. semicoloratus sp. nov.: Pseudopores distribution: a, dorsal pseudopores; bc, ventral pesudopores." figureDoi-0="http://doi.org/10.5281/zenodo.6224610" figureDoi-1="http://doi.org/10.5281/zenodo.6224616" figureDoi-2="http://doi.org/10.5281/zenodo.6224624" figureDoi-3="http://doi.org/10.5281/zenodo.6224632" figureDoi-4="http://doi.org/10.5281/zenodo.6224640" figureDoi-5="http://doi.org/10.5281/zenodo.6224652" figureDoi-6="http://doi.org/10.5281/zenodo.6224662" figureDoi-7="http://doi.org/10.5281/zenodo.6224666" httpUri-0="https://zenodo.org/record/6224610/files/figure.png" httpUri-1="https://zenodo.org/record/6224616/files/figure.png" httpUri-2="https://zenodo.org/record/6224624/files/figure.png" httpUri-3="https://zenodo.org/record/6224632/files/figure.png" httpUri-4="https://zenodo.org/record/6224640/files/figure.png" httpUri-5="https://zenodo.org/record/6224652/files/figure.png" httpUri-6="https://zenodo.org/record/6224662/files/figure.png" httpUri-7="https://zenodo.org/record/6224666/files/figure.png" pageId="6" pageNumber="457">Figs 823</figureCitation>
,
<tableCitation id="C6C80378FF97FF9A6542FD2DBC70808C" box="[276,377,727,753]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="2.[151,239,151,175]" captionStartId-1="14.[151,239,476,502]" captionTargetBox-0="[166,1370,227,2028]" captionTargetPageId-0="2" captionTargetPageId-1="14" captionText-0="TABLE 1. Species included in molecular analysis indicating population code, locality (province, country), and GenBank" captionText-1="TABLE 2. Characters differentiating between the European Lepidocyrtus species of lignorum-group." httpUri-0="http://table.plazi.org/id/DF35664BFF93FF9E64C1FF6DBF2782AE" httpUri-1="http://table.plazi.org/id/DF35664BFF9FFF9264C1FE26B9C6838B" pageId="6" pageNumber="457" tableUuid-0="DF35664BFF93FF9E64C1FF6DBF2782AE" tableUuid-1="DF35664BFF9FFF9264C1FE26B9C6838B">Tabs 12</tableCitation>
</paragraph>
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Zoobank:
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urn:lsid:zoobank.org:act:
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</uri>
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<emphasis id="B93EEAD1FF97FF9A64C1FCB9BC4C8121" bold="true" box="[151,325,834,861]" pageId="6" pageNumber="457">Type material.</emphasis>
<materialsCitation id="3B223C9EFF97FF9A651DFCB9BEAD81D8" ID-GBIF-Occurrence="3470323310" collectingDate="2007-04-18" collectorName="E. Mateos." country="Spain" county="Barcelona" elevation="610" latitude="41.66449" location="Male" longLatPrecision="1" longitude="2.56279" municipality="Sant Celoni" pageId="6" pageNumber="457" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F18861FF97FF9A651DFCB9BCB18120" box="[331,440,835,861]" pageId="6" pageNumber="457" type="holotype">Holotype</typeStatus>
:
<location id="8E956018FF97FF9A6594FCB9BCF58120" LSID="urn:lsid:plazi:treatment:03E387D5FF97FF9164C1FD48BC3482A8:8E956018FF97FF9A6594FCB9BCF58120" box="[450,508,835,861]" country="Spain" county="Barcelona" latitude="41.66449" longLatPrecision="1" longitude="2.56279" municipality="Sant Celoni" name="Male" pageId="6" pageNumber="457">Male</location>
on two slides (CRBA-90744a (head) and CRBA-90744b(body)),
<location id="8E956018FF97FF9A608DFCB9BDFC81FC" LSID="urn:lsid:plazi:treatment:03E387D5FF97FF9164C1FD48BC3482A8:8E956018FF97FF9A608DFCB9BDFC81FC" country="Spain" county="Barcelona" latitude="41.66449" longLatPrecision="1" longitude="2.56279" municipality="Sant Celoni" name="Montnegre Natural Park" pageId="6" pageNumber="457">Montnegre Natural Park</location>
,
<collectingMunicipality id="6B91ACB9FF97FF9A6555FC9DBC8581FC" box="[259,396,871,897]" pageId="6" pageNumber="457">Sant Celoni</collectingMunicipality>
,
<collectingCounty id="62944E4FFF97FF9A65CDFC9DBF0481FC" box="[411,525,871,897]" pageId="6" pageNumber="457">Barcelona</collectingCounty>
(
<collectingCountry id="F35D7653FF97FF9A6649FC9DBF6D81FF" box="[543,612,871,898]" name="Spain" pageId="6" pageNumber="457">Spain</collectingCountry>
),
<elevation id="0067D1F0FF97FF9A662CFC9DBE8B81FC" box="[634,898,871,897]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="457" unit="m" value="610.0">
<quantity id="4CB29B26FF97FF9A662CFC9DBFCD81FC" box="[634,708,871,897]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="457" unit="m" value="610.0">610 m</quantity>
above sea level
</elevation>
, lat/long coordinates
<geoCoordinate id="EE7E5004FF97FF9A602AFC9DB9F181FC" box="[1148,1272,871,897]" degrees="41.66449" direction="north" orientation="latitude" pageId="6" pageNumber="457" precision="1" value="41.66449">N41.66449</geoCoordinate>
<geoCoordinate id="EE7E5004FF97FF9A6155FC9CB87B81FC" box="[1283,1394,870,897]" degrees="2.56279" direction="east" orientation="longitude" pageId="6" pageNumber="457" precision="1" value="2.56279">E2.56279</geoCoordinate>
, on soil litter under oak trees, hand collecting,
<date id="FFF41003FF97FF9A6620FC71BFE781D8" box="[630,750,907,933]" pageId="6" pageNumber="457" value="2007-04-18">
<collectingDate id="EFB0E9EBFF97FF9A6620FC71BFE781D8" box="[630,750,907,933]" pageId="6" pageNumber="457" value="2007-04-18">18.iv.2007</collectingDate>
</date>
, leg.
<collectorName id="26BF5315FF97FF9A677CFC71BEAD81D8" box="[810,932,907,933]" pageId="6" pageNumber="457">E. Mateos.</collectorName>
</materialsCitation>
<typeStatus id="54F18861FF97FF9A67FCFC71B91281D8" box="[938,1051,907,933]" pageId="6" pageNumber="457" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D4CFD4AFF97FF9A6070FC71B9B981D8" box="[1062,1200,907,933]" pageId="6" pageNumber="457" type="generic">8 specimens</specimenCount>
on slides (
<specimenCount id="9D4CFD4AFF97FF9A617CFC71B88A81D8" box="[1322,1411,907,933]" pageId="6" pageNumber="457" type="male">3 males</specimenCount>
,
<specimenCount id="9D4CFD4AFF97FF9A61D8FC71BDEF81B4" pageId="6" pageNumber="457" type="female">1 female</specimenCount>
, and 4 without visible sexual plate) and
<materialsCitation id="3B223C9EFF97FF9A66FCFC55B89081B4" ID-GBIF-Occurrence="3470323302" box="[682,1433,943,969]" collectingDate="2007-04-18" collectorName="E. Mateos." country="Spain" county="Barcelona" elevation="610" latitude="41.66449" location="Male" longLatPrecision="1" longitude="2.56279" municipality="Sant Celoni" pageId="6" pageNumber="457" specimenCount="17" typeStatus="paratype">
<specimenCount id="9D4CFD4AFF97FF9A66FCFC55BE4B81B4" box="[682,834,943,969]" pageId="6" pageNumber="457" type="generic">17 specimens</specimenCount>
preserved in absolute alcohol, same data as holotype
</materialsCitation>
.
<materialsCitation id="3B223C9EFF97FF9A64C1FC29BF868648" ID-GBIF-Occurrence="3470323301" collectionCode="LP" collectorName="E. Mateos'" country="Spain" county="University of Barcelona" location="Centre de Recursos de Biodiversitat Animal" municipality="Faculty of Biology" pageId="6" pageNumber="457" specimenCode="LP125" specimenCount="3" typeStatus="holotype">
<typeStatus id="54F18861FF97FF9A64C1FC29BDF68190" box="[151,255,979,1005]" pageId="6" pageNumber="457" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54F18861FF97FF9A6561FC29BC9E8190" box="[311,407,979,1005]" pageId="6" pageNumber="457" type="paratype">paratype</typeStatus>
slides CRBA-90745a and CRBA-90745b saved in the collection of the
<location id="8E956018FF97FF9A6094FC29BCA5866C" LSID="urn:lsid:plazi:treatment:03E387D5FF97FF9164C1FD48BC3482A8:8E956018FF97FF9A6094FC29BCA5866C" country="Spain" county="University of Barcelona" municipality="Faculty of Biology" name="Centre de Recursos de Biodiversitat Animal" pageId="6" pageNumber="457">Centre de Recursos de Biodiversitat Animal</location>
,
<collectingMunicipality id="6B91ACB9FF97FF9A65ECFC0DBF9C866C" box="[442,661,1015,1041]" pageId="6" pageNumber="457">Faculty of Biology</collectingMunicipality>
,
<collectingCounty id="62944E4FFF97FF9A66F2FC0DBEB1866C" box="[676,952,1015,1041]" pageId="6" pageNumber="457">University of Barcelona</collectingCounty>
(http://www.crba.ub.edu); other
<typeStatus id="54F18861FF97FF9A6167FC0DB895866C" box="[1329,1436,1015,1041]" pageId="6" pageNumber="457" type="paratype">paratypes</typeStatus>
kept in the
<collectorName id="26BF5315FF97FF9A6542FBE1BC878648" box="[276,398,1051,1077]" pageId="6" pageNumber="457">E. Mateos</collectorName>
collection (lot
<specimenCode id="DBEC9EB8FF97FF9A6661FBE0BF8E8648" box="[567,647,1050,1077]" collectionCode="LP" country="Russia" name="Laboratory of Palaeontology" pageId="6" pageNumber="457">LP125</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF536C3FF97FF9A6491FBC4BEC686DC" blockId="6.[151,1437,834,1185]" pageId="6" pageNumber="457">
<emphasis id="B93EEAD1FF97FF9A6491FBC4BC888625" bold="true" box="[199,385,1086,1112]" pageId="6" pageNumber="457">Other material.</emphasis>
<materialsCitation id="3B223C9EFF97FF9A65DEFBC5BEC286DC" ID-GBIF-Occurrence="3470323312" collectingDate="2007-04-18" collectionCode="LP" collectorName="E. Mateos. All &amp; E. Mateos'" country="Spain" county="Barcelona" elevation="427" latitude="41.6663" location="Montnegre Natural Park" longLatPrecision="6" longitude="2.5793" municipality="Sant Celoni" pageId="6" pageNumber="457" specimenCode="LP134" specimenCount="40">
<specimenCount id="9D4CFD4AFF97FF9A65DEFBC5BF298624" box="[392,544,1087,1113]" pageId="6" pageNumber="457" type="generic">40 specimens</specimenCount>
preserved in absolute alcohol,
<location id="8E956018FF97FF9A672AFBC5B9878624" LSID="urn:lsid:plazi:treatment:03E387D5FF97FF9164C1FD48BC3482A8:8E956018FF97FF9A672AFBC5B9878624" box="[892,1166,1087,1113]" country="Spain" county="Barcelona" latitude="41.6663" longLatPrecision="6" longitude="2.5793" municipality="Sant Celoni" name="Montnegre Natural Park" pageId="6" pageNumber="457">Montnegre Natural Park</location>
,
<collectingMunicipality id="6B91ACB9FF97FF9A60CFFBC5B8168624" box="[1177,1311,1087,1113]" pageId="6" pageNumber="457">Sant Celoni</collectingMunicipality>
,
<collectingCounty id="62944E4FFF97FF9A617DFBC5B8948624" box="[1323,1437,1087,1113]" pageId="6" pageNumber="457">Barcelona</collectingCounty>
(
<collectingCountry id="F35D7653FF97FF9A64C9FB99BDED8603" box="[159,228,1123,1150]" name="Spain" pageId="6" pageNumber="457">Spain</collectingCountry>
),
<elevation id="0067D1F0FF97FF9A64AFFB99BCF08600" box="[249,505,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="4.27" pageId="6" pageNumber="457" unit="m" value="427.0">
<quantity id="4CB29B26FF97FF9A64AFFB99BC488600" box="[249,321,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="4.27" pageId="6" pageNumber="457" unit="m" value="427.0">427 m</quantity>
above sea level
</elevation>
, lat/long coordinates
<geoCoordinate id="EE7E5004FF97FF9A66BAFB99BE528600" box="[748,859,1123,1149]" degrees="41.6663" direction="north" orientation="latitude" pageId="6" pageNumber="457" precision="5" value="41.6663">N41.6663</geoCoordinate>
<geoCoordinate id="EE7E5004FF97FF9A6732FB98BECD8600" box="[868,964,1122,1149]" degrees="2.5793" direction="east" orientation="longitude" pageId="6" pageNumber="457" precision="5" value="2.5793">E2.5793</geoCoordinate>
, on soil litter under oak trees,
<date id="FFF41003FF97FF9A6149FB99B89E8600" box="[1311,1431,1123,1149]" pageId="6" pageNumber="457" value="2007-04-18">
<collectingDate id="EFB0E9EBFF97FF9A6149FB99B89E8600" box="[1311,1431,1123,1149]" pageId="6" pageNumber="457" value="2007-04-18">18.iv.2007</collectingDate>
</date>
, leg.
<collectorName id="26BF5315FF97FF9A6491FB7DBC6086DC" box="[199,361,1159,1185]" pageId="6" pageNumber="457">E. Mateos. All</collectorName>
material kept in the
<collectorName id="26BF5315FF97FF9A6606FB7DBFC086DC" box="[592,713,1159,1185]" pageId="6" pageNumber="457">E. Mateos</collectorName>
collection (lot
<specimenCode id="DBEC9EB8FF97FF9A6725FB7DBECB86DC" box="[883,962,1159,1185]" collectionCode="LP" country="Russia" name="Laboratory of Palaeontology" pageId="6" pageNumber="457">LP134</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF35664BFF97FF9A64C1F880B9BA85EF" ID-DOI="http://doi.org/10.5281/zenodo.6224610" ID-Zenodo-Dep="6224610" box="[151,1203,1914,1938]" httpUri="https://zenodo.org/record/6224610/files/figure.png" pageId="6" pageNumber="457" startId="6.[151,250,1914,1938]" targetBox="[152,1436,1209,1891]" targetPageId="6">
<paragraph id="8BF536C3FF97FF9A64C1F880B9BA85EF" blockId="6.[151,1203,1914,1938]" box="[151,1203,1914,1938]" pageId="6" pageNumber="457">
<emphasis id="B93EEAD1FF97FF9A64C1F880BC1A85EF" bold="true" box="[151,275,1914,1938]" pageId="6" pageNumber="457">FIGURE 8.</emphasis>
<taxonomicName id="4C4A4D40FF97FF9A654FF880BF3D85EF" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[281,564,1914,1938]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF97FF9A654FF880BF3D85EF" box="[281,564,1914,1938]" italics="true" pageId="6" pageNumber="457">Lepidocyrtus fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF97FF9A666CF880BF8385EF" bold="true" box="[570,650,1914,1938]" pageId="6" pageNumber="457">
<taxonomicNameLabel id="A20D57AAFF97FF9A666CF880BF8385EF" box="[570,650,1914,1938]" pageId="6" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: Habitus lateral (specimen in alcohol, without scales).
</paragraph>
</caption>
<subSubSection id="C3506548FF96FF9B6491FF6CB88383F5" pageId="7" pageNumber="458" type="diagnosis">
<paragraph id="8BF536C3FF96FF9B6491FF6CBC87833D" blockId="7.[151,1437,150,789]" pageId="7" pageNumber="458">
<emphasis id="B93EEAD1FF96FF9B6491FF6CBC4B82CD" bold="true" box="[199,322,150,176]" pageId="7" pageNumber="458">Diagnosis.</emphasis>
With dark blue pigment on all head, Ant.IIIV, coxae-trocanter-femur of legs IIII, and ventral tube; disperse blue pigment also on Abd.IV and Abd.V. Th.II slightly projecting over the head.Ant.IIII, legs, ventral tube and posterior region of manubrium with scales. Labial chaetotaxy
<emphasis id="B93EEAD1FF96FF9B672CFF24B9158283" bold="true" box="[890,1052,222,254]" pageId="7" pageNumber="458">
M
<subScript id="17CE3486FF96FF9B67C3FF0ABE948283" attach="right" box="[917,925,240,254]" fontSize="6" pageId="7" pageNumber="458">1</subScript>
M
<subScript id="17CE3486FF96FF9B67E1FF0ABEB68283" attach="both" box="[951,959,240,254]" fontSize="6" pageId="7" pageNumber="458">2</subScript>
REL
<subScript id="17CE3486FF96FF9B67AFFF0AB9088283" attach="both" box="[1017,1025,240,254]" fontSize="6" pageId="7" pageNumber="458">1</subScript>
L
<subScript id="17CE3486FF96FF9B6042FF0AB9158283" attach="left" box="[1044,1052,240,254]" fontSize="6" pageId="7" pageNumber="458">2</subScript>
</emphasis>
,
<emphasis id="B93EEAD1FF96FF9B607CFF24B9378285" bold="true" box="[1066,1086,222,248]" pageId="7" pageNumber="458">R</emphasis>
shortened. Dorsal cephalic and body macrochaetae formula
<emphasis id="B93EEAD1FF96FF9B658EFEF8BE1D8361" bold="true" box="[472,788,258,291]" pageId="7" pageNumber="458">
A
<subScript id="17CE3486FF96FF9B65BAFEEEBCFD835F" attach="left" box="[492,500,276,290]" fontSize="6" pageId="7" pageNumber="458">0</subScript>
[A
<subScript id="17CE3486FF96FF9B6647FEEEBF2B835F" attach="left" box="[529,546,276,290]" fontSize="6" pageId="7" pageNumber="458">2a</subScript>
]A
<subScript id="17CE3486FF96FF9B6669FEEEBF4E835F" attach="both" box="[575,583,276,290]" fontSize="6" pageId="7" pageNumber="458">2</subScript>
A
<subScript id="17CE3486FF96FF9B660AFEEEBF6D835F" attach="right" box="[604,612,276,290]" fontSize="6" pageId="7" pageNumber="458">3</subScript>
Pa
<subScript id="17CE3486FF96FF9B66D5FEEFBF82835E" attach="left" box="[643,651,277,291]" fontSize="6" pageId="7" pageNumber="458">5</subScript>
/00/0101+3.
</emphasis>
Abd.IV without chaeta
<emphasis id="B93EEAD1FF96FF9B6076FEF8B9228361" bold="true" box="[1056,1067,258,284]" pageId="7" pageNumber="458">s</emphasis>
. Unguiculus lanceolate and with serrated outer margin.
</paragraph>
<paragraph id="8BF536C3FF96FF9B6491FEB0B88383F5" blockId="7.[151,1437,150,789]" pageId="7" pageNumber="458">
<emphasis id="B93EEAD1FF96FF9B6491FEB0BCC98319" bold="true" box="[199,448,330,356]" pageId="7" pageNumber="458">Molecular diagnosis.</emphasis>
This species includes all populations that cluster with CoxII and EF sequences of the individuals LP125-1 to LP125-5 (
<tableCitation id="C6C80378FF96FF9B6647FE94BF6C83F5" box="[529,613,366,392]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionTargetBox="[166,1370,227,2028]" captionTargetPageId="2" captionText="TABLE 1. Species included in molecular analysis indicating population code, locality (province, country), and GenBank" httpUri="http://table.plazi.org/id/DF35664BFF93FF9E64C1FF6DBF2782AE" pageId="7" pageNumber="458" tableUuid="DF35664BFF93FF9E64C1FF6DBF2782AE">Table 1</tableCitation>
), with significant support in an adequate molecular delimitation model.
</paragraph>
</subSubSection>
<subSubSection id="C3506548FF96FF9B6491FE68BCAD83AD" pageId="7" pageNumber="458" type="etymology">
<paragraph id="8BF536C3FF96FF9B6491FE68BCAD83AD" blockId="7.[151,1437,150,789]" pageId="7" pageNumber="458">
<emphasis id="B93EEAD1FF96FF9B6491FE68BC4783D1" bold="true" box="[199,334,402,428]" pageId="7" pageNumber="458">Etymology.</emphasis>
The species name refers to the dark blue colour of the head. In latin “fuscus” means dark, and “cephalus” means head.
</paragraph>
</subSubSection>
<subSubSection id="C3506548FF96FF906491FE20B908856C" lastPageId="12" lastPageNumber="463" pageId="7" pageNumber="458" type="description">
<paragraph id="8BF536C3FF96FF9B6491FE20B99E801D" blockId="7.[151,1437,150,789]" pageId="7" pageNumber="458">
<emphasis id="B93EEAD1FF96FF9B6491FE20BC588389" bold="true" box="[199,337,474,500]" pageId="7" pageNumber="458">Description</emphasis>
.
<typeStatus id="54F18861FF96FF9B6536FE20BCC18389" box="[352,456,474,500]" pageId="7" pageNumber="458" type="holotype">Holotype</typeStatus>
body length (without head nor furca)
<quantity id="4CB29B26FF96FF9B6725FE20BEC28388" box="[883,971,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="458" unit="mm" value="2.2">2.2 mm</quantity>
,
<typeStatus id="54F18861FF96FF9B6781FE21B94B8388" box="[983,1090,475,501]" pageId="7" pageNumber="458" type="paratype">paratypes</typeStatus>
<quantity id="4CB29B26FF96FF9B601CFE20B9DD8388" box="[1098,1236,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="7" pageNumber="458" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
. Body colour pattern (
<figureCitation id="13712A46FF96FF9B6484FE04BC1C8065" box="[210,277,510,536]" captionStart="FIGURE 8" captionStartId="6.[151,250,1914,1938]" captionTargetBox="[152,1436,1209,1891]" captionTargetId="figure-385@6.[152,1436,1209,1891]" captionTargetPageId="6" captionText="FIGURE 8. Lepidocyrtus fuscocephalus sp. nov.: Habitus lateral (specimen in alcohol, without scales)." figureDoi="http://doi.org/10.5281/zenodo.6224610" httpUri="https://zenodo.org/record/6224610/files/figure.png" pageId="7" pageNumber="458">Fig. 8</figureCitation>
) with dark blue pigment on all head, Ant.IIIV (with increasing colour intensity towards the distal part of each segment), coxae, and ventral tube; disperse blue pigment also on trochanter-femur of legs IIII, Abd.IV and Abd.V; densely black pigmented ocular areas. Mesothorax slightly projected over the head.
</paragraph>
<paragraph id="8BF536C3FF96FF9B6491FD90B8668169" blockId="7.[151,1437,150,789]" pageId="7" pageNumber="458">Antenna with scales on dorsal and ventral region of Ant.IIII. Ratio antenna:cephalic diagonal = 1.81.9 (head diagonal measured from cervical edge to apex of mouth part); ratio Ant.I:II:III:IV as 1:1.7:1.6:2.5. Basis of Ant.I dorsally with three microchaetae arranged in triangle (Ant.I-organ); apex of Ant.I with a short curved S-chaeta in the membranous area of the ventral region. Ant.III organ composed of two subcilindrical and curved sensory rods. Ant. IV without apical bulb. Eyes 8+8; eyes A to F of equal size, G and H slightly smaller, ratio A/G and A/H = 1.6.</paragraph>
<caption id="DF35664BFF96FF9B64C1F873B94E8597" ID-DOI="http://doi.org/10.5281/zenodo.6224616" ID-Zenodo-Dep="6224616" httpUri="https://zenodo.org/record/6224616/files/figure.png" pageId="7" pageNumber="458" startId="7.[151,264,1929,1953]" targetBox="[151,1436,812,1906]" targetPageId="7">
<paragraph id="8BF536C3FF96FF9B64C1F873B94E8597" blockId="7.[151,1437,1929,2026]" pageId="7" pageNumber="458">
<emphasis id="B93EEAD1FF96FF9B64C1F873BC4F85DF" bold="true" box="[151,326,1929,1954]" pageId="7" pageNumber="458">FIGURES 912.</emphasis>
<taxonomicName id="4C4A4D40FF96FF9B651DF870BF6D85DF" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[331,612,1930,1954]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF96FF9B651DF870BF6D85DF" box="[331,612,1930,1954]" italics="true" pageId="7" pageNumber="458">Lepidocyrtus fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF96FF9B663FF870BFB185DF" bold="true" box="[617,696,1930,1954]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="A20D57AAFF96FF9B663FF870BFB185DF" box="[617,696,1930,1954]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 9, clypeal and labral chaetotaxy; 10, maxillary palp (right side); 11, outer labial papilla (right side); 12, labial and postlabial chaetotaxy (the arrow points to a supernumerary M chaeta present in one paratype). Black dotsciliated chaetae, crossscales, p.c.—proximal chaetae of labial palp.
</paragraph>
</caption>
<caption id="DF35664BFF99FF9464C1F8A8BF8B85F3" ID-DOI="http://doi.org/10.5281/zenodo.6224624" ID-Zenodo-Dep="6224624" httpUri="https://zenodo.org/record/6224624/files/figure.png" pageId="8" pageNumber="459" startId="8.[151,250,1874,1898]" targetBox="[151,1436,181,1850]" targetPageId="8">
<paragraph id="8BF536C3FF99FF9464C1F8A8BF8B85F3" blockId="8.[151,1436,1874,1934]" pageId="8" pageNumber="459">
<emphasis id="B93EEAD1FF99FF9464C1F8A8BC298517" bold="true" box="[151,288,1874,1898]" pageId="8" pageNumber="459">FIGURE 13.</emphasis>
<taxonomicName id="4C4A4D40FF99FF946571F8A8BF4B8517" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[295,578,1874,1898]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF99FF946571F8A8BF4B8517" box="[295,578,1874,1898]" italics="true" pageId="8" pageNumber="459">Lepidocyrtus fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF99FF94661EF8A8BF908517" bold="true" box="[584,665,1874,1898]" pageId="8" pageNumber="459">
<taxonomicNameLabel id="A20D57AAFF99FF94661EF8A8BF908517" box="[584,665,1874,1898]" pageId="8" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: Dorsal head chaetotaxy (left side). Broad circleslong ciliated macrochaetae, small circlesshort ciliated macrochaetae.
</paragraph>
</caption>
<caption id="DF35664BFF98FF9564C1F86DBC3285A9" ID-DOI="http://doi.org/10.5281/zenodo.6224632" ID-Zenodo-Dep="6224632" httpUri="https://zenodo.org/record/6224632/files/figure.png" pageId="9" pageNumber="460" startId="9.[151,264,1943,1967]" targetBox="[169,1414,183,1919]" targetPageId="9">
<paragraph id="8BF536C3FF98FF9564C1F86DBC3285A9" blockId="9.[151,1436,1943,2004]" pageId="9" pageNumber="460">
<emphasis id="B93EEAD1FF98FF9564C1F86DBC5C85D2" bold="true" box="[151,341,1943,1967]" pageId="9" pageNumber="460">FIGURES 1416.</emphasis>
<taxonomicName id="4C4A4D40FF98FF95650DF862BF7F85CD" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[347,630,1944,1968]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF98FF95650DF862BF7F85CD" box="[347,630,1944,1968]" italics="true" pageId="9" pageNumber="460">Lepidocyrtus fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF98FF95662AF862BFC485CD" bold="true" box="[636,717,1944,1968]" pageId="9" pageNumber="460">
<taxonomicNameLabel id="A20D57AAFF98FF95662AF862BFC485CD" box="[636,717,1944,1968]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
dorsal chaetotaxy (left side): 14, Th.II; 15, Th.III; 16 Abd.I. Circlesciliated chaetae.
</paragraph>
</caption>
<caption id="DF35664BFF9BFF9664C1FA68BE6387B2" ID-DOI="http://doi.org/10.5281/zenodo.6224640" ID-Zenodo-Dep="6224640" httpUri="https://zenodo.org/record/6224640/files/figure.png" pageId="10" pageNumber="461" startId="10.[151,264,1426,1450]" targetBox="[156,1429,190,1402]" targetPageId="10">
<paragraph id="8BF536C3FF9BFF9664C1FA68BE6387B2" blockId="10.[151,1436,1426,1487]" pageId="10" pageNumber="461">
<emphasis id="B93EEAD1FF9BFF9664C1FA68BC5C87D7" bold="true" box="[151,341,1426,1450]" pageId="10" pageNumber="461">FIGURES 1718.</emphasis>
<taxonomicName id="4C4A4D40FF9BFF96650BFA69BF7187D6" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[349,632,1427,1451]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF9BFF96650BFA69BF7187D6" box="[349,632,1427,1451]" italics="true" pageId="10" pageNumber="461">Lepidocyrtus fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF9BFF966629FA69BFD887D6" bold="true" box="[639,721,1427,1451]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="A20D57AAFF9BFF966629FA69BFD887D6" box="[639,721,1427,1451]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
dorsal chaetotaxy (left side): 17, Abd.II; 18, Abd.III. Broad circlesbroad ciliated macrochaetae, small circlesthin ciliated macrochaetae.
</paragraph>
</caption>
<paragraph id="8BF536C3FF9BFF966491FA07BEBB84B6" blockId="10.[151,1437,1533,2028]" pageId="10" pageNumber="461">
Clypeus (
<figureCitation id="13712A46FF9BFF966564FA07BC7C846A" box="[306,373,1533,1559]" captionStart="FIGURES 912" captionStartId="7.[151,264,1929,1953]" captionTargetBox="[151,1436,812,1906]" captionTargetId="figure-313@7.[151,1436,812,1906]" captionTargetPageId="7" captionText="FIGURES 912. Lepidocyrtus fuscocephalus sp. nov.: 9, clypeal and labral chaetotaxy; 10, maxillary palp (right side); 11, outer labial papilla (right side); 12, labial and postlabial chaetotaxy (the arrow points to a supernumerary M chaeta present in one paratype). Black dotsciliated chaetae, crossscales, p.c.—proximal chaetae of labial palp." figureDoi="http://doi.org/10.5281/zenodo.6224616" httpUri="https://zenodo.org/record/6224616/files/figure.png" pageId="10" pageNumber="461">Fig. 9</figureCitation>
) with three prefrontal chaetae (1 pf0 and 2 pf1), four facial chaetae (f), and four lateral chaetae (2 L1 and 2 L2), all these chaetae ciliated. Prelabral and labral chaetae in typical number 4/554 (
<figureCitation id="13712A46FF9BFF96608BF9DBB82A8446" box="[1245,1315,1569,1595]" captionStart="FIGURES 912" captionStartId="7.[151,264,1929,1953]" captionTargetBox="[151,1436,812,1906]" captionTargetId="figure-313@7.[151,1436,812,1906]" captionTargetPageId="7" captionText="FIGURES 912. Lepidocyrtus fuscocephalus sp. nov.: 9, clypeal and labral chaetotaxy; 10, maxillary palp (right side); 11, outer labial papilla (right side); 12, labial and postlabial chaetotaxy (the arrow points to a supernumerary M chaeta present in one paratype). Black dotsciliated chaetae, crossscales, p.c.—proximal chaetae of labial palp." figureDoi="http://doi.org/10.5281/zenodo.6224616" httpUri="https://zenodo.org/record/6224616/files/figure.png" pageId="10" pageNumber="461">Fig. 9</figureCitation>
), prelabral chaetae ciliated, all labral chaetae smooth and pointed, apical row curved; inverted U-shaped labral apical intrusion; four rounded labral papillae with a central small pointed expansion. Maxillary palp outer lobe with smooth apical appendage and basal chaeta, and three smooth sublobal appendages (
<figureCitation id="13712A46FF9BFF9667C1F977BEE084DA" box="[919,1001,1677,1703]" captionStart="FIGURES 912" captionStartId="7.[151,264,1929,1953]" captionTargetBox="[151,1436,812,1906]" captionTargetId="figure-313@7.[151,1436,812,1906]" captionTargetPageId="7" captionText="FIGURES 912. Lepidocyrtus fuscocephalus sp. nov.: 9, clypeal and labral chaetotaxy; 10, maxillary palp (right side); 11, outer labial papilla (right side); 12, labial and postlabial chaetotaxy (the arrow points to a supernumerary M chaeta present in one paratype). Black dotsciliated chaetae, crossscales, p.c.—proximal chaetae of labial palp." figureDoi="http://doi.org/10.5281/zenodo.6224616" httpUri="https://zenodo.org/record/6224616/files/figure.png" pageId="10" pageNumber="461">Fig. 10</figureCitation>
). Lateral process of outer labial papilla finger-shape, slightly curved, tip not reaching apex of papilla (
<figureCitation id="13712A46FF9BFF966704F94BBEAD84B6" box="[850,932,1713,1739]" captionStart="FIGURES 912" captionStartId="7.[151,264,1929,1953]" captionTargetBox="[151,1436,812,1906]" captionTargetId="figure-313@7.[151,1436,812,1906]" captionTargetPageId="7" captionText="FIGURES 912. Lepidocyrtus fuscocephalus sp. nov.: 9, clypeal and labral chaetotaxy; 10, maxillary palp (right side); 11, outer labial papilla (right side); 12, labial and postlabial chaetotaxy (the arrow points to a supernumerary M chaeta present in one paratype). Black dotsciliated chaetae, crossscales, p.c.—proximal chaetae of labial palp." figureDoi="http://doi.org/10.5281/zenodo.6224616" httpUri="https://zenodo.org/record/6224616/files/figure.png" pageId="10" pageNumber="461">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BF536C3FF9BFF966491F92FBFAF8502" blockId="10.[151,1437,1533,2028]" pageId="10" pageNumber="461">
Labial and postlabial chaetotaxy as in
<figureCitation id="13712A46FF9BFF9666D7F92FBFDC8492" box="[641,725,1749,1775]" captionStart="FIGURES 912" captionStartId="7.[151,264,1929,1953]" captionTargetBox="[151,1436,812,1906]" captionTargetId="figure-313@7.[151,1436,812,1906]" captionTargetPageId="7" captionText="FIGURES 912. Lepidocyrtus fuscocephalus sp. nov.: 9, clypeal and labral chaetotaxy; 10, maxillary palp (right side); 11, outer labial papilla (right side); 12, labial and postlabial chaetotaxy (the arrow points to a supernumerary M chaeta present in one paratype). Black dotsciliated chaetae, crossscales, p.c.—proximal chaetae of labial palp." figureDoi="http://doi.org/10.5281/zenodo.6224616" httpUri="https://zenodo.org/record/6224616/files/figure.png" pageId="10" pageNumber="461">Fig. 12</figureCitation>
; with five smooth proximal chaetae at the base of labial palp; labial anterior row formed by five smooth chaetae (a1a5); posterior row formed by ciliated chaetae with formula
<emphasis id="B93EEAD1FF9BFF9664C1F8E7BC308540" bold="true" box="[151,313,1821,1853]" pageId="10" pageNumber="461">
M
<subScript id="17CE3486FF9BFF9664E4F8D5BDB38540" attach="right" box="[178,186,1839,1853]" fontSize="6" pageId="10" pageNumber="461">1</subScript>
M
<subScript id="17CE3486FF9BFF966482F8D5BDD58540" attach="both" box="[212,220,1839,1853]" fontSize="6" pageId="10" pageNumber="461">2</subScript>
REL
<subScript id="17CE3486FF9BFF966540F8D5BC178540" attach="both" box="[278,286,1839,1853]" fontSize="6" pageId="10" pageNumber="461">1</subScript>
L
<subScript id="17CE3486FF9BFF966567F8D5BC308540" attach="left" box="[305,313,1839,1853]" fontSize="6" pageId="10" pageNumber="461">2</subScript>
</emphasis>
; chaeta
<emphasis id="B93EEAD1FF9BFF9665CEF8E7BCA5854A" bold="true" box="[408,428,1821,1847]" pageId="10" pageNumber="461">R</emphasis>
shorter, ratio
<emphasis id="B93EEAD1FF9BFF96661CF8E7BF80854A" bold="true" box="[586,649,1821,1853]" pageId="10" pageNumber="461">
M
<subScript id="17CE3486FF9BFF966633F8D5BF648540" attach="right" box="[613,621,1839,1853]" fontSize="6" pageId="10" pageNumber="461">2</subScript>
/R
</emphasis>
≈ 2.4;
<specimenCount id="9D4CFD4AFF9BFF96668CF8E7BE62854A" box="[730,875,1821,1847]" pageId="10" pageNumber="461" type="generic" typeStatus="paratype">one paratype</specimenCount>
with a supernumerary reduced
<emphasis id="B93EEAD1FF9BFF966087F8E7B9E2854A" bold="true" box="[1233,1259,1821,1847]" pageId="10" pageNumber="461">M</emphasis>
chaeta (as long as chaeta
<emphasis id="B93EEAD1FF9BFF966550F8BBBC138526" bold="true" box="[262,282,1857,1883]" pageId="10" pageNumber="461">R</emphasis>
) between
<emphasis id="B93EEAD1FF9BFF9665C6F8BBBCBB851C" bold="true" box="[400,434,1857,1889]" pageId="10" pageNumber="461">
M
<subScript id="17CE3486FF9BFF9665FCF8A9BCBB851C" attach="left" box="[426,434,1875,1889]" fontSize="6" pageId="10" pageNumber="461">1</subScript>
</emphasis>
and
<emphasis id="B93EEAD1FF9BFF9665B1F8BBBF03851C" bold="true" box="[487,522,1857,1889]" pageId="10" pageNumber="461">
M
<subScript id="17CE3486FF9BFF966654F8A9BF03851C" attach="left" box="[514,522,1875,1889]" fontSize="6" pageId="10" pageNumber="461">2</subScript>
</emphasis>
present on one side (
<figureCitation id="13712A46FF9BFF9666ADF8BBBE598526" box="[763,848,1857,1883]" captionStart="FIGURES 912" captionStartId="7.[151,264,1929,1953]" captionTargetBox="[151,1436,812,1906]" captionTargetId="figure-313@7.[151,1436,812,1906]" captionTargetPageId="7" captionText="FIGURES 912. Lepidocyrtus fuscocephalus sp. nov.: 9, clypeal and labral chaetotaxy; 10, maxillary palp (right side); 11, outer labial papilla (right side); 12, labial and postlabial chaetotaxy (the arrow points to a supernumerary M chaeta present in one paratype). Black dotsciliated chaetae, crossscales, p.c.—proximal chaetae of labial palp." figureDoi="http://doi.org/10.5281/zenodo.6224616" httpUri="https://zenodo.org/record/6224616/files/figure.png" pageId="10" pageNumber="461">Fig. 12</figureCitation>
); postlabial chaetaxy with all chaetae ciliated, row I (along ventral cephalic groove) with 4 chaetae.
</paragraph>
<paragraph id="8BF536C3FF9BFF966491F873BF568596" blockId="10.[151,1437,1533,2028]" pageId="10" pageNumber="461">
Dorsal cephalic macrochaetae formula
<emphasis id="B93EEAD1FF9BFF9666D4F873BFF785D4" bold="true" box="[642,766,1929,1961]" pageId="10" pageNumber="461">
A
<subScript id="17CE3486FF9BFF9666C0F861BF9785D4" attach="both" box="[662,670,1947,1961]" fontSize="6" pageId="10" pageNumber="461">0</subScript>
A
<subScript id="17CE3486FF9BFF9666E4F861BFB385D4" attach="both" box="[690,698,1947,1961]" fontSize="6" pageId="10" pageNumber="461">2</subScript>
A
<subScript id="17CE3486FF9BFF966699F861BFDE85D4" attach="both" box="[719,727,1947,1961]" fontSize="6" pageId="10" pageNumber="461">3</subScript>
Pa
<subScript id="17CE3486FF9BFF9666A0F861BFF785D4" attach="left" box="[758,766,1947,1961]" fontSize="6" pageId="10" pageNumber="461">5</subScript>
</emphasis>
, but also with pair of smaller supplementary macrochaetae
<emphasis id="B93EEAD1FF9BFF9664C1F857BDB585B0" bold="true" box="[151,188,1965,1997]" pageId="10" pageNumber="461">
A
<subScript id="17CE3486FF9BFF9664FDF845BDB585B0" attach="left" box="[171,188,1983,1997]" fontSize="6" pageId="10" pageNumber="461">2a</subScript>
</emphasis>
between
<emphasis id="B93EEAD1FF9BFF96657EF857BC4C85B0" bold="true" box="[296,325,1965,1997]" pageId="10" pageNumber="461">
A
<subScript id="17CE3486FF9BFF96656BF845BC4C85B0" attach="left" box="[317,325,1983,1997]" fontSize="6" pageId="10" pageNumber="461">0</subScript>
</emphasis>
and
<emphasis id="B93EEAD1FF9BFF96652FF857BC9F85B0" bold="true" box="[377,406,1965,1997]" pageId="10" pageNumber="461">
A
<subScript id="17CE3486FF9BFF9665D8F845BC9F85B0" attach="right" box="[398,406,1983,1997]" fontSize="6" pageId="10" pageNumber="461">2</subScript>
</emphasis>
; maximum number of macrochaetae
<emphasis id="B93EEAD1FF9BFF96676AF857BE6985BA" bold="true" box="[828,864,1965,1991]" pageId="10" pageNumber="461">An</emphasis>
on head 15+15. Interocular chaetotaxy with
<emphasis id="B93EEAD1FF9BFF96610DF857B86F85BA" bold="true" box="[1371,1382,1965,1991]" pageId="10" pageNumber="461">s</emphasis>
,
<emphasis id="B93EEAD1FF9BFF966123F857B87785BA" bold="true" box="[1397,1406,1965,1991]" pageId="10" pageNumber="461">t</emphasis>
,
<emphasis id="B93EEAD1FF9BFF9661DBF857B89485BA" bold="true" box="[1421,1437,1965,1991]" pageId="10" pageNumber="461">p</emphasis>
ciliated chaetae and 57 scales (
<figureCitation id="13712A46FF9BFF9665A8F82BBF5B8596" box="[510,594,2001,2027]" captionStart="FIGURE 13" captionStartId="8.[151,250,1874,1898]" captionTargetBox="[151,1436,181,1850]" captionTargetId="figure-34@8.[151,1436,181,1850]" captionTargetPageId="8" captionText="FIGURE 13. Lepidocyrtus fuscocephalus sp. nov.: Dorsal head chaetotaxy (left side). Broad circleslong ciliated macrochaetae, small circlesshort ciliated macrochaetae." figureDoi="http://doi.org/10.5281/zenodo.6224624" httpUri="https://zenodo.org/record/6224624/files/figure.png" pageId="10" pageNumber="461">Fig. 13</figureCitation>
).
</paragraph>
<caption id="DF35664BFF9AFF9764C1F86DBFDE85A9" ID-DOI="http://doi.org/10.5281/zenodo.6224652" ID-Zenodo-Dep="6224652" httpUri="https://zenodo.org/record/6224652/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,250,1943,1967]" targetBox="[243,1320,181,1919]" targetPageId="11">
<paragraph id="8BF536C3FF9AFF9764C1F86DBFDE85A9" blockId="11.[151,1436,1943,2004]" pageId="11" pageNumber="462">
<emphasis id="B93EEAD1FF9AFF9764C1F86DBC2885D2" bold="true" box="[151,289,1943,1967]" pageId="11" pageNumber="462">FIGURE 19.</emphasis>
<taxonomicName id="4C4A4D40FF9AFF97657EF862BF4D85CD" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[296,580,1944,1968]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF9AFF97657EF862BF4D85CD" box="[296,580,1944,1968]" italics="true" pageId="11" pageNumber="462">Lepidocyrtus fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF9AFF97661DF862BF9585CD" bold="true" box="[587,668,1944,1968]" pageId="11" pageNumber="462">
<taxonomicNameLabel id="A20D57AAFF9AFF97661DF862BF9585CD" box="[587,668,1944,1968]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: Abd.IV chaetotaxy (left side). Broad black circlesbroad ciliated macrochaetae, small black circlesthin ciliated macrochaetae.
</paragraph>
</caption>
<caption id="DF35664BFF9DFF9064C1FD83B9D180C8" ID-DOI="http://doi.org/10.5281/zenodo.6224662" ID-Zenodo-Dep="6224662" httpUri="https://zenodo.org/record/6224662/files/figure.png" pageId="12" pageNumber="463" startId="12.[151,264,633,657]" targetBox="[151,1436,189,609]" targetPageId="12">
<paragraph id="8BF536C3FF9DFF9064C1FD83B9D180C8" blockId="12.[151,1436,633,693]" pageId="12" pageNumber="463">
<emphasis id="B93EEAD1FF9DFF9064C1FD83BC5D80EC" bold="true" box="[151,340,633,657]" pageId="12" pageNumber="463">FIGURES 2022.</emphasis>
<taxonomicName id="4C4A4D40FF9DFF90650CFD83BF7C80EC" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[346,629,633,657]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF9DFF90650CFD83BF7C80EC" box="[346,629,633,657]" italics="true" pageId="12" pageNumber="463">Lepidocyrtus fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF9DFF90662DFD83BFC280EC" bold="true" box="[635,715,633,657]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A20D57AAFF9DFF90662DFD83BFC280EC" box="[635,715,633,657]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 20, Abd.V chaetotaxy (left side), broad circleslong ciliated chaetae, small circlesshort ciliated or smooth chaetae; 21, trochanteral organ; 22, third leg unguis and unguiculus.
</paragraph>
</caption>
<paragraph id="8BF536C3FF9DFF906491FD19BC8B8764" blockId="12.[151,1437,739,2026]" pageId="12" pageNumber="463">
Dorsal body macrochaetae formula
<emphasis id="B93EEAD1FF9DFF90660FFD19BFDB8080" bold="true" box="[601,722,739,765]" pageId="12" pageNumber="463">00/0101+3</emphasis>
(macrochaetae
<emphasis id="B93EEAD1FF9DFF9067D2FD19BEA38080" bold="true" box="[900,938,739,765]" pageId="12" pageNumber="463">m3</emphasis>
on Abd.II, and
<emphasis id="B93EEAD1FF9DFF90600CFD19B8078080" bold="true" box="[1114,1294,739,766]" pageId="12" pageNumber="463">Sm+B4, B5, B6</emphasis>
on Abd.IV). Dorsal chaetotaxy of Th.IIIII and Abd.I as in
<figureCitation id="13712A46FF9DFF9066FCFCFDBE22815C" box="[682,811,775,801]" captionStart="FIGURES 1416" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[169,1414,183,1919]" captionTargetId="figure-17@9.[169,1418,181,1919]" captionTargetPageId="9" captionText="FIGURES 1416. Lepidocyrtus fuscocephalus sp. nov. dorsal chaetotaxy (left side): 14, Th.II; 15, Th.III; 16 Abd.I. Circlesciliated chaetae." figureDoi="http://doi.org/10.5281/zenodo.6224632" httpUri="https://zenodo.org/record/6224632/files/figure.png" pageId="12" pageNumber="463">Figs 1416</figureCitation>
. Th.II with 2 lateral S-chaetae (
<emphasis id="B93EEAD1FF9DFF9060C2FCFDB9A3815C" bold="true" box="[1172,1194,775,801]" pageId="12" pageNumber="463">al</emphasis>
and
<emphasis id="B93EEAD1FF9DFF9060B4FCFDB80D815C" bold="true" box="[1250,1284,775,801]" pageId="12" pageNumber="463">ms</emphasis>
) and without macrochaetae in dorsal position. Th.III with a lateral sensillum (
<emphasis id="B93EEAD1FF9DFF906724FCD1BE818138" bold="true" box="[882,904,811,837]" pageId="12" pageNumber="463">al</emphasis>
) close to several ciliated chaetae. Abd.I with a lateral S-microchaeta (
<emphasis id="B93EEAD1FF9DFF9065CFFCB5BCB28114" bold="true" box="[409,443,847,873]" pageId="12" pageNumber="463">ms</emphasis>
) external to
<emphasis id="B93EEAD1FF9DFF90661BFCB5BF608114" bold="true" box="[589,617,847,873]" pageId="12" pageNumber="463">a6</emphasis>
. Chaetotaxy of Abd.IIIII as in
<figureCitation id="13712A46FF9DFF906781FCB5B9518114" box="[983,1112,847,873]" captionStart="FIGURES 1718" captionStartId="10.[151,264,1426,1450]" captionTargetBox="[156,1429,190,1402]" captionTargetId="figure-34@10.[151,1436,181,1402]" captionTargetPageId="10" captionText="FIGURES 1718. Lepidocyrtus fuscocephalus sp. nov. dorsal chaetotaxy (left side): 17, Abd.II; 18, Abd.III. Broad circlesbroad ciliated macrochaetae, small circlesthin ciliated macrochaetae." figureDoi="http://doi.org/10.5281/zenodo.6224640" httpUri="https://zenodo.org/record/6224640/files/figure.png" pageId="12" pageNumber="463">Figs 1718</figureCitation>
. Abd.II chaeta
<emphasis id="B93EEAD1FF9DFF906153FCB5B82D8114" bold="true" box="[1285,1316,847,873]" pageId="12" pageNumber="463">ml</emphasis>
present or absent depending on the specimens; macrochaeta
<emphasis id="B93EEAD1FF9DFF90669EFC89BFE781F0" bold="true" box="[712,750,883,909]" pageId="12" pageNumber="463">m3</emphasis>
with socket diameter 1.6 times higher than macrochaeta
<emphasis id="B93EEAD1FF9DFF906126FC89B89C81F0" bold="true" box="[1392,1429,883,909]" pageId="12" pageNumber="463">m5</emphasis>
. Abd. III chaeta
<emphasis id="B93EEAD1FF9DFF90651FFC6DBC6181CC" bold="true" box="[329,360,919,945]" pageId="12" pageNumber="463">mi</emphasis>
present or absent depending on the specimens, with chaeta
<emphasis id="B93EEAD1FF9DFF90605AFC6DB92081CC" bold="true" box="[1036,1065,919,945]" pageId="12" pageNumber="463">d3</emphasis>
between macrochaetae
<emphasis id="B93EEAD1FF9DFF906161FC6DB86581CC" bold="true" box="[1335,1388,919,945]" pageId="12" pageNumber="463">pm6</emphasis>
and
<emphasis id="B93EEAD1FF9DFF9064C1FC41BDBC81A8" bold="true" box="[151,181,955,981]" pageId="12" pageNumber="463">p6</emphasis>
, and with S-chaetae
<emphasis id="B93EEAD1FF9DFF9065F5FC41BCB581A8" bold="true" box="[419,444,955,981]" pageId="12" pageNumber="463">as</emphasis>
and
<emphasis id="B93EEAD1FF9DFF9065A3FC41BF1E81A8" bold="true" box="[501,535,955,981]" pageId="12" pageNumber="463">ms</emphasis>
. All chaetae associated with the trichobothria on Abd.IIIII strongly ciliate and partially fan-shaped. Chaetotaxy of Abd.IV as in
<figureCitation id="13712A46FF9DFF906695FC25BE1E8184" box="[707,791,991,1017]" captionStart="FIGURE 19" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[243,1320,181,1919]" captionTargetId="figure-17@11.[243,1343,181,1919]" captionTargetPageId="11" captionText="FIGURE 19. Lepidocyrtus fuscocephalus sp. nov.: Abd.IV chaetotaxy (left side). Broad black circlesbroad ciliated macrochaetae, small black circlesthin ciliated macrochaetae." figureDoi="http://doi.org/10.5281/zenodo.6224652" httpUri="https://zenodo.org/record/6224652/files/figure.png" pageId="12" pageNumber="463">Fig. 19</figureCitation>
; macrochaetae
<emphasis id="B93EEAD1FF9DFF906793FC25BEE58184" bold="true" box="[965,1004,991,1017]" pageId="12" pageNumber="463">Sm</emphasis>
,
<emphasis id="B93EEAD1FF9DFF9067ACFC25BDBF8660" bold="true" pageId="12" pageNumber="463">B4, B5, B6, D3, E2, E3, E4, F1, F2, F3</emphasis>
broader and with broad socket; macrochaetae
<emphasis id="B93EEAD1FF9DFF90669FFBF9B9A38660" bold="true" box="[713,1194,1027,1053]" pageId="12" pageNumber="463">T6, T7, D2, De3, E1, E4p, Fe4, Fe5, F3p</emphasis>
shorter or longer but always thinner and with socket of minor diameter; macrochaeta
<emphasis id="B93EEAD1FF9DFF90672BFBDDBE95863C" bold="true" box="[893,924,1063,1089]" pageId="12" pageNumber="463">F2</emphasis>
inserted above macrochaeta
<emphasis id="B93EEAD1FF9DFF9060BDFBDDB802863C" bold="true" box="[1259,1291,1063,1089]" pageId="12" pageNumber="463">E3</emphasis>
; the ratio of distances between macrochaetae
<emphasis id="B93EEAD1FF9DFF906646FBB1BF6C8618" bold="true" box="[528,613,1099,1126]" pageId="12" pageNumber="463">SmB4</emphasis>
/
<emphasis id="B93EEAD1FF9DFF90663BFBB1BFB58618" bold="true" box="[621,700,1099,1126]" pageId="12" pageNumber="463">B4B6</emphasis>
is 0.80.9; the ratio of distances between macrochaetae
<emphasis id="B93EEAD1FF9DFF906113FBB1B89D8618" bold="true" box="[1349,1428,1099,1126]" pageId="12" pageNumber="463">B4B5</emphasis>
/
<emphasis id="B93EEAD1FF9DFF9064C1FB95BDEE86F4" bold="true" box="[151,231,1135,1162]" pageId="12" pageNumber="463">B5B6</emphasis>
is 1.11.3; accessory chaeta
<emphasis id="B93EEAD1FF9DFF90667CFB95BF3C86F4" bold="true" box="[554,565,1135,1161]" pageId="12" pageNumber="463">s</emphasis>
associated with trichobotrium
<emphasis id="B93EEAD1FF9DFF9067D8FB95BEA786F4" bold="true" box="[910,942,1135,1161]" pageId="12" pageNumber="463">T2</emphasis>
absent; chaetae
<emphasis id="B93EEAD1FF9DFF906033FB95B98E86F4" bold="true" box="[1125,1159,1135,1161]" pageId="12" pageNumber="463">D1</emphasis>
,
<emphasis id="B93EEAD1FF9DFF9060C3FB95B9A586F4" bold="true" box="[1173,1196,1135,1161]" pageId="12" pageNumber="463">m</emphasis>
,
<emphasis id="B93EEAD1FF9DFF9060ECFB95B9DF86F4" bold="true" box="[1210,1238,1135,1161]" pageId="12" pageNumber="463">pe</emphasis>
and
<emphasis id="B93EEAD1FF9DFF90615DFB95B82B86F4" bold="true" box="[1291,1314,1135,1161]" pageId="12" pageNumber="463">pi</emphasis>
associated with trichobotria
<emphasis id="B93EEAD1FF9DFF90650DFB69BC7286D0" bold="true" box="[347,379,1171,1197]" pageId="12" pageNumber="463">T2</emphasis>
and
<emphasis id="B93EEAD1FF9DFF9065E5FB69BCDD86D0" bold="true" box="[435,468,1171,1197]" pageId="12" pageNumber="463">T4</emphasis>
fan-shaped and strongly ciliate, chaeta
<emphasis id="B93EEAD1FF9DFF9067C5FB69BEA886D0" bold="true" box="[915,929,1171,1197]" pageId="12" pageNumber="463">a</emphasis>
pointed and strongly ciliate; sens chaetotaxy composed of 2 anterior dorsomedial elongate
<emphasis id="B93EEAD1FF9DFF9066F3FB4DBFBC86AC" bold="true" box="[677,693,1207,1233]" pageId="12" pageNumber="463">S</emphasis>
-chaetae, and short chaetae
<emphasis id="B93EEAD1FF9DFF9067A4FB4DB90286AC" bold="true" box="[1010,1035,1207,1233]" pageId="12" pageNumber="463">as</emphasis>
and
<emphasis id="B93EEAD1FF9DFF906010FB4DB96986AC" bold="true" box="[1094,1120,1207,1233]" pageId="12" pageNumber="463">ps</emphasis>
; posterior margin with 9+9 smooth mesochaetae; lateral region BP4 without pseudopori. Dorsal chaetotaxy of Abd.V with S-chaetae
<emphasis id="B93EEAD1FF9DFF90617DFB21B84D8688" bold="true" box="[1323,1348,1243,1269]" pageId="12" pageNumber="463">as</emphasis>
,
<emphasis id="B93EEAD1FF9DFF906107FB21B8958688" bold="true" box="[1361,1436,1243,1269]" pageId="12" pageNumber="463">acc.p4</emphasis>
and
<emphasis id="B93EEAD1FF9DFF906491FB05BC1B8764" bold="true" box="[199,274,1279,1305]" pageId="12" pageNumber="463">acc.p5</emphasis>
(
<figureCitation id="13712A46FF9DFF906577FB05BC7F8764" box="[289,374,1279,1305]" captionStart="FIGURES 2022" captionStartId="12.[151,264,633,657]" captionTargetBox="[151,1436,189,609]" captionTargetId="figure-34@12.[151,1436,181,609]" captionTargetPageId="12" captionText="FIGURES 2022. Lepidocyrtus fuscocephalus sp. nov.: 20, Abd.V chaetotaxy (left side), broad circleslong ciliated chaetae, small circlesshort ciliated or smooth chaetae; 21, trochanteral organ; 22, third leg unguis and unguiculus." figureDoi="http://doi.org/10.5281/zenodo.6224662" httpUri="https://zenodo.org/record/6224662/files/figure.png" pageId="12" pageNumber="463">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8BF536C3FF9DFF906491FADEBC618468" blockId="12.[151,1437,739,2026]" pageId="12" pageNumber="463">
Legs with scales except in claws. Rectangular-shaped trochanteral organ formed by a maximum of 28 smooth straight chaetae (
<figureCitation id="13712A46FF9DFF90650EFABDBCA4871C" box="[344,429,1351,1377]" captionStart="FIGURES 2022" captionStartId="12.[151,264,633,657]" captionTargetBox="[151,1436,189,609]" captionTargetId="figure-34@12.[151,1436,181,609]" captionTargetPageId="12" captionText="FIGURES 2022. Lepidocyrtus fuscocephalus sp. nov.: 20, Abd.V chaetotaxy (left side), broad circleslong ciliated chaetae, small circlesshort ciliated or smooth chaetae; 21, trochanteral organ; 22, third leg unguis and unguiculus." figureDoi="http://doi.org/10.5281/zenodo.6224662" httpUri="https://zenodo.org/record/6224662/files/figure.png" pageId="12" pageNumber="463">Fig. 21</figureCitation>
). Unguis (
<figureCitation id="13712A46FF9DFF906673FABDBF7E871C" box="[549,631,1351,1377]" captionStart="FIGURES 2022" captionStartId="12.[151,264,633,657]" captionTargetBox="[151,1436,189,609]" captionTargetId="figure-34@12.[151,1436,181,609]" captionTargetPageId="12" captionText="FIGURES 2022. Lepidocyrtus fuscocephalus sp. nov.: 20, Abd.V chaetotaxy (left side), broad circleslong ciliated chaetae, small circlesshort ciliated or smooth chaetae; 21, trochanteral organ; 22, third leg unguis and unguiculus." figureDoi="http://doi.org/10.5281/zenodo.6224662" httpUri="https://zenodo.org/record/6224662/files/figure.png" pageId="12" pageNumber="463">Fig. 22</figureCitation>
) with basal pair of teeth at 49% from base of the inner edge, and with two inner unpaired teeth at 71% and 88% from the base of the inner edge respectively; on unguis III the apical inner tooth tiny and sometimes difficult to see; one external tooth and a pair of lateral teeth also present. Unguiculus lanceolate with serrated outer margin (serration less evident on unguiculus III). Tibiotarsal tenent hair spatulate, smooth and as long as claw; ratio of supra-empodial chaeta (smooth chaeta on tibiotarsus III opposite to tenent hair) / unguiculus ≈ 1.2.
</paragraph>
<paragraph id="8BF536C3FF9DFF906491F9E5BF8484FF" blockId="12.[151,1437,739,2026]" pageId="12" pageNumber="463">Ventral tube with 6+6 ciliated chaetae on anterior side (4+4 proximal and 2+2 distal) and 11+11 weakly ciliated chaetae on posterior side; scales present on anterior and posterior sides; lateral flap with a maximum of 26 laterodistal chaetae (2023 ciliated and 3 smooth).</paragraph>
<paragraph id="8BF536C3FF9DFF906491F971B84F8490" blockId="12.[151,1437,739,2026]" pageId="12" pageNumber="463">Manubrium with scales on anterior and posterior surfaces, with 2+2 ciliated apical chaetae on anterior side. The ratio manubrium:dens:mucro is 19:20:1. Manubrial plate with 34 inner chaetae and a maximum of 16 outer chaetae. Dental tubercle absent. Mucro with the two teeth of the same size, without spinelet on basal spine.</paragraph>
<paragraph id="8BF536C3FF9DFF906491F90DB908856C" blockId="12.[151,1437,739,2026]" box="[199,1025,1783,1810]" pageId="12" pageNumber="463">
Pseudopores distribution on dorsal and ventral positions as in
<figureCitation id="13712A46FF9DFF906728F90DBEF4856C" box="[894,1021,1783,1809]" captionStart="FIGURE 23" captionStartId="13.[151,250,1892,1916]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-40@13.[151,1436,237,1869]" captionTargetPageId="13" captionText="FIGURE 23. Lepidocyrtus fuscocephalus sp. nov, L. milagrosae sp. nov. and L. semicoloratus sp. nov.: Pseudopores distribution: a, dorsal pseudopores; bc, ventral pesudopores." figureDoi="http://doi.org/10.5281/zenodo.6224666" httpUri="https://zenodo.org/record/6224666/files/figure.png" pageId="12" pageNumber="463">Figs 23ab</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3506548FF9DFF906491F8E1B827854B" box="[199,1326,1819,1846]" pageId="12" pageNumber="463" type="biology_ecology">
<paragraph id="8BF536C3FF9DFF906491F8E1B827854B" blockId="12.[151,1437,739,2026]" box="[199,1326,1819,1846]" pageId="12" pageNumber="463">
<emphasis id="B93EEAD1FF9DFF906491F8E1BCE68548" bold="true" box="[199,495,1819,1845]" pageId="12" pageNumber="463">Ecology and distribution</emphasis>
. All specimens were obtained by sifting soil surface litter under oak trees.
</paragraph>
</subSubSection>
<subSubSection id="C3506548FF9DFF916491F8C5BC3482A8" lastPageId="13" lastPageNumber="464" pageId="12" pageNumber="463" type="discussion">
<paragraph id="8BF536C3FF9DFF916491F8C5BC3482A8" blockId="12.[151,1437,739,2026]" lastBlockId="13.[151,1436,151,213]" lastPageId="13" lastPageNumber="464" pageId="12" pageNumber="463">
<emphasis id="B93EEAD1FF9DFF906491F8C5BC4C8524" bold="true" box="[199,325,1855,1881]" pageId="12" pageNumber="463">Discussion</emphasis>
. Morphological characters clearly assign
<taxonomicName id="4C4A4D40FF9DFF906748F8C5B9518524" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[798,1112,1855,1881]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF9DFF906748F8C5B9518524" box="[798,1112,1855,1881]" italics="true" pageId="12" pageNumber="463">Lepidocyrtus fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF9DFF906009F8C5B9B38524" bold="true" box="[1119,1210,1855,1881]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A20D57AAFF9DFF906009F8C5B9B38524" box="[1119,1210,1855,1881]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
to the
<taxonomicName id="4C4A4D40FF9DFF90615CF8C5BDF78500" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="lignorum">
<emphasis id="B93EEAD1FF9DFF90615CF8C5BDF78500" italics="true" pageId="12" pageNumber="463">Lepidocyrtus lignorum</emphasis>
</taxonomicName>
-group (
<emphasis id="B93EEAD1FF9DFF90650CF89EBC918500" box="[346,408,1892,1917]" italics="true" pageId="12" pageNumber="463">sensu</emphasis>
<bibRefCitation id="EFDB4B32FF9DFF9065F7F89EBF318500" author="Mateos, E." box="[417,568,1891,1918]" pageId="12" pageNumber="463" pagination="25 - 40" refId="ref12259" refString="Mateos, E. (2011) New Lepidocyrtus Bourlet, 1839 taxa from Greece (Collembola: Entomobryidae). Zootaxa, 3108 (1), 25 - 40. https: // doi. org / 10.11646 / zootaxa. 3108.1.2" type="journal article" year="2011">Mateos 2011</bibRefCitation>
). By the characteristic body colour pattern
<taxonomicName id="4C4A4D40FF9DFF906070F89EB9ED8500" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[1062,1252,1891,1917]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF9DFF906070F89EB9ED8500" box="[1062,1252,1891,1917]" italics="true" pageId="12" pageNumber="463">L. fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF9DFF9060BAF899B84E8500" bold="true" box="[1260,1351,1891,1917]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A20D57AAFF9DFF9060BAF899B84E8500" box="[1260,1351,1891,1917]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
clearly differs from all the other species of the
<taxonomicName id="4C4A4D40FF9DFF906609F872BFEF85DC" box="[607,742,1927,1953]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="lignorum">
<emphasis id="B93EEAD1FF9DFF906609F872BFEF85DC" box="[607,742,1927,1953]" italics="true" pageId="12" pageNumber="463">L. lignorum</emphasis>
</taxonomicName>
-group. Other differences between all species included in the group are summarized in
<tableCitation id="C6C80378FF9DFF9065E4F851BF0E85B8" box="[434,519,1963,1989]" captionStart="TABLE 2" captionStartId="14.[151,239,476,502]" captionTargetPageId="14" captionText="TABLE 2. Characters differentiating between the European Lepidocyrtus species of lignorum-group." httpUri="http://table.plazi.org/id/DF35664BFF9FFF9264C1FE26B9C6838B" pageId="12" pageNumber="463" tableUuid="DF35664BFF9FFF9264C1FE26B9C6838B">Table 2</tableCitation>
. By having body partially blue pigmented and lanceolate unguiculus
<taxonomicName id="4C4A4D40FF9DFF906150F856BDCF8594" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="fuscocephalus" status="sp. nov.">
<emphasis id="B93EEAD1FF9DFF906150F856BDCF8594" italics="true" pageId="12" pageNumber="463">L. fuscocephalus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF9DFF906498F835BC208594" bold="true" box="[206,297,1999,2025]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A20D57AAFF9DFF906498F835BC208594" box="[206,297,1999,2025]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is close to species
<taxonomicName id="4C4A4D40FF9DFF906652F82ABF8C8594" authorityName="Handschin" authorityYear="1924" box="[516,645,2000,2025]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="instratus">
<emphasis id="B93EEAD1FF9DFF906652F82ABF8C8594" box="[516,645,2000,2025]" italics="true" pageId="12" pageNumber="463">L. instratus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D40FF9DFF9066C2F82ABE288594" authorityName="Baquero &amp; Jordana" authorityYear="2021" box="[660,801,1999,2025]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="labyrinthi">
<emphasis id="B93EEAD1FF9DFF9066C2F82ABE288594" box="[660,801,1999,2025]" italics="true" pageId="12" pageNumber="463">L. labyrinthi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D40FF9DFF906767F82ABE938594" box="[817,922,2000,2025]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="traseri">
<emphasis id="B93EEAD1FF9DFF906767F82ABE938594" box="[817,922,2000,2025]" italics="true" pageId="12" pageNumber="463">L. traseri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D40FF9DFF9067FFF82AB94C8594" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[937,1093,1999,2025]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="milagrosae" status="sp. nov.">
<emphasis id="B93EEAD1FF9DFF9067FFF82AB94C8594" box="[937,1093,1999,2025]" italics="true" pageId="12" pageNumber="463">L. milagrosae</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF9DFF90601BF835B9A18594" bold="true" box="[1101,1192,1999,2025]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A20D57AAFF9DFF90601BF835B9A18594" box="[1101,1192,1999,2025]" pageId="12" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C4A4D40FF9DFF9060B6F82AB8958594" authority="Mateos &amp; Álvarez-Presas, 2022" authorityName="Mateos &amp; Álvarez-Presas" authorityYear="2022" box="[1248,1436,1999,2025]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="semicoloratus" status="sp. nov.">
<emphasis id="B93EEAD1FF9DFF9060B6F82AB8958594" box="[1248,1436,1999,2025]" italics="true" pageId="12" pageNumber="463">L. semicoloratus</emphasis>
</taxonomicName>
<emphasis id="B93EEAD1FF9CFF9164C1FF6DBDF882CC" bold="true" box="[151,241,151,177]" pageId="13" pageNumber="464">
<taxonomicNameLabel id="A20D57AAFF9CFF9164C1FF6DBDF882CC" box="[151,241,151,177]" pageId="13" pageNumber="464" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Of these, the new species differ by the colour pattern, the presence of scales on Ant.III and pointed labral apical chaetae.
</paragraph>
</subSubSection>
</treatment>
</document>