226 lines
34 KiB
XML
226 lines
34 KiB
XML
<document id="95EF19013B58A3916D129F3B8CAE9B29" ID-CLB-Dataset="110330" ID-DOI="10.17109/AZH.65.1.1.2019" ID-GBIF-Dataset="dedc13e2-6231-4a23-bf04-b37b23379c97" ID-ISSN="2064-2474" ID-Zenodo-Dep="5734674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638188679242" checkinUser="felipe" docAuthor="Kóbor, Péter" docDate="2019" docId="03E387D2FFE0FFBB48DECF84FC58C27E" docLanguage="en" docName="ActaZoolAcadSciH.65.1.1-8.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 65 (1)" docSource="http://dx.doi.org/10.17109/azh.65.1.1.2019" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.4:ActaZoolAcadSciH.2015-.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-.journal_article" docStyleVersion="4" docTitle="Umbrageocoris Kóbor 2019, gen. n." docType="treatment" docUuid="EB1CD010-9CA6-45E9-9C79-7EEB330C8597" docUuidSource="ZooBank" docVersion="7" lastPageNumber="4" masterDocId="FFDAFFAAFFE2FFBE490ECE12FFEEC012" masterDocTitle="Umbrageocoris Kondorosyi: A New Genus And Species Of Big-Eyed Bugs From New Guinea (Heteroptera: Lygaeoidea: Geocoridae)" masterLastPageNumber="8" masterPageNumber="1" pageNumber="3" updateTime="1699470944118" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="5852CF5B6E7317197DE6AD5F34E6EE1B">Umbrageocoris Kondorosyi: A New Genus And Species Of Big-Eyed Bugs From New Guinea (Heteroptera: Lygaeoidea: Geocoridae)</mods:title>
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<mods:namePart id="A8D86E7BA22C46A481ED3F1C3B37FDFD">Kóbor, Péter</mods:namePart>
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<mods:title id="84B0F36A8B49A4E9E60FC20E46942CB2">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date id="A812013A1335C2D7C232F538E6EBEDBB">2019</mods:date>
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<treatment id="03E387D2FFE0FFBB48DECF84FC58C27E" ID-DOI="http://doi.org/10.5281/zenodo.6949280" ID-GBIF-Taxon="198138328" ID-Zenodo-Dep="6949280" LSID="urn:lsid:zoobank.org:act:EB1CD010-9CA6-45E9-9C79-7EEB330C8597" httpUri="http://treatment.plazi.org/id/03E387D2FFE0FFBB48DECF84FC58C27E" lastPageId="5" lastPageNumber="4" pageId="2" pageNumber="3">
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<subSubSection id="C350654FFFE0FFBC48DECF84FD17C1A3" box="[464,761,406,433]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BF536C4FFE0FFBC48DECF84FD17C1A3" blockId="2.[464,761,406,433]" box="[464,761,406,433]" pageId="2" pageNumber="3">
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<heading id="D0BD81A8FFE0FFBC48DECF84FD17C1A3" box="[464,761,406,433]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="4C4A4D47FFE0FFBC48DECF84FD72C1A2" ID-CoL="8QNXC" authority="Kóbor, 2019" authorityName="Kóbor" authorityYear="2019" box="[464,668,406,432]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B93EEAD6FFE0FFBC48DECF84FD72C1A2" bold="true" box="[464,668,406,432]" pageId="2" pageNumber="3">Umbrageocoris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20D57ADFFE0FFBC4BADCF85FD17C1A3" box="[675,761,407,433]" pageId="2" pageNumber="3" rank="genus">gen. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C350654FFFE0FFBC4B24CFABFD71C1C6" box="[554,671,441,468]" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BF536C4FFE0FFBC4B24CFABFD71C1C6" blockId="2.[554,671,441,468]" box="[554,671,441,468]" pageId="2" pageNumber="3">
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<heading id="D0BD81A8FFE0FFBC4B24CFABFD71C1C6" box="[554,671,441,468]" centered="true" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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(
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<figureCitation id="13712A41FFE0FFBC4B3DCFABFD78C1C6" box="[563,662,441,468]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="3.[174,210,1524,1546]" captionStartId-1="4.[136,172,952,974]" captionStartId-2="4.[205,241,1516,1538]" captionStartId-3="5.[174,210,1496,1518]" captionTargetBox-0="[174,1126,1236,1504]" captionTargetBox-1="[136,1088,189,928]" captionTargetBox-2="[141,1081,1100,1488]" captionTargetBox-3="[178,1126,1203,1466]" captionTargetId-0="figure-346@3.[174,1126,1236,1504]" captionTargetId-1="figure-23@4.[136,1088,189,929]" captionTargetId-2="figure-59@4.[136,1088,1088,1493]" captionTargetId-3="figure-400@5.[174,1127,1203,1468]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Fig. 1. Habitus of Umbrageocoris kondorosyi sp. n.: A = dorsal (holotype, HNHM), B = lateral (paratype, BMNH)" captionText-1="Fig. 2. Photos of Umbrageocoris kondorosyi sp. n.: A = dorsum of head, B = venter of head, C = peritreme and evaporative area of metathoracic scent gland, D = dorsolateral trichobotria of abdomen" captionText-2="Fig. 3. Male paramere of Umbrageocoris kondorosyi sp. n. from different angles" captionText-3="Fig. 4. Peritreme of metathoracic scent gland in New Guinean Geocorinae: A = Germalus Stål, 1862; B = Umbrageocoris gen. n.; C = Geocoris Fallén, 1814. Legend: cx2 = mesocoxa, ev = evaporatorium, pe = peritreme, o = ostiole, ves = vestibular scar" figureDoi-0="http://doi.org/10.5281/zenodo.5734676" figureDoi-1="http://doi.org/10.5281/zenodo.5734678" figureDoi-2="http://doi.org/10.5281/zenodo.5734682" figureDoi-3="http://doi.org/10.5281/zenodo.5734684" httpUri-0="https://zenodo.org/record/5734676/files/figure.png" httpUri-1="https://zenodo.org/record/5734678/files/figure.png" httpUri-2="https://zenodo.org/record/5734682/files/figure.png" httpUri-3="https://zenodo.org/record/5734684/files/figure.png" pageId="2" pageNumber="3">Figs1–4</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C350654FFFE0FFBC4986CFE0FDB2C215" box="[136,604,497,519]" pageId="2" pageNumber="3" type="type_taxon">
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<paragraph id="8BF536C4FFE0FFBC4986CFE0FDB2C215" blockId="2.[136,604,497,519]" box="[136,604,497,519]" pageId="2" pageNumber="3">
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<typeStatus id="54F18866FFE0FFBC4986CFE0FF50C215" box="[136,190,498,519]" pageId="2" pageNumber="3">Type</typeStatus>
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species:
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<taxonomicName id="4C4A4D47FFE0FFBC4815CFE3FDF3C214" ID-CoL="8QNHD" authority="Kóbor, 2019" authorityName="Kóbor" authorityYear="2019" box="[283,541,497,518]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kondorosyi" status="sp. nov.">
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<emphasis id="B93EEAD6FFE0FFBC4815CFE3FDF3C214" box="[283,541,497,518]" italics="true" pageId="2" pageNumber="3">Umbrageocoris kondorosyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20D57ADFFE0FFBC4B2DCFE0FDB2C215" box="[547,604,498,519]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C350654FFFE0FFBC49CBCC39FD8DC558" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BF536C4FFE0FFBC49CBCC39FD8DC558" blockId="2.[136,1089,555,1469]" pageId="2" pageNumber="3">
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Description – Body elongate oval; integument shiny, lacking conspicuous pubescence (
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<figureCitation id="13712A41FFE0FFBC49DCCC55FECFC24C" box="[210,289,583,606]" captionStart="Fig" captionStartId="3.[174,210,1524,1546]" captionTargetBox="[174,1126,1236,1504]" captionTargetId="figure-346@3.[174,1126,1236,1504]" captionTargetPageId="3" captionText="Fig. 1. Habitus of Umbrageocoris kondorosyi sp. n.: A = dorsal (holotype, HNHM), B = lateral (paratype, BMNH)" figureDoi="http://doi.org/10.5281/zenodo.5734676" httpUri="https://zenodo.org/record/5734676/files/figure.png" pageId="2" pageNumber="3">Fig. 1A</figureCitation>
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). Head pentagonal; surface smooth, shiny; vertex with a thin longitudinal furrow extending from between the ocelli to middle of clypeus. Eyes big, reniform, moderately stylate; ocular sulcus slightly visible but complete (
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<figureCitation id="13712A41FFE0FFBC4BFCCC93FCD1C285" box="[754,831,641,663]" captionStart="Fig" captionStartId="4.[136,172,952,974]" captionTargetBox="[136,1088,189,928]" captionTargetId="figure-23@4.[136,1088,189,929]" captionTargetPageId="4" captionText="Fig. 2. Photos of Umbrageocoris kondorosyi sp. n.: A = dorsum of head, B = venter of head, C = peritreme and evaporative area of metathoracic scent gland, D = dorsolateral trichobotria of abdomen" figureDoi="http://doi.org/10.5281/zenodo.5734678" httpUri="https://zenodo.org/record/5734678/files/figure.png" pageId="2" pageNumber="3">Fig. 2B</figureCitation>
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). Proportion of antennal segments: segment I shortest, segment II longest, segments III and IV subequal in length. Margins of jugae somewhat projecting, forming a Ushaped rim; labial trough closes at midline of head, suture formed by margins extending to base of head. Labial segment I not reaching base of head, segment II shortest, segment III subequal to I, segment IV longest, apex reaching about the line of metacoxae. Thorax. Pronotum trapezoidal with anterior margin and edges broadly convex. Surface shiny with coarse, dense punctation except anterior and posterior margins, callosities and humeral angles. Margins, callosities and humeral angles slightly elevated. Anterior half of lateral margins of pronotum slightly carinate and impressed behind callosities. Scutellum almost equilateral, sharply pointed apically, surface with dense punctuation except trifurcate median carina. Apical part of carina expressed, elevated, basal parts reduced but visible. Hemelytron macropterous with welldeveloped membrane, length of membrane subequal to length of corium. Margins of clavus converging to apex; claval commissure reduced, inconspicuous. Costal margin suddenly widening at third of length; costal region widest at about half of length. Thoracic pleurites and sternites with dense, strong punctation except supracoxal lobes, peritreme of metathoracic scent gland and dorsal edge of metapleurite (
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<figureCitation id="13712A41FFE0FFBC4A75CA5CFC24C476" box="[891,970,1102,1124]" captionStart="Fig" captionStartId="4.[136,172,952,974]" captionTargetBox="[136,1088,189,928]" captionTargetId="figure-23@4.[136,1088,189,929]" captionTargetPageId="4" captionText="Fig. 2. Photos of Umbrageocoris kondorosyi sp. n.: A = dorsum of head, B = venter of head, C = peritreme and evaporative area of metathoracic scent gland, D = dorsolateral trichobotria of abdomen" figureDoi="http://doi.org/10.5281/zenodo.5734678" httpUri="https://zenodo.org/record/5734678/files/figure.png" pageId="2" pageNumber="3">Fig. 2A</figureCitation>
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). Peritreme of metathoracic scent gland situated ventrolaterally, near anterior margin of metathorax; shape earlike, dorsal rim slightly curved downwards. Evaporative area small, narrow, not reaching dorsal margin of supracoxal lobes (
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<figureCitation id="13712A41FFE0FFBC4B52CAB6FD45C4AB" box="[604,683,1188,1210]" captionStart="Fig" captionStartId="4.[136,172,952,974]" captionTargetBox="[136,1088,189,928]" captionTargetId="figure-23@4.[136,1088,189,929]" captionTargetPageId="4" captionText="Fig. 2. Photos of Umbrageocoris kondorosyi sp. n.: A = dorsum of head, B = venter of head, C = peritreme and evaporative area of metathoracic scent gland, D = dorsolateral trichobotria of abdomen" figureDoi="http://doi.org/10.5281/zenodo.5734678" httpUri="https://zenodo.org/record/5734678/files/figure.png" pageId="2" pageNumber="3">Figs 2B</figureCitation>
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,
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<figureCitation id="13712A41FFE0FFBC4BBBCAB6FD3DC4A8" box="[693,723,1188,1210]" captionStart="Fig" captionStartId="5.[174,210,1496,1518]" captionTargetBox="[178,1126,1203,1466]" captionTargetId="figure-400@5.[174,1127,1203,1468]" captionTargetPageId="5" captionText="Fig. 4. Peritreme of metathoracic scent gland in New Guinean Geocorinae: A = Germalus Stål, 1862; B = Umbrageocoris gen. n.; C = Geocoris Fallén, 1814. Legend: cx2 = mesocoxa, ev = evaporatorium, pe = peritreme, o = ostiole, ves = vestibular scar" figureDoi="http://doi.org/10.5281/zenodo.5734684" httpUri="https://zenodo.org/record/5734684/files/figure.png" pageId="2" pageNumber="3">4B</figureCitation>
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). Abdomen shiny, with sparse fine pubescence. Abdominal tergites III–V with surface strongly wrinkled (
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<figureCitation id="13712A41FFE0FFBC4A79CAD3FC29C4C5" box="[887,967,1217,1239]" captionStart="Fig" captionStartId="4.[136,172,952,974]" captionTargetBox="[136,1088,189,928]" captionTargetId="figure-23@4.[136,1088,189,929]" captionTargetPageId="4" captionText="Fig. 2. Photos of Umbrageocoris kondorosyi sp. n.: A = dorsum of head, B = venter of head, C = peritreme and evaporative area of metathoracic scent gland, D = dorsolateral trichobotria of abdomen" figureDoi="http://doi.org/10.5281/zenodo.5734678" httpUri="https://zenodo.org/record/5734678/files/figure.png" pageId="2" pageNumber="3">Fig. 2D</figureCitation>
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). Trichobotria on connexiva V–VI conspicuously bulging (
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<figureCitation id="13712A41FFE0FFBC4B8DCACCFD3CC4E6" box="[643,722,1246,1268]" captionStart="Fig" captionStartId="4.[136,172,952,974]" captionTargetBox="[136,1088,189,928]" captionTargetId="figure-23@4.[136,1088,189,929]" captionTargetPageId="4" captionText="Fig. 2. Photos of Umbrageocoris kondorosyi sp. n.: A = dorsum of head, B = venter of head, C = peritreme and evaporative area of metathoracic scent gland, D = dorsolateral trichobotria of abdomen" figureDoi="http://doi.org/10.5281/zenodo.5734678" httpUri="https://zenodo.org/record/5734678/files/figure.png" pageId="2" pageNumber="3">Fig. 2C</figureCitation>
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). Abdominal venter finely rugous. Genitalia: Male pygophore with conspicuous lateral processes on dorsal half of the opening; paramere (
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<figureCitation id="13712A41FFE0FFBC4826CB05FE8CC53C" box="[296,354,1303,1326]" captionStart="Fig" captionStartId="4.[205,241,1516,1538]" captionTargetBox="[141,1081,1100,1488]" captionTargetId="figure-59@4.[136,1088,1088,1493]" captionTargetPageId="4" captionText="Fig. 3. Male paramere of Umbrageocoris kondorosyi sp. n. from different angles" figureDoi="http://doi.org/10.5281/zenodo.5734682" httpUri="https://zenodo.org/record/5734682/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
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) with blade elongate, somewhat curved, lacking conspicuous setae or pubescence; gonoporal process with 10 coils.
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</paragraph>
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</subSubSection>
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<subSubSection id="C350654FFFE0FFBC49CBCB43FF15C5AF" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="8BF536C4FFE0FFBC49CBCB43FF15C5AF" blockId="2.[136,1089,555,1469]" pageId="2" pageNumber="3">
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Etymology – The name of the is composed of the Latin noun umbra (= shade, night, darkness), referring to the dorsum being dark brownish laterally margined with ochraceous, endowing it with a “shaded look”, and
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<taxonomicName id="4C4A4D47FFE0FFBC4B87CB99FD32C5B2" authorityName="Fallen" authorityYear="1814" box="[649,732,1419,1440]" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B93EEAD6FFE0FFBC4B87CB99FD32C5B2" box="[649,732,1419,1440]" italics="true" pageId="2" pageNumber="3">Geocoris</emphasis>
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</taxonomicName>
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, a closely related genus. Gender masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C350654FFFE0FFBD49CBCBF0FE4BC16D" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="8BF536C4FFE0FFBD49CBCBF0FE4BC16D" blockId="2.[136,1089,1506,1566]" lastBlockId="3.[173,1128,189,1190]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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Diagnosis –
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<taxonomicName id="4C4A4D47FFE0FFBC486DCBF0FDE0C5EE" authority="Kóbor, 2019" authorityName="Kóbor" authorityYear="2019" box="[355,526,1506,1532]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B93EEAD6FFE0FFBC486DCBF0FDE0C5EE" box="[355,526,1506,1532]" italics="true" pageId="2" pageNumber="3">Umbrageocoris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20D57ADFFE0FFBC4B17CBF1FD81C5EF" box="[537,623,1507,1533]" pageId="2" pageNumber="3" rank="genus">gen. n.</taxonomicNameLabel>
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can be separated from other geocorines of the region with combination of following characters: labial through Yshaped with suture reaching base of head; labial segment II shorter than III, segment IV longest; clavus gradually narrowing towards apex, claval commissure reduced; corium punctate along veins; peritreme and evaporatorium of metathoracic scent gland as in
|
||
<figureCitation id="13712A41FFE1FFBD4B5DCF30FD42C12E" box="[595,684,290,316]" captionStart="Fig" captionStartId="5.[174,210,1496,1518]" captionTargetBox="[178,1126,1203,1466]" captionTargetId="figure-400@5.[174,1127,1203,1468]" captionTargetPageId="5" captionText="Fig. 4. Peritreme of metathoracic scent gland in New Guinean Geocorinae: A = Germalus Stål, 1862; B = Umbrageocoris gen. n.; C = Geocoris Fallén, 1814. Legend: cx2 = mesocoxa, ev = evaporatorium, pe = peritreme, o = ostiole, ves = vestibular scar" figureDoi="http://doi.org/10.5281/zenodo.5734684" httpUri="https://zenodo.org/record/5734684/files/figure.png" pageId="3" pageNumber="4">Fig. 4B</figureCitation>
|
||
; surface of abdominal terga III and IV rugous medially; male paramere slender, blade narrow, curved, surface lacking pubescence.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C350654FFFE1FFBB49E4CF95FC58C27E" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="8BF536C4FFE1FFBD49E4CF95FCF5C471" blockId="3.[173,1128,189,1190]" pageId="3" pageNumber="4">
|
||
Remarks –
|
||
<taxonomicName id="4C4A4D47FFE1FFBD487FCF95FDF2C1B3" authority="Kóbor, 2019" authorityName="Kóbor" authorityYear="2019" box="[369,540,391,417]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93EEAD6FFE1FFBD487FCF95FDF2C1B3" box="[369,540,391,417]" italics="true" pageId="3" pageNumber="4">Umbrageocoris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20D57ADFFE1FFBD4B2CCF9AFD9CC1B0" box="[546,626,392,418]" pageId="3" pageNumber="4" rank="genus">gen. n.</taxonomicNameLabel>
|
||
resembling most to the Australian genus
|
||
<taxonomicName id="4C4A4D47FFE1FFBD49A3CFBBFDCDC1D1" authority="Montandon, 1907" authorityName="Montandon" authorityYear="1907" box="[173,547,425,451]" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD49A3CFBBFEACC1D1" box="[173,322,425,451]" italics="true" pageId="3" pageNumber="4">Stylogeocoris</emphasis>
|
||
Montandon, 1907
|
||
</taxonomicName>
|
||
, nevertheless there are some remarkable differences separating these genera. Based on
|
||
<bibRefCitation id="EFDB4B35FFE1FFBD4BAACFD8FC64C1F6" author="MALIPATIL, M. B." box="[676,906,458,484]" pageId="3" pageNumber="4" pagination="299 - 327" refId="ref3554" refString="MALIPATIL, M. B. (1994): Revision of Australian Geocoris Fallen and Stylogeocoris Montandon (Heteroptera: Lygaeidae: Geocorinae). - Invertebrate Systematics 8 (2): 299 - 327. https: // doi. org / 10.1071 / IT 9940299" type="journal article" year="1994">MALIPATIL’s (1994)</bibRefCitation>
|
||
revisionary work on
|
||
<taxonomicName id="4C4A4D47FFE1FFBD49D9CFFEFE82C214" authorityName="Montandon" authorityYear="1907" box="[215,364,492,518]" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD49D9CFFEFE82C214" box="[215,364,492,518]" italics="true" pageId="3" pageNumber="4">Stylogeocoris</emphasis>
|
||
</taxonomicName>
|
||
and studying the
|
||
<typeStatus id="54F18866FFE1FFBD4B59CFFEFD2BC214" box="[599,709,492,518]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
of
|
||
<emphasis id="B93EEAD6FFE1FFBD4BFFCFFEFC2AC214" box="[753,964,492,518]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C4A4D47FFE1FFBD4BFFCFFEFC68C214" authorityName="Montandon" authorityYear="1907" box="[753,902,492,518]" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Stylogeocoris</taxonomicName>
|
||
biroi
|
||
</emphasis>
|
||
Montandon, 1907 (HNHM), this genus can be diagnosed with the combination of following characters: labial segment 2 and 3 subequal in length; ocular sulcus complete, distinct; margins of clavus almost parallel, claval commissure developed; abdominal terga III and IV punctate. In
|
||
<taxonomicName id="4C4A4D47FFE1FFBD4B86CC60FCDDC29E" authorityName="Kóbor" authorityYear="2019" box="[648,819,626,652]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD4B86CC60FCDDC29E" box="[648,819,626,652]" italics="true" pageId="3" pageNumber="4">Umbrageocoris</emphasis>
|
||
</taxonomicName>
|
||
labial segment II is conspicuously shorter than segment III; margins of clavus gradually converging, claval commissure reduced; abdominal terga III and IV rather wrinkly than punctate. Male paramere is stouter, more curved in
|
||
<taxonomicName id="4C4A4D47FFE1FFBD4A20CCC5FC2DC2E3" authorityName="Montandon" authorityYear="1907" box="[814,963,727,753]" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD4A20CCC5FC2DC2E3" box="[814,963,727,753]" italics="true" pageId="3" pageNumber="4">Stylogeocoris</emphasis>
|
||
</taxonomicName>
|
||
compared to
|
||
<taxonomicName id="4C4A4D47FFE1FFBD49A0CCEBFEB7C301" authorityName="Kóbor" authorityYear="2019" box="[174,345,761,787]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD49A0CCEBFEB7C301" box="[174,345,761,787]" italics="true" pageId="3" pageNumber="4">Umbrageocoris</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, paramere of
|
||
<taxonomicName id="4C4A4D47FFE1FFBD4BA6CCEBFCBDC301" authorityName="Kóbor" authorityYear="2019" box="[680,851,761,787]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD4BA6CCEBFCBDC301" box="[680,851,761,787]" italics="true" pageId="3" pageNumber="4">Umbrageocoris</emphasis>
|
||
</taxonomicName>
|
||
is lacking pubescence. Comparing to the
|
||
<taxonomicName id="4C4A4D47FFE1FFBD4882CD09FD6AC326" authority="Fallen, 1814" authorityName="Fallen" authorityYear="1814" box="[396,644,794,821]" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD4882CD09FE03C327" box="[396,493,795,821]" italics="true" pageId="3" pageNumber="4">Geocoris</emphasis>
|
||
Fallén, 1814
|
||
</taxonomicName>
|
||
distributed in the archipelago (
|
||
<emphasis id="B93EEAD6FFE1FFBD4D0BCD09FF14C344" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C4A4D47FFE1FFBD4D0BCD09FB88C327" authorityName="Fallen" authorityYear="1814" box="[1029,1126,795,821]" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Geocoris</taxonomicName>
|
||
willeyi
|
||
</emphasis>
|
||
Kirkaldy, 1905 and
|
||
<emphasis id="B93EEAD6FFE1FFBD48F9CD2EFD9CC344" box="[503,626,828,854]" italics="true" pageId="3" pageNumber="4">G. leopoldi</emphasis>
|
||
Schouteden, 1933;
|
||
<typeStatus id="54F18866FFE1FFBD4A6DCD2EFC33C344" box="[867,989,828,854]" pageId="3" pageNumber="4" type="holotype">holotypes</typeStatus>
|
||
examined) the following differences can be observed: the new genus is more slender in general appearance than the other two species; head more pentagonal, ocular sulcus slight, but complete, not reduced; pronotal callosities more developed in
|
||
<taxonomicName id="4C4A4D47FFE1FFBD49DECDD0FE95C3CE" authorityName="Kóbor" authorityYear="2019" box="[208,379,962,988]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD49DECDD0FE95C3CE" box="[208,379,962,988]" italics="true" pageId="3" pageNumber="4">Umbrageocoris</emphasis>
|
||
</taxonomicName>
|
||
; apical part of trifurcate carina of scutellum slight, but not reduced; corium of hemelytron without punctation in intervanals. In terms of
|
||
<taxonomicName id="4C4A4D47FFE1FFBD49A0CA17FEF2C40D" authorityName="Stal" authorityYear="1862" box="[174,284,1029,1055]" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD49A0CA17FEF2C40D" box="[174,284,1029,1055]" italics="true" pageId="3" pageNumber="4">Germalus</emphasis>
|
||
</taxonomicName>
|
||
, the most remarkable differences are the proportions of labial segment II–III (similarly as in the case of
|
||
<taxonomicName id="4C4A4D47FFE1FFBD4B9ACA35FCC7C453" authorityName="Montandon" authorityYear="1907" box="[660,809,1063,1089]" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD4B9ACA35FCC7C453" box="[660,809,1063,1089]" italics="true" pageId="3" pageNumber="4">Stylogeocoris</emphasis>
|
||
</taxonomicName>
|
||
) and the arrangement of peritreme of metathoracic scent gland (
|
||
<figureCitation id="13712A41FFE1FFBD4B9CCA5BFCE3C471" box="[658,781,1097,1123]" captionStart="Fig" captionStartId="5.[174,210,1496,1518]" captionTargetBox="[178,1126,1203,1466]" captionTargetId="figure-400@5.[174,1127,1203,1468]" captionTargetPageId="5" captionText="Fig. 4. Peritreme of metathoracic scent gland in New Guinean Geocorinae: A = Germalus Stål, 1862; B = Umbrageocoris gen. n.; C = Geocoris Fallén, 1814. Legend: cx2 = mesocoxa, ev = evaporatorium, pe = peritreme, o = ostiole, ves = vestibular scar" figureDoi="http://doi.org/10.5281/zenodo.5734684" httpUri="https://zenodo.org/record/5734684/files/figure.png" pageId="3" pageNumber="4">Fig. 4A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF536C4FFE1FFBB49E4CA78FC58C27E" blockId="3.[173,1128,189,1190]" lastBlockId="5.[174,1128,189,620]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
|
||
If looking at genus
|
||
<taxonomicName id="4C4A4D47FFE1FFBD48ECCA79FDADC497" authorityName="Fallen" authorityYear="1814" box="[482,579,1131,1157]" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD48ECCA79FDADC497" box="[482,579,1131,1157]" italics="true" pageId="3" pageNumber="4">Geocoris</emphasis>
|
||
</taxonomicName>
|
||
in general, species similar to
|
||
<taxonomicName id="4C4A4D47FFE1FFBD4AB2CA78FB89C496" authorityName="Kóbor" authorityYear="2019" box="[956,1127,1130,1156]" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE1FFBD4AB2CA78FB89C496" box="[956,1127,1130,1156]" italics="true" pageId="3" pageNumber="4">Umbrageocoris</emphasis>
|
||
</taxonomicName>
|
||
can be found in other biogeographical regions. Nevertheless, it has to be noted that more authors consider the genus
|
||
<taxonomicName id="4C4A4D47FFE7FFBB4B72CEACFD33C0CA" authorityName="Fallen" authorityYear="1814" box="[636,733,190,216]" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE7FFBB4B72CEACFD33C0CA" box="[636,733,190,216]" italics="true" pageId="5" pageNumber="6">Geocoris</emphasis>
|
||
</taxonomicName>
|
||
“an illdefined group of species belonging to perhaps several distinct genera” (
|
||
<bibRefCitation id="EFDB4B35FFE7FFBB4BE3CECDFC1BC0EB" author="READIO, J. & SWEET, M. H." box="[749,1013,223,249]" pageId="5" pageNumber="6" refId="ref3756" refString="READIO, J. & SWEET, M. H. (1982): A review of the Geocorinae of the United States east of 100 th meridian (Hemiptera: Lygaeidae). - Entomological Society of America Miscellaneous Publications 12. 92 pp." type="book" year="1982">READIO & SWEET 1982</bibRefCitation>
|
||
, MALIPATIL 1994) which is in need of a thorough revision. Based on the literature available and own observation, the author of recent study supports this opinion and concludes that the given combination of characters is eligible to separate this taxon on generic level from other members of New Guinean fauna. One of the most convincing characters is the arrangement and shape of the outer structures of metathoracic scent efferent apparatus (
|
||
<figureCitation id="13712A41FFE7FFBB4A3CCFBBFC96C1D1" box="[818,888,425,451]" captionStart="Fig" captionStartId="5.[174,210,1496,1518]" captionTargetBox="[178,1126,1203,1466]" captionTargetId="figure-400@5.[174,1127,1203,1468]" captionTargetPageId="5" captionText="Fig. 4. Peritreme of metathoracic scent gland in New Guinean Geocorinae: A = Germalus Stål, 1862; B = Umbrageocoris gen. n.; C = Geocoris Fallén, 1814. Legend: cx2 = mesocoxa, ev = evaporatorium, pe = peritreme, o = ostiole, ves = vestibular scar" figureDoi="http://doi.org/10.5281/zenodo.5734684" httpUri="https://zenodo.org/record/5734684/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
|
||
), which is showing remarkable differences if compared to
|
||
<taxonomicName id="4C4A4D47FFE7FFBB4B93CFD9FD10C1F7" authorityName="Fallen" authorityYear="1814" box="[669,766,459,485]" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93EEAD6FFE7FFBB4B93CFD9FD10C1F7" box="[669,766,459,485]" italics="true" pageId="5" pageNumber="6">Geocoris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4A4D47FFE7FFBB4A4CCFD8FBFCC1F6" box="[834,1042,458,484]" class="Insecta" family="Geocoridae" genus="Germalus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B93EEAD6FFE7FFBB4A4CCFD8FC5EC1F6" box="[834,944,458,484]" italics="true" pageId="5" pageNumber="6">Germalus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of the region. Based on the results of recent study and unpublished findings of the author this character might have significant value in taxonomy and phylogeny of
|
||
<taxonomicName id="4C4A4D47FFE7FFBB480CCC3DFE7DC25B" box="[258,403,559,585]" class="Insecta" family="Geocoridae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Geocoridae</taxonomicName>
|
||
. Similar findings were concluded by
|
||
<bibRefCitation id="EFDB4B35FFE7FFBB4A53CC22FF17C279" author="KMENT, P. & VILIMOVA, J." pageId="5" pageNumber="6" pagination="1 - 59" refId="ref3375" refString="KMENT, P. & VILIMOVA, J. (2010): Thoracic scent efferent system of the Tessaratomidae sen- su lato (Hemiptera: Heteroptera: Pentatomoidea) with implication to the phylogeny of the family. - Zootaxa 2363: 1 - 59. https: // doi. org / 10.11646 / zootaxa. 2363.1.1" type="journal article" year="2010">KMENT and VILIMOVÁ (2010)</bibRefCitation>
|
||
in family
|
||
<taxonomicName id="4C4A4D47FFE7FFBB4878CC43FDD9C279" authorityName="Stal" authorityYear="1864" box="[374,567,593,619]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Tessaratomidae</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C4A4D47FFE7FFBB4B48CC43FD0EC279" box="[582,736,593,619]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
|
||
: Pentatomoidea).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |