187 lines
20 KiB
XML
187 lines
20 KiB
XML
<document id="6E9C5FE1C58EE9E54AD28494744E870C" ID-CLB-Dataset="6390" ID-DOI="10.5252/g2014n4a4" ID-GBIF-Dataset="8bc26961-9853-4113-a63d-4cf691aa7100" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538625" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613150850454" checkinUser="felipe" docAuthor="Vianey-Liaud, Monique, Comte, Bernard, Marandat, Bernard, Peigné, Stéphane, Rage, Jean-Claude & Sudre, Jean" docDate="2014" docId="03D6987B4476B12BFD781139FB3BE3AB" docLanguage="en" docName="Geodiversitas.36.4.565-622.pdf" docOrigin="Geodiversitas 36 (4)" docSource="http://dx.doi.org/10.5252/g2014n4a4" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Pseudeumeces Hoffstetter 1944" docType="treatment" docVersion="7" lastPageNumber="572" masterDocId="FFEFE0034471B12CFF8F106FFF81E50F" masterDocTitle="A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)" masterLastPageNumber="622" masterPageNumber="565" pageNumber="572" updateTime="1698929460103" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="08FDC4102464F48A3C7199174814C028">A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)</mods:title>
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<mods:name id="B7F41D468D36DA1D392D41A0F5D7FCE3" type="personal">
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<mods:namePart id="328CB1C1C3A3E243DF64DD3943BD1F29">Vianey-Liaud, Monique</mods:namePart>
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<mods:namePart id="9AC46521ED0A439D3740F91591195F9C">Comte, Bernard</mods:namePart>
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<mods:namePart id="96C63D88BCC986E61E0DCCA2CBB118FE">Marandat, Bernard</mods:namePart>
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<mods:namePart id="20C4B4181CEE2B270FE8F78343C1D3D4">Peigné, Stéphane</mods:namePart>
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<mods:namePart id="F8C8CDC31C3D96F74E6A292CA8B63B9E">Rage, Jean-Claude</mods:namePart>
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<mods:namePart id="F281B64ABA11C04F01122A0778A131CA">Sudre, Jean</mods:namePart>
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<mods:title id="F00A0442A72D255DD2B01D8FB4B941A4">Geodiversitas</mods:title>
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<mods:date id="F726B2F8C6021CC48165858532DBF3E9">2014</mods:date>
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<mods:number id="09B626137B9E964F8BD07D775ACCF54C">2014-12-26</mods:number>
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<mods:number id="F248E2540D4C29DC83A076430E1EFD5D">36</mods:number>
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<mods:identifier id="4FA0E54A19648AF80E4ACB69033ACE5C" type="DOI">10.5252/g2014n4a4</mods:identifier>
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<treatment id="03D6987B4476B12BFD781139FB3BE3AB" ID-DOI="http://doi.org/10.5281/zenodo.4837344" ID-GBIF-Taxon="182405820" ID-Zenodo-Dep="4837344" LSID="urn:lsid:plazi:treatment:03D6987B4476B12BFD781139FB3BE3AB" httpUri="http://treatment.plazi.org/id/03D6987B4476B12BFD781139FB3BE3AB" lastPageNumber="572" pageId="7" pageNumber="572">
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<subSubSection id="C3657AE64476B12BFD781139FBE6E47E" box="[759,1127,342,369]" pageId="7" pageNumber="572" type="nomenclature">
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<paragraph id="8BC0296D4476B12BFD781139FBE6E47E" blockId="7.[759,1127,342,402]" box="[759,1127,342,369]" pageId="7" pageNumber="572">
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<heading id="D0889E014476B12BFD781139FBE6E47E" box="[759,1127,342,369]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="572" reason="2">
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cf.
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<taxonomicName id="4C7F52EE4476B12BFC971139FBE6E47E" authority="Hoffstetter, 1944" authorityName="Hoffstetter" authorityYear="1944" box="[792,1127,342,369]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFC971139FC25E47F" box="[792,932,342,368]" italics="true" pageId="7" pageNumber="572">Pseudeumeces</emphasis>
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Hoffstetter, 1944
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3657AE64476B12BFC0F1117FC5FE49D" box="[896,990,376,402]" pageId="7" pageNumber="572" type="description">
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<paragraph id="8BC0296D4476B12BFC0F1117FC5FE49D" blockId="7.[759,1127,342,402]" box="[896,990,376,402]" pageId="7" pageNumber="572">
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(
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<figureCitation id="134435E84476B12BFC061117FC54E49D" box="[905,981,376,402]" captionStart="FIG" captionStartId="6.[113,124,1085,1103]" captionTargetBox="[113,1209,215,1042]" captionTargetId="figure-223@6.[387,662,490,767]" captionTargetPageId="6" captionText="FIG. 1. — Amphibians and lizards from Saint-Privat-des-Vieux (Gard; Lower Chattian,MP 26): A-C, Discoglossus Otth, 1837 or Latonia Meyer, 1843: A, left ilium (SPV701) in lateral view; B, right scapula (SPV702) in lateral view; C, right angulare (SPV715) in dorsal view; D, E, Pelophylax sp.: D, right humerus (SPV705) in ventral view; E, left ilium (SPV704) in lateral view; F, cf. Pseudeumeces Hoffstetter, 1944, left dentary (SPV706) in medial view; G, H,?Shinisauria indet., right maxilla (SPV709): G, lateral view; H, medial view; I, am- phisbaenian A, left dentary (SPV707) in medial view; J, amphisbaenian B, right dentary (SPV708) in medial view. Scale bars: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4538629" httpUri="https://zenodo.org/record/4538629/files/figure.png" pageId="7" pageNumber="572">Fig. 1F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3657AE64476B12BFD2A11DFFC8FE71E" pageId="7" pageNumber="572" type="materials_examined">
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<paragraph id="8BC0296D4476B12BFD2A11DFFB3AE4C8" blockId="7.[677,1211,431,455]" box="[677,1211,431,455]" pageId="7" pageNumber="572">
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<collectionCode id="ED6EB1A84476B12BFD2A11DFFC9FE4C9" box="[677,798,432,455]" pageId="7" pageNumber="572">
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<typeStatus id="54C497CF4476B12BFD2A11DFFD54E4CA" box="[677,725,432,455]" pageId="7" pageNumber="572">TYPE</typeStatus>
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SPECIES
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</collectionCode>
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. —
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<taxonomicName id="4C7F52EE4476B12BFCC911C0FB37E4C8" authority="(Filhol, 1877)" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[838,1206,431,455]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="cadurcensis">
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<emphasis id="B90BF57F4476B12BFCC911C0FBAAE4C8" box="[838,1067,431,455]" italics="true" pageId="7" pageNumber="572">Pseudeumeces cadurcensis</emphasis>
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(
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<bibRefCitation id="EFEE549C4476B12BFBB911C0FB2EE4C8" author="FILHOL H." box="[1078,1199,431,455]" pageId="7" pageNumber="572" refId="ref32651" refString="FILHOL H. 1877. - Recherches sur les phosphorites du Quercy. Etude des fossiles qu'on y rencontre et specialement des mammiferes. G. Masson, Paris, 561 p." type="book" year="1877">Filhol, 1877</bibRefCitation>
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)
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BC0296D4476B12BFD2A11B0FC8FE71E" blockId="7.[677,1209,479,529]" pageId="7" pageNumber="572">
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MATERIAL EXAMINED. — One anterior part of dentary (
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<collectionCode id="ED6EB1A84476B12BFD241196FD5AE71E" box="[683,731,505,529]" pageId="7" pageNumber="572">SPV</collectionCode>
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706).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3657AE64476B12BFD2A125AFB35E664" pageId="7" pageNumber="572" type="description">
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<paragraph id="8BC0296D4476B12BFD2A125AFCB4E742" blockId="7.[675,1212,565,876]" box="[677,821,565,591]" pageId="7" pageNumber="572">DESCRIPTION</paragraph>
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<paragraph id="8BC0296D4476B12BFD2A123BFB35E664" blockId="7.[675,1212,565,876]" pageId="7" pageNumber="572">
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Ŋe dentary is deep and its subdental shelf (sensu
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<bibRefCitation id="EFEE549C4476B12BFD2A121BFCF2E781" author="RAGE J. C. & AUGE M." box="[677,883,628,654]" pageId="7" pageNumber="572" pagination="253 - 268" refId="ref34338" refString="RAGE J. C. & AUGE M. 2010. - Squamate reptiles from the middle Eocene of Lissieu (France). A landmark in the middle Eocene of Europe. Geobios 43: 253 - 268." type="journal article" year="2010">Rage & Augé 2010</bibRefCitation>
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) markedly thickens anteriorly. Teeth are pleurodont; the most anterior ones are not enlarged but the last tooth of the preserved row is amblyodont, i.e. it is larger than normal teeth. Ŋe enlargement, from the most anterior to the more posterior teeth is progressive. Ŋe apices are round and devoid of cuspids. Striae are present on the apices but they are weak on the largest tooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3657AE64476B12BFD2A13FEFB3BE3AB" pageId="7" pageNumber="572" type="discussion">
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<paragraph id="8BC0296D4476B12BFD2A13FEFCA7E6A6" blockId="7.[674,1212,913,1700]" box="[677,806,913,939]" pageId="7" pageNumber="572">DISCUSSION</paragraph>
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<paragraph id="8BC0296D4476B12BFD2A13DFFB3BE3AB" blockId="7.[674,1212,913,1700]" pageId="7" pageNumber="572">
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Several lizard taxa with amblyodont teeth, or showing tendency toward amblyodonty, are known in the European Cenozoic, mainly from the Oligocene (
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<bibRefCitation id="EFEE549C4476B12BFB09139FFD5DE125" author="RAGE J. C." pageId="7" pageNumber="572" pagination="145 - 149" refId="ref34257" refString="RAGE J. C. 1987. - Extinctions chez les Squamates (Reptiles) a la fin de l'Oligocene en France. Adaptations et modifications de l'environnement. Memoires de la Societe geologique de France 150: 145 - 149." type="journal article" year="1987">Rage 1987</bibRefCitation>
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,
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<bibRefCitation id="EFEE549C4476B12BFD6B147FFCD4E125" author="AUGE M." box="[740,853,1039,1066]" pageId="7" pageNumber="572" pagination="5 - 369" refId="ref31134" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Histoire naturelle 192: 5 - 369." type="journal article" year="2005">Augé 2005</bibRefCitation>
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).Ŋe amblyodont condition is weak in
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<taxonomicName id="4C7F52EE4476B12BFD4F1440FB87E145" authority="Auge, 2005" authorityName="Auge" authorityYear="2005" box="[704,1030,1071,1098]" class="Reptilia" family="Lacertidae" genus="Mediolacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="roceki">
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<emphasis id="B90BF57F4476B12BFD4F1440FC07E146" box="[704,902,1071,1097]" italics="true" pageId="7" pageNumber="572">Mediolacerta roceki</emphasis>
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<bibRefCitation id="EFEE549C4476B12BFC041440FB87E145" author="AUGE M." box="[907,1030,1071,1098]" pageId="7" pageNumber="572" pagination="5 - 369" refId="ref31134" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Histoire naturelle 192: 5 - 369." type="journal article" year="2005">Augé, 2005</bibRefCitation>
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</taxonomicName>
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(MP 23-MP 30) and is very prominent in
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<taxonomicName id="4C7F52EE4476B12BFC4E143FFCB5E186" authority="Gervais, 1848" authorityName="Gervais" authorityYear="1848" class="Reptilia" family="Lacertidae" genus="Dracaenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="croizeti">
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<emphasis id="B90BF57F4476B12BFC4E143FFB38E166" box="[961,1209,1104,1129]" italics="true" pageId="7" pageNumber="572">Dracaenosaurus croizeti</emphasis>
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<bibRefCitation id="EFEE549C4476B12BFD291401FCB5E186" author="GERVAIS P." box="[678,820,1134,1161]" pageId="7" pageNumber="572" refId="ref32885" refString="GERVAIS P. 1848 - 1852. - Zoologie et Paleontologie francaises (animaux vertebres), Bertrand, Paris, 3 volumes: 271 p., 45 pls." type="book" year="1848">Gervais, 1848</bibRefCitation>
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</taxonomicName>
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-52 (MP28-MP 30). Morphologically, between these two species occur the intermediate taxa
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<taxonomicName id="4C7F52EE4476B12BFD5614C1FC7EE1C7" authority="Rocek, 1884" authorityName="Rocek" authorityYear="1884" box="[729,1023,1198,1224]" class="Reptilia" family="Lacertidae" genus="Amblyolacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFD5614C1FCEDE1C7" box="[729,876,1198,1224]" italics="true" pageId="7" pageNumber="572">Amblyolacerta</emphasis>
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Roček, 1884
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</taxonomicName>
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and
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<taxonomicName id="4C7F52EE4476B12BFBB614C0FC51E1E7" authority="Auge, Bailon & Malfay, 2003" authorityName="Auge, Bailon & Malfay" authorityYear="2003" class="Reptilia" family="Lacertidae" genus="Ligerosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFBB614C0FB3BE1C7" box="[1081,1210,1199,1224]" italics="true" pageId="7" pageNumber="572">Ligerosaurus</emphasis>
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Augé, Bailon & Malfay, 2003
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</taxonomicName>
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from the Miocene and
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<taxonomicName id="4C7F52EE4476B12BFD2D1482FCAAE008" authorityName="Hoffstetter" authorityYear="1944" box="[674,811,1261,1287]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFD2D1482FCAAE008" box="[674,811,1261,1287]" italics="true" pageId="7" pageNumber="572">Pseudeumeces</emphasis>
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</taxonomicName>
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from the Oligocene. Ŋe fossil from Saint-Privat-des-Vieux may be distinguished from
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<taxonomicName id="4C7F52EE4476B12BFD2A1542FCB7E048" authorityName="Rocek" authorityYear="1884" box="[677,822,1325,1351]" class="Reptilia" family="Lacertidae" genus="Amblyolacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFD2A1542FCB7E048" box="[677,822,1325,1351]" italics="true" pageId="7" pageNumber="572">Amblyolacerta</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C7F52EE4476B12BFCE21542FBF9E048" authority="Auge, 2005" authorityName="Auge" authorityYear="2005" box="[877,1144,1324,1351]" class="Reptilia" family="Lacertidae" genus="Mediolacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFCE21542FC73E048" box="[877,1010,1325,1351]" italics="true" pageId="7" pageNumber="572">Mediolacerta</emphasis>
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<bibRefCitation id="EFEE549C4476B12BFC751542FBF9E048" author="AUGE M." box="[1018,1144,1324,1351]" pageId="7" pageNumber="572" pagination="5 - 369" refId="ref31134" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Memoires du Museum national d'Histoire naturelle 192: 5 - 369." type="journal article" year="2005">Augé, 2005</bibRefCitation>
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</taxonomicName>
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by its stronger amblyodonty. It differs from
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<taxonomicName id="4C7F52EE4476B12BFBAA1522FB20E069" authorityName="Auge, Bailon & Malfay" authorityYear="2003" box="[1061,1185,1357,1382]" class="Reptilia" family="Lacertidae" genus="Ligerosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFBAA1522FB20E069" box="[1061,1185,1357,1382]" italics="true" pageId="7" pageNumber="572">Ligerosaurus</emphasis>
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</taxonomicName>
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in having a subdental shelf that grows clearly thinner posteriorly and a facet for contact with the splenial that does not reach the anterior part of the bone. In contrast, the fossil does not show any difference with
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<taxonomicName id="4C7F52EE4476B12BFD521584FCE7E30A" authorityName="Hoffstetter" authorityYear="1944" box="[733,870,1515,1541]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFD521584FCE7E30A" box="[733,870,1515,1541]" italics="true" pageId="7" pageNumber="572">Pseudeumeces</emphasis>
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</taxonomicName>
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and only the incomplete nature of the specimen prevents confident referral to this genus.
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<taxonomicName id="4C7F52EE4476B12BFCAD1645FC2AE34B" authorityName="Hoffstetter" authorityYear="1944" box="[802,939,1578,1604]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="genus">
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<emphasis id="B90BF57F4476B12BFCAD1645FC2AE34B" box="[802,939,1578,1604]" italics="true" pageId="7" pageNumber="572">Pseudeumeces</emphasis>
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</taxonomicName>
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|
includes a single species,
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<taxonomicName id="4C7F52EE4476B12BFD291624FC59E36B" authority="(Filhol, 1877)" baseAuthorityName="Filhol" baseAuthorityYear="1877" box="[678,984,1610,1636]" class="Reptilia" family="Lacertidae" genus="Pseudeumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="572" phylum="Chordata" rank="species" species="cadurcensis">
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<emphasis id="B90BF57F4476B12BFD291624FD35E36B" box="[678,692,1611,1636]" italics="true" pageId="7" pageNumber="572">P</emphasis>
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.
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<emphasis id="B90BF57F4476B12BFD4B1625FCB4E36B" box="[708,821,1610,1636]" italics="true" pageId="7" pageNumber="572">cadurcensis</emphasis>
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(
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<bibRefCitation id="EFEE549C4476B12BFCCA1625FC51E36B" author="FILHOL H." box="[837,976,1610,1636]" pageId="7" pageNumber="572" refId="ref32651" refString="FILHOL H. 1877. - Recherches sur les phosphorites du Quercy. Etude des fossiles qu'on y rencontre et specialement des mammiferes. G. Masson, Paris, 561 p." type="book" year="1877">Filhol, 1877</bibRefCitation>
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)
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</taxonomicName>
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, that is known from MP 25 to MP 28. Apparently, amblyodont lizards fed at least partly on shelled mollusks (gastropods).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |