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<document id="0E1A52592D64FDC0092154DC9B5F2C09" ID-CLB-Dataset="3794" ID-DOI="10.11646/zootaxa.5060.1.7" ID-GBIF-Dataset="03e2aa8b-e28f-427d-9be9-b311e591c2f0" ID-ISSN="1175-5326" ID-Zenodo-Dep="5606848" ID-ZooBank="B7FD8234-A78D-47AC-93D5-8D6DF80356E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635407248973" checkinUser="plazi" docAuthor="Gong, Yan-An, Peng, Li-Fang, Huang, Song, Lin, Yan-Feng, Huang, Ru-Yi, Xu, Yu-Hao, Yang, Dian-Cheng &amp; Nie, Liu-Wang" docDate="2021" docId="03B5E551FFE1FFA4FF56E5F0FE12F9B5" docLanguage="en" docName="zootaxa.5060.1.7.pdf" docOrigin="Zootaxa 5060 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelodiscus huangshanensis Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie 2021, sp. nov." docType="treatment" docUuid="0C377CC0-9B47-420E-BB22-D4BF1D113014" docUuidSource="ZooBank" docVersion="5" lastPageNumber="143" masterDocId="FF8C9D29FFE2FFA2FFC1E175FF88FFFB" masterDocTitle="A new species of the Genus Pelodiscus Fitzinger, 1835 (Testudines: Trionychidae) from Huangshan, Anhui, China" masterLastPageNumber="145" masterPageNumber="137" pageNumber="140" updateTime="1699467049518" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D425C5435F9003CE0230018EF8748EEA">A new species of the Genus Pelodiscus Fitzinger, 1835 (Testudines: Trionychidae) from Huangshan, Anhui, China</mods:title>
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<mods:namePart id="09B41E8118B3EB4D1752CE73C5FCB505">Gong, Yan-An</mods:namePart>
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<mods:namePart id="837B2BA548867B88A86A4BECB800F6FC">Xu, Yu-Hao</mods:namePart>
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<mods:namePart id="5197573C66775A07A773013EAC5307D8">Nie, Liu-Wang</mods:namePart>
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<mods:affiliation id="E4A4D93A343E17EBE49CFFCEDC0EE862">Anhui Province Key Laboratory of the Conservation and Exploitation of Biological Resource, College of Life Sciences, Anhui Normal University, Wuhu, 241000, China &amp; lwnie @ ahnu. edu. cn; https: // orcid. org / 0000 - 0003 - 2660 - 5799</mods:affiliation>
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<treatment id="03B5E551FFE1FFA4FF56E5F0FE12F9B5" ID-DOI="http://doi.org/10.5281/zenodo.5606912" ID-GBIF-Taxon="189550194" ID-Zenodo-Dep="5606912" LSID="urn:lsid:zoobank.org:act:0C377CC0-9B47-420E-BB22-D4BF1D113014" httpUri="http://treatment.plazi.org/id/03B5E551FFE1FFA4FF56E5F0FE12F9B5" lastPageId="6" lastPageNumber="143" pageId="3" pageNumber="140">
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<taxonomicName id="4C1C2FC4FFE1FFA1FF56E5F0FE6CFB5B" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[151,484,1157,1184]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="140" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE1FFA1FF56E5F0FE6CFB5B" bold="true" box="[151,484,1157,1184]" italics="true" pageId="3" pageNumber="140">Pelodiscus huangshanensis</emphasis>
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<taxonomicNameLabel id="A25B352EFFE1FFA1FE2DE5F3FDC1FB5B" box="[492,585,1158,1184]" pageId="3" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
Gong, Peng, Huang &amp; Nie, 2021
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(
<figureCitation id="132748C2FFE1FFA1FF5EE5DCFE82FB3F" box="[159,266,1193,1220]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[151,250,1427,1451]" captionStartId-1="5.[151,250,1080,1104]" captionStartId-2="5.[151,250,1807,1831]" captionStartId-3="6.[151,250,689,713]" captionTargetBox-0="[163,1424,756,1393]" captionTargetBox-1="[165,1420,196,1044]" captionTargetBox-2="[151,1436,1128,1784]" captionTargetBox-3="[152,1436,181,665]" captionTargetId-0="figure-225@4.[151,1437,740,1403]" captionTargetId-1="figure-19@5.[151,1435,181,1056]" captionTargetId-2="figure-39@5.[151,1436,1128,1784]" captionTargetId-3="figure-23@6.[151,1436,181,665]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 2. Dorsal and ventral aspects of the holotype of Pelodiscus huangshanensis sp. nov. (ANU20210001, adult male)" captionText-1="FIGURE 3. Dorsal and ventral aspects of the paratypes of Pelodiscus huangshanensis sp. nov. (ANU20210002, adult female)" captionText-2="FIGURE 4. Head and neck of Pelodiscus huangshanensis sp. nov.; A, B ANU20210001, male; C, D ANU20210004, juvenile." captionText-3="FIGURE 5. The three months old Huangshan soft-shelled turtles." figureDoi-0="http://doi.org/10.5281/zenodo.5606856" figureDoi-1="http://doi.org/10.5281/zenodo.5606860" figureDoi-2="http://doi.org/10.5281/zenodo.5606864" figureDoi-3="http://doi.org/10.5281/zenodo.5606866" httpUri-0="https://zenodo.org/record/5606856/files/figure.png" httpUri-1="https://zenodo.org/record/5606860/files/figure.png" httpUri-2="https://zenodo.org/record/5606864/files/figure.png" httpUri-3="https://zenodo.org/record/5606866/files/figure.png" pageId="3" pageNumber="140">Figs. 25</figureCitation>
)
</paragraph>
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<paragraph id="8BA35447FFE1FFA1FF56E584FCA2FAF7" blockId="3.[151,1437,1265,2012]" box="[151,810,1265,1292]" pageId="3" pageNumber="140">
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<emphasis id="B9688855FFE1FFA1FF56E584FCA2FAF7" bold="true" box="[151,810,1265,1292]" pageId="3" pageNumber="140">Suggested English name: Huangshan soft-shelled turtle</emphasis>
</heading>
</paragraph>
<paragraph id="8BA35447FFE1FFA1FF06E460FC25FAD4" blockId="3.[151,1437,1265,2012]" box="[199,941,1301,1328]" pageId="3" pageNumber="140">
<emphasis id="B9688855FFE1FFA1FF06E460FC25FAD4" bold="true" box="[199,941,1301,1328]" pageId="3" pageNumber="140">Suggested Chinese name: •山ƼÞ× (Huáng Shān Mǎ tí Biē)</emphasis>
</paragraph>
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</subSubSection>
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<paragraph id="8BA35447FFE1FFA1FF06E44FFBD8FA3B" blockId="3.[151,1437,1265,2012]" pageId="3" pageNumber="140">
<materialsCitation id="3B745E1AFFE1FFA1FF06E44FFBD8FA3B" ID-GBIF-Occurrence="3395067301" collectingDate="2017-06-10" collectionCode="ANU" country="China" county="Xiuning County" elevation="170" location="The" municipality="The" pageId="3" pageNumber="140" specimenCode="ANU20210001" specimenCount="3" specimenCount-adult="1" specimenCount-male="1" stateProvince="Anhui Province" typeStatus="holotype">
<emphasis id="B9688855FFE1FFA1FF06E44FFEB3FAAF" bold="true" box="[199,315,1338,1364]" pageId="3" pageNumber="140">
<typeStatus id="54A7EAE5FFE1FFA1FF06E44FFEBEFAAF" box="[199,310,1338,1364]" pageId="3" pageNumber="140" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBBAFC3CFFE1FFA1FE82E44FFE7BFAAF" box="[323,499,1338,1364]" pageId="3" pageNumber="140">ANU20210001</specimenCode>
, an
<specimenCount id="9D1A9FCEFFE1FFA1FDE0E44CFDD1FAA8" box="[545,601,1337,1363]" pageId="3" pageNumber="140" type="adult">adult</specimenCount>
<specimenCount id="9D1A9FCEFFE1FFA1FDA0E44CFD1FFAA8" box="[609,663,1337,1363]" pageId="3" pageNumber="140" type="male">male</specimenCount>
(
<figureCitation id="132748C2FFE1FFA1FD69E44CFD66FAAF" box="[680,750,1337,1364]" captionStart="FIGURE 2" captionStartId="4.[151,250,1427,1451]" captionTargetBox="[163,1424,756,1393]" captionTargetId="figure-225@4.[151,1437,740,1403]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal and ventral aspects of the holotype of Pelodiscus huangshanensis sp. nov. (ANU20210001, adult male)" figureDoi="http://doi.org/10.5281/zenodo.5606856" httpUri="https://zenodo.org/record/5606856/files/figure.png" pageId="3" pageNumber="140">Fig. 2</figureCitation>
), was found in a stream from Xikou Town,
<collectingCounty id="62C22CCBFFE1FFA1FB20E44CFA10FAAF" box="[1249,1432,1337,1364]" pageId="3" pageNumber="140">Xiuning County</collectingCounty>
, Huangshan City,
<collectingRegion id="49D89AA5FFE1FFA1FE99E428FD84FA8C" box="[344,524,1373,1399]" country="China" name="Anhui" pageId="3" pageNumber="140">Anhui Province</collectingRegion>
,
<collectingCountry id="F30B14D7FFE1FFA1FDD9E428FDD3FA8C" box="[536,603,1373,1399]" name="China" pageId="3" pageNumber="140">China</collectingCountry>
(118.02°N, 29.69°E, and
<elevation id="0031B374FFE1FFA1FC41E42BFC40FA83" box="[896,968,1374,1400]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="140" unit="m" value="170.0">170 m</elevation>
A. S. L.). The coordinates are given only with a precision of 0.01 degrees to avoid private or commercial sampling. The specimen was collected on
<date id="FFA27287FFE1FFA1FA84E4F7FF47FA3B" pageId="3" pageNumber="140" value="2017-06-10">
<collectingDate id="EFE68B6FFFE1FFA1FA84E4F7FF47FA3B" pageId="3" pageNumber="140" value="2017-06-10">10 June 2017</collectingDate>
</date>
and deposited at Anhui Normal University Museum (voucher:
<specimenCode id="DBBAFC3CFFE1FFA1FC53E4D3FBCCFA3B" box="[914,1092,1446,1472]" pageId="3" pageNumber="140">ANU20210001</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA35447FFE1FFA1FF06E4BFFE43F9D0" blockId="3.[151,1437,1265,2012]" pageId="3" pageNumber="140">
<emphasis id="B9688855FFE1FFA1FF06E4BFFECEFA1F" bold="true" box="[199,326,1482,1508]" pageId="3" pageNumber="140">
<typeStatus id="54A7EAE5FFE1FFA1FF06E4BFFEC9FA1F" box="[199,321,1482,1508]" pageId="3" pageNumber="140" type="paratype">Paratypes</typeStatus>
.
</emphasis>
ANU20210002, adult female (
<figureCitation id="132748C2FFE1FFA1FD79E4BCFD73FA1F" box="[696,763,1481,1508]" captionStart="FIGURE 3" captionStartId="5.[151,250,1080,1104]" captionTargetBox="[165,1420,196,1044]" captionTargetId="figure-19@5.[151,1435,181,1056]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal and ventral aspects of the paratypes of Pelodiscus huangshanensis sp. nov. (ANU20210002, adult female)" figureDoi="http://doi.org/10.5281/zenodo.5606860" httpUri="https://zenodo.org/record/5606860/files/figure.png" pageId="3" pageNumber="140">Fig 3</figureCitation>
); ANU20210003, adult female; ANU20210004, juvenile; ANU20210005, juvenile; CIB116368, juvenile; SNHM5414, juvenile. They were sampled from May to June, 2017 around the site of
<typeStatus id="54A7EAE5FFE1FFA1FEA0E764FE4FF9D0" box="[353,455,1553,1579]" pageId="3" pageNumber="140" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30607CCFFE1FFA1FF06E740FD99F8F8" pageId="3" pageNumber="140" type="diagnosis">
<paragraph id="8BA35447FFE1FFA1FF06E740FD99F8F8" blockId="3.[151,1437,1265,2012]" pageId="3" pageNumber="140">
<emphasis id="B9688855FFE1FFA1FF06E740FECAF9B4" bold="true" box="[199,322,1589,1615]" pageId="3" pageNumber="140">Diagnosis.</emphasis>
<taxonomicName id="4C1C2FC4FFE1FFA1FE8AE740FD09F9B4" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[331,641,1589,1615]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="140" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE1FFA1FE8AE740FD09F9B4" box="[331,641,1589,1615]" italics="true" pageId="3" pageNumber="140">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE1FFA1FD4BE743FD6EF9AB" bold="true" box="[650,742,1590,1616]" pageId="3" pageNumber="140">
<taxonomicNameLabel id="A25B352EFFE1FFA1FD4BE743FD6EF9AB" box="[650,742,1590,1616]" pageId="3" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the five known congeners by a combination of the following characters: (1) Small size; maximum carapace length of 84.0 mm and maximum body length of
<quantity id="4CE4F9A2FFE1FFA1FAB3E72FFF4EF963" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="140" unit="mm" value="190.0">190 mm</quantity>
; (2) keel high; (3) tiny yellowish-white spots on the throat; (4) no black pinstripes around the eyes; (5) white longitudinal bands on both sides of the neck in juveniles, absent in adults (
<figureCitation id="132748C2FFE1FFA1FC10E7D4FB9EF947" box="[977,1046,1697,1724]" captionStart="FIGURE 4" captionStartId="5.[151,250,1807,1831]" captionTargetBox="[151,1436,1128,1784]" captionTargetId="figure-39@5.[151,1436,1128,1784]" captionTargetPageId="5" captionText="FIGURE 4. Head and neck of Pelodiscus huangshanensis sp. nov.; A, B ANU20210001, male; C, D ANU20210004, juvenile." figureDoi="http://doi.org/10.5281/zenodo.5606864" httpUri="https://zenodo.org/record/5606864/files/figure.png" pageId="3" pageNumber="140">Fig. 4</figureCitation>
); (6) plastron yellowish-white, and only a dark patch on each side of the armpit; (7) many tubercles on the dorsal surface, but indistinct in the center; and (8) entoplastron “⌒” shaped.
</paragraph>
</subSubSection>
<subSubSection id="C30607CCFFE1FFA6FF06E678FBFFF97F" lastPageId="4" lastPageNumber="141" pageId="3" pageNumber="140" type="description">
<paragraph id="8BA35447FFE1FFA6FF06E678FD9EFEE6" blockId="3.[151,1437,1265,2012]" lastBlockId="4.[151,1437,151,286]" lastPageId="4" lastPageNumber="141" pageId="3" pageNumber="140">
<emphasis id="B9688855FFE1FFA1FF06E678FD05F8DC" bold="true" box="[199,653,1805,1832]" pageId="3" pageNumber="140">
Description of
<typeStatus id="54A7EAE5FFE1FFA1FEB4E67BFE54F8D3" box="[373,476,1806,1832]" pageId="3" pageNumber="140" type="holotype">holotype</typeStatus>
and Variation.
</emphasis>
Where they differ from the
<typeStatus id="54A7EAE5FFE1FFA1FC02E678FBA0F8DC" box="[963,1064,1805,1831]" pageId="3" pageNumber="140" type="holotype">holotype</typeStatus>
, features of the
<typeStatus id="54A7EAE5FFE1FFA1FB16E67BFACAF8D3" box="[1239,1346,1806,1832]" pageId="3" pageNumber="140" type="paratype">paratypes</typeStatus>
are listed in parentheses (ANU202100025, CIB116368, SNHM5414).
<typeStatus id="54A7EAE5FFE1FFA1FCB2E644FC53F8B0" box="[883,987,1841,1867]" pageId="3" pageNumber="140" type="holotype">Holotype</typeStatus>
of an adult male (the first two are adult females and the last four are juveniles); carapace length
<quantity id="4CE4F9A2FFE1FFA1FCCAE620FCE7F88B" box="[779,879,1877,1904]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" pageId="3" pageNumber="140" unit="mm" value="82.5">82.5 mm</quantity>
(adult females 73.484.0 mm, juvenile
<quantity id="4CE4F9A2FFE1FFA1FAEDE620FF4FF86F" metricMagnitude="-2" metricUnit="m" metricValue="4.82" metricValueMax="5.59" metricValueMin="4.05" pageId="3" pageNumber="140" unit="mm" value="48.2" valueMax="55.9" valueMin="40.5">40.555.9 mm</quantity>
); plastron length
<quantity id="4CE4F9A2FFE1FFA1FE47E60FFE61F86F" box="[390,489,1914,1940]" metricMagnitude="-2" metricUnit="m" metricValue="7.93" pageId="3" pageNumber="140" unit="mm" value="79.3">79.3 mm</quantity>
(adult females
<quantity id="4CE4F9A2FFE1FFA1FD52E60CFCBFF86F" box="[659,823,1913,1940]" metricMagnitude="-2" metricUnit="m" metricValue="8.09" metricValueMax="8.61" metricValueMin="7.57" pageId="3" pageNumber="140" unit="mm" value="80.9" valueMax="86.1" valueMin="75.7">75.786.1 mm</quantity>
, juvenile
<quantity id="4CE4F9A2FFE1FFA1FC63E60CFBC0F86F" box="[930,1096,1913,1940]" metricMagnitude="-2" metricUnit="m" metricValue="4.784999999999999" metricValueMax="5.58" metricValueMin="3.9899999999999998" pageId="3" pageNumber="140" unit="mm" value="47.849999999999994" valueMax="55.8" valueMin="39.9">39.955.8 mm</quantity>
); body length 190.0 mm (adult females 144.9160.0 mm, juvenile
<quantity id="4CE4F9A2FFE1FFA1FDE0E6EBFD59F843" box="[545,721,1950,1976]" metricMagnitude="-1" metricUnit="m" metricValue="1.0145" metricValueMax="1.147" metricValueMin="0.882" pageId="3" pageNumber="140" unit="mm" value="101.45" valueMax="114.7" valueMin="88.2">88.2114.7 mm</quantity>
) (
<tableCitation id="C69E61FCFFE1FFA1FD26E6E8FCB5F843" box="[743,829,1949,1976]" captionStart="TABLE 3" captionStartId="4.[151,239,332,356]" captionTargetPageId="4" captionText="TABLE 3. Measurements of Pelodiscus huangshanensis sp. nov. (mm), for abbreviations, see the Materials and Methods." httpUri="http://table.plazi.org/id/DF6304CFFFE6FFA6FF56E039FA15FE9F" pageId="3" pageNumber="140" tableUuid="DF6304CFFFE6FFA6FF56E039FA15FE9F">Table 3</tableCitation>
). Back and side of head have many fine dark brown to black spots; no black pinstripes around the eyes. The neck shows no white longitudinal striae bilaterally (there are white longitudinal bands on both sides of the neck in juveniles). The throat has many tiny yellowish-white spots. The carapace slightly oval with a high keel. The carapace is olive brown to dark brown. The dorsal surface has many tubercles, but it is indistinct in the center. The plastron yellowish-white, and only a dark patch on each side of the armpit. Entoplastron “⌒” shaped.
</paragraph>
<caption id="DF6304CFFFE6FFA6FF56E039FA15FE9F" ID-Table-UUID="DF6304CFFFE6FFA6FF56E039FA15FE9F" box="[151,1437,332,357]" httpUri="http://table.plazi.org/id/DF6304CFFFE6FFA6FF56E039FA15FE9F" pageId="4" pageNumber="141" startId="4.[151,239,332,356]" targetBox="[166,1392,373,678]" targetIsTable="true" targetPageId="4">
<paragraph id="8BA35447FFE6FFA6FF56E039FA15FE9F" blockId="4.[151,1437,332,357]" box="[151,1437,332,357]" pageId="4" pageNumber="141">
<emphasis id="B9688855FFE6FFA6FF56E039FE81FE9F" bold="true" box="[151,265,332,356]" pageId="4" pageNumber="141">TABLE 3.</emphasis>
Measurements of
<taxonomicName id="4C1C2FC4FFE6FFA6FE0AE039FD66FE9F" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[459,750,332,356]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="141" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE6FFA6FE0AE039FD66FE9F" box="[459,750,332,356]" italics="true" pageId="4" pageNumber="141">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE6FFA6FD35E038FCC0FE9E" bold="true" box="[756,840,333,357]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="A25B352EFFE6FFA6FD35E038FCC0FE9E" box="[756,840,333,357]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(mm), for abbreviations, see the Materials and Methods.
</paragraph>
</caption>
<paragraph id="8BA35447FFE6FFA6FE62E000FAE7FD5D" pageId="4" pageNumber="141">
<table id="F91CA6E7FFE6005DFF67E000FAF8FD5D" box="[166,1392,373,678]" gridcols="11" gridrows="8" pageId="4" pageNumber="141">
<tr id="352C5605FFE6005DFF67E000FAF8FE76" box="[166,1392,373,397]" gridrow="0" pageId="4" pageNumber="141" rowspan-0="1">
<th id="76FD3F79FFE6005DFE62E000FE7DFE76" box="[419,501,373,397]" gridcol="1" gridrow="0" pageId="4" pageNumber="141">Sex</th>
<th id="76FD3F79FFE6005DFDF6E000FDEBFE76" box="[567,611,373,397]" gridcol="2" gridrow="0" pageId="4" pageNumber="141">HL</th>
<th id="76FD3F79FFE6005DFD57E000FD4AFE76" box="[662,706,373,397]" gridcol="3" gridrow="0" pageId="4" pageNumber="141">HW</th>
<th id="76FD3F79FFE6005DFD34E000FC9DFE76" box="[757,789,373,397]" gridcol="4" gridrow="0" pageId="4" pageNumber="141">SL</th>
<th id="76FD3F79FFE6005DFC83E000FCECFE76" box="[834,868,373,397]" gridcol="5" gridrow="0" pageId="4" pageNumber="141">ED</th>
<th id="76FD3F79FFE6005DFC4EE000FC48FE76" box="[911,960,373,397]" gridcol="6" gridrow="0" pageId="4" pageNumber="141">DEL</th>
<th id="76FD3F79FFE6005DFC34E000FBB3FE76" box="[1013,1083,373,397]" gridcol="7" gridrow="0" pageId="4" pageNumber="141">CL</th>
<th id="76FD3F79FFE6005DFBB9E000FB2CFE76" box="[1144,1188,373,397]" gridcol="8" gridrow="0" pageId="4" pageNumber="141">CW</th>
<th id="76FD3F79FFE6005DFB16E000FA8BFE76" box="[1239,1283,373,397]" gridcol="9" gridrow="0" pageId="4" pageNumber="141">PL</th>
<th id="76FD3F79FFE6005DFAF6E000FAF8FE76" box="[1335,1392,373,397]" gridcol="10" gridrow="0" pageId="4" pageNumber="141">BL</th>
</tr>
<tr id="352C5605FFE6005DFF67E0E8FAF8FE4E" box="[166,1392,413,437]" gridrow="1" pageId="4" pageNumber="141">
<th id="76FD3F79FFE6005DFF67E0E8FECAFE4E" box="[166,322,413,437]" gridcol="0" gridrow="1" pageId="4" pageNumber="141">ANU20210001</th>
<td id="76FD3F79FFE6005DFE62E0E8FE7DFE4E" box="[419,501,413,437]" gridcol="1" gridrow="1" pageId="4" pageNumber="141">male</td>
<td id="76FD3F79FFE6005DFDF6E0E8FDEBFE4E" box="[567,611,413,437]" gridcol="2" gridrow="1" pageId="4" pageNumber="141">51.7</td>
<td id="76FD3F79FFE6005DFD57E0E8FD4AFE4E" box="[662,706,413,437]" gridcol="3" gridrow="1" pageId="4" pageNumber="141">25.8</td>
<td id="76FD3F79FFE6005DFD34E0E8FC9DFE4E" box="[757,789,413,437]" gridcol="4" gridrow="1" pageId="4" pageNumber="141">5.9</td>
<td id="76FD3F79FFE6005DFC83E0E8FCECFE4E" box="[834,868,413,437]" gridcol="5" gridrow="1" pageId="4" pageNumber="141">5.3</td>
<td id="76FD3F79FFE6005DFC4EE0E8FC48FE4E" box="[911,960,413,437]" gridcol="6" gridrow="1" pageId="4" pageNumber="141">7.0</td>
<td id="76FD3F79FFE6005DFC34E0E8FBB3FE4E" box="[1013,1083,413,437]" gridcol="7" gridrow="1" pageId="4" pageNumber="141">90.69</td>
<td id="76FD3F79FFE6005DFBB9E0E8FB2CFE4E" box="[1144,1188,413,437]" gridcol="8" gridrow="1" pageId="4" pageNumber="141">80.1</td>
<td id="76FD3F79FFE6005DFB16E0E8FA8BFE4E" box="[1239,1283,413,437]" gridcol="9" gridrow="1" pageId="4" pageNumber="141">79.3</td>
<td id="76FD3F79FFE6005DFAF6E0E8FAF8FE4E" box="[1335,1392,413,437]" gridcol="10" gridrow="1" pageId="4" pageNumber="141">190.0</td>
</tr>
<tr id="352C5605FFE6005DFF67E0B0FAF8FE26" box="[166,1392,453,477]" gridrow="2" pageId="4" pageNumber="141">
<th id="76FD3F79FFE6005DFF67E0B0FECAFE26" box="[166,322,453,477]" gridcol="0" gridrow="2" pageId="4" pageNumber="141">ANU20210002</th>
<td id="76FD3F79FFE6005DFE62E0B0FE7DFE26" box="[419,501,453,477]" gridcol="1" gridrow="2" pageId="4" pageNumber="141">female</td>
<td id="76FD3F79FFE6005DFDF6E0B0FDEBFE26" box="[567,611,453,477]" gridcol="2" gridrow="2" pageId="4" pageNumber="141">38.5</td>
<td id="76FD3F79FFE6005DFD57E0B0FD4AFE26" box="[662,706,453,477]" gridcol="3" gridrow="2" pageId="4" pageNumber="141">20.8</td>
<td id="76FD3F79FFE6005DFD34E0B0FC9DFE26" box="[757,789,453,477]" gridcol="4" gridrow="2" pageId="4" pageNumber="141">6.6</td>
<td id="76FD3F79FFE6005DFC83E0B0FCECFE26" box="[834,868,453,477]" gridcol="5" gridrow="2" pageId="4" pageNumber="141">5.4</td>
<td id="76FD3F79FFE6005DFC4EE0B0FC48FE26" box="[911,960,453,477]" gridcol="6" gridrow="2" pageId="4" pageNumber="141">5.2</td>
<td id="76FD3F79FFE6005DFC34E0B0FBB3FE26" box="[1013,1083,453,477]" gridcol="7" gridrow="2" pageId="4" pageNumber="141">101.16</td>
<td id="76FD3F79FFE6005DFBB9E0B0FB2CFE26" box="[1144,1188,453,477]" gridcol="8" gridrow="2" pageId="4" pageNumber="141">84.3</td>
<td id="76FD3F79FFE6005DFB16E0B0FA8BFE26" box="[1239,1283,453,477]" gridcol="9" gridrow="2" pageId="4" pageNumber="141">86.1</td>
<td id="76FD3F79FFE6005DFAF6E0B0FAF8FE26" box="[1335,1392,453,477]" gridcol="10" gridrow="2" pageId="4" pageNumber="141">160.0</td>
</tr>
<tr id="352C5605FFE6005DFF67E098FAF8FDFD" box="[166,1392,493,518]" gridrow="3" pageId="4" pageNumber="141">
<th id="76FD3F79FFE6005DFF67E098FECAFDFD" box="[166,322,493,518]" gridcol="0" gridrow="3" pageId="4" pageNumber="141">ANU20210003</th>
<td id="76FD3F79FFE6005DFE62E098FE7DFDFD" box="[419,501,493,518]" gridcol="1" gridrow="3" pageId="4" pageNumber="141">female</td>
<td id="76FD3F79FFE6005DFDF6E098FDEBFDFD" box="[567,611,493,518]" gridcol="2" gridrow="3" pageId="4" pageNumber="141">41.5</td>
<td id="76FD3F79FFE6005DFD57E098FD4AFDFD" box="[662,706,493,518]" gridcol="3" gridrow="3" pageId="4" pageNumber="141">21.4</td>
<td id="76FD3F79FFE6005DFD34E098FC9DFDFD" box="[757,789,493,518]" gridcol="4" gridrow="3" pageId="4" pageNumber="141">4.4</td>
<td id="76FD3F79FFE6005DFC83E098FCECFDFD" box="[834,868,493,518]" gridcol="5" gridrow="3" pageId="4" pageNumber="141">4.8</td>
<td id="76FD3F79FFE6005DFC4EE098FC48FDFD" box="[911,960,493,518]" gridcol="6" gridrow="3" pageId="4" pageNumber="141">4.8</td>
<td id="76FD3F79FFE6005DFC34E098FBB3FDFD" box="[1013,1083,493,518]" gridcol="7" gridrow="3" pageId="4" pageNumber="141">93.6</td>
<td id="76FD3F79FFE6005DFBB9E098FB2CFDFD" box="[1144,1188,493,518]" gridcol="8" gridrow="3" pageId="4" pageNumber="141">73.5</td>
<td id="76FD3F79FFE6005DFB16E098FA8BFDFD" box="[1239,1283,493,518]" gridcol="9" gridrow="3" pageId="4" pageNumber="141">75.7</td>
<td id="76FD3F79FFE6005DFAF6E098FAF8FDFD" box="[1335,1392,493,518]" gridcol="10" gridrow="3" pageId="4" pageNumber="141">144.9</td>
</tr>
<tr id="352C5605FFE6005DFF67E360FAF8FDD5" box="[166,1392,533,558]" gridrow="4" pageId="4" pageNumber="141">
<th id="76FD3F79FFE6005DFF67E360FECAFDD5" box="[166,322,533,558]" gridcol="0" gridrow="4" pageId="4" pageNumber="141">ANU20210004</th>
<td id="76FD3F79FFE6005DFE62E360FE7DFDD5" box="[419,501,533,558]" gridcol="1" gridrow="4" pageId="4" pageNumber="141">juvenile</td>
<td id="76FD3F79FFE6005DFDF6E360FDEBFDD5" box="[567,611,533,558]" gridcol="2" gridrow="4" pageId="4" pageNumber="141">27.8</td>
<td id="76FD3F79FFE6005DFD57E360FD4AFDD5" box="[662,706,533,558]" gridcol="3" gridrow="4" pageId="4" pageNumber="141">14.2</td>
<td id="76FD3F79FFE6005DFD34E360FC9DFDD5" box="[757,789,533,558]" gridcol="4" gridrow="4" pageId="4" pageNumber="141">2.8</td>
<td id="76FD3F79FFE6005DFC83E360FCECFDD5" box="[834,868,533,558]" gridcol="5" gridrow="4" pageId="4" pageNumber="141">2.5</td>
<td id="76FD3F79FFE6005DFC4EE360FC48FDD5" box="[911,960,533,558]" gridcol="6" gridrow="4" pageId="4" pageNumber="141">4.3</td>
<td id="76FD3F79FFE6005DFC34E360FBB3FDD5" box="[1013,1083,533,558]" gridcol="7" gridrow="4" pageId="4" pageNumber="141">46.1</td>
<td id="76FD3F79FFE6005DFBB9E360FB2CFDD5" box="[1144,1188,533,558]" gridcol="8" gridrow="4" pageId="4" pageNumber="141">44.7</td>
<td id="76FD3F79FFE6005DFB16E360FA8BFDD5" box="[1239,1283,533,558]" gridcol="9" gridrow="4" pageId="4" pageNumber="141">39.9</td>
<td id="76FD3F79FFE6005DFAF6E360FAF8FDD5" box="[1335,1392,533,558]" gridcol="10" gridrow="4" pageId="4" pageNumber="141">91.6</td>
</tr>
<tr id="352C5605FFE6005DFF67E34BFAF8FDAD" box="[166,1392,574,598]" gridrow="5" pageId="4" pageNumber="141">
<th id="76FD3F79FFE6005DFF67E34BFECAFDAD" box="[166,322,574,598]" gridcol="0" gridrow="5" pageId="4" pageNumber="141">ANU20210005</th>
<td id="76FD3F79FFE6005DFE62E34BFE7DFDAD" box="[419,501,574,598]" gridcol="1" gridrow="5" pageId="4" pageNumber="141">juvenile</td>
<td id="76FD3F79FFE6005DFDF6E34BFDEBFDAD" box="[567,611,574,598]" gridcol="2" gridrow="5" pageId="4" pageNumber="141">28.4</td>
<td id="76FD3F79FFE6005DFD57E34BFD4AFDAD" box="[662,706,574,598]" gridcol="3" gridrow="5" pageId="4" pageNumber="141">13.2</td>
<td id="76FD3F79FFE6005DFD34E34BFC9DFDAD" box="[757,789,574,598]" gridcol="4" gridrow="5" pageId="4" pageNumber="141">3.0</td>
<td id="76FD3F79FFE6005DFC83E34BFCECFDAD" box="[834,868,574,598]" gridcol="5" gridrow="5" pageId="4" pageNumber="141">2.6</td>
<td id="76FD3F79FFE6005DFC4EE34BFC48FDAD" box="[911,960,574,598]" gridcol="6" gridrow="5" pageId="4" pageNumber="141">4.1</td>
<td id="76FD3F79FFE6005DFC34E34BFBB3FDAD" box="[1013,1083,574,598]" gridcol="7" gridrow="5" pageId="4" pageNumber="141">46.8</td>
<td id="76FD3F79FFE6005DFBB9E34BFB2CFDAD" box="[1144,1188,574,598]" gridcol="8" gridrow="5" pageId="4" pageNumber="141">42.1</td>
<td id="76FD3F79FFE6005DFB16E34BFA8BFDAD" box="[1239,1283,574,598]" gridcol="9" gridrow="5" pageId="4" pageNumber="141">42.5</td>
<td id="76FD3F79FFE6005DFAF6E34BFAF8FDAD" box="[1335,1392,574,598]" gridcol="10" gridrow="5" pageId="4" pageNumber="141">88.2</td>
</tr>
<tr id="352C5605FFE6005DFF67E313FAF8FD85" box="[166,1392,614,638]" gridrow="6" pageId="4" pageNumber="141">
<th id="76FD3F79FFE6005DFF67E313FECAFD85" box="[166,322,614,638]" gridcol="0" gridrow="6" pageId="4" pageNumber="141">CIB116368</th>
<td id="76FD3F79FFE6005DFE62E313FE7DFD85" box="[419,501,614,638]" gridcol="1" gridrow="6" pageId="4" pageNumber="141">juvenile</td>
<td id="76FD3F79FFE6005DFDF6E313FDEBFD85" box="[567,611,614,638]" gridcol="2" gridrow="6" pageId="4" pageNumber="141">34.1</td>
<td id="76FD3F79FFE6005DFD57E313FD4AFD85" box="[662,706,614,638]" gridcol="3" gridrow="6" pageId="4" pageNumber="141">17.3</td>
<td id="76FD3F79FFE6005DFD34E313FC9DFD85" box="[757,789,614,638]" gridcol="4" gridrow="6" pageId="4" pageNumber="141">5.0</td>
<td id="76FD3F79FFE6005DFC83E313FCECFD85" box="[834,868,614,638]" gridcol="5" gridrow="6" pageId="4" pageNumber="141">4.0</td>
<td id="76FD3F79FFE6005DFC4EE313FC48FD85" box="[911,960,614,638]" gridcol="6" gridrow="6" pageId="4" pageNumber="141">5.2</td>
<td id="76FD3F79FFE6005DFC34E313FBB3FD85" box="[1013,1083,614,638]" gridcol="7" gridrow="6" pageId="4" pageNumber="141">63.2</td>
<td id="76FD3F79FFE6005DFBB9E313FB2CFD85" box="[1144,1188,614,638]" gridcol="8" gridrow="6" pageId="4" pageNumber="141">57.4</td>
<td id="76FD3F79FFE6005DFB16E313FA8BFD85" box="[1239,1283,614,638]" gridcol="9" gridrow="6" pageId="4" pageNumber="141">55.8</td>
<td id="76FD3F79FFE6005DFAF6E313FAF8FD85" box="[1335,1392,614,638]" gridcol="10" gridrow="6" pageId="4" pageNumber="141">114.7</td>
</tr>
<tr id="352C5605FFE6005DFF67E3FBFAF8FD5D" box="[166,1392,654,678]" gridrow="7" pageId="4" pageNumber="141">
<th id="76FD3F79FFE6005DFF67E3FBFECAFD5D" box="[166,322,654,678]" gridcol="0" gridrow="7" pageId="4" pageNumber="141">SNHM5414</th>
<td id="76FD3F79FFE6005DFE62E3FBFE7DFD5D" box="[419,501,654,678]" gridcol="1" gridrow="7" pageId="4" pageNumber="141">juvenile</td>
<td id="76FD3F79FFE6005DFDF6E3FBFDEBFD5D" box="[567,611,654,678]" gridcol="2" gridrow="7" pageId="4" pageNumber="141">32.2</td>
<td id="76FD3F79FFE6005DFD57E3FBFD4AFD5D" box="[662,706,654,678]" gridcol="3" gridrow="7" pageId="4" pageNumber="141">16.7</td>
<td id="76FD3F79FFE6005DFD34E3FBFC9DFD5D" box="[757,789,654,678]" gridcol="4" gridrow="7" pageId="4" pageNumber="141">4.3</td>
<td id="76FD3F79FFE6005DFC83E3FBFCECFD5D" box="[834,868,654,678]" gridcol="5" gridrow="7" pageId="4" pageNumber="141">3.6</td>
<td id="76FD3F79FFE6005DFC4EE3FBFC48FD5D" box="[911,960,654,678]" gridcol="6" gridrow="7" pageId="4" pageNumber="141">4.9</td>
<td id="76FD3F79FFE6005DFC34E3FBFBB3FD5D" box="[1013,1083,654,678]" gridcol="7" gridrow="7" pageId="4" pageNumber="141">60.6</td>
<td id="76FD3F79FFE6005DFBB9E3FBFB2CFD5D" box="[1144,1188,654,678]" gridcol="8" gridrow="7" pageId="4" pageNumber="141">57.3</td>
<td id="76FD3F79FFE6005DFB16E3FBFA8BFD5D" box="[1239,1283,654,678]" gridcol="9" gridrow="7" pageId="4" pageNumber="141">52.7</td>
<td id="76FD3F79FFE6005DFAF6E3FBFAF8FD5D" box="[1335,1392,654,678]" gridcol="10" gridrow="7" pageId="4" pageNumber="141">113.0</td>
</tr>
</table>
</paragraph>
<caption id="DF6304CFFFE6FFA6FF56E4E6FAF5FA50" ID-DOI="http://doi.org/10.5281/zenodo.5606856" ID-Zenodo-Dep="5606856" box="[151,1405,1427,1451]" httpUri="https://zenodo.org/record/5606856/files/figure.png" pageId="4" pageNumber="141" startId="4.[151,250,1427,1451]" targetBox="[163,1424,756,1393]" targetPageId="4">
<paragraph id="8BA35447FFE6FFA6FF56E4E6FAF5FA50" blockId="4.[151,1405,1427,1451]" box="[151,1405,1427,1451]" pageId="4" pageNumber="141">
<emphasis id="B9688855FFE6FFA6FF56E4E6FE9BFA50" bold="true" box="[151,275,1427,1451]" pageId="4" pageNumber="141">FIGURE 2.</emphasis>
Dorsal and ventral aspects of the holotype of
<taxonomicName id="4C1C2FC4FFE6FFA6FD24E4E6FC74FA50" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[741,1020,1427,1451]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="141" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE6FFA6FD24E4E6FC74FA50" box="[741,1020,1427,1451]" italics="true" pageId="4" pageNumber="141">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE6FFA6FBC3E4E6FBDAFA50" bold="true" box="[1026,1106,1427,1451]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="A25B352EFFE6FFA6FBC3E4E6FBDAFA50" box="[1026,1106,1427,1451]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ANU20210001, adult male)
</paragraph>
</caption>
<paragraph id="8BA35447FFE6FFA6FF06E4ACFBFFF97F" blockId="4.[151,1437,1497,2028]" pageId="4" pageNumber="141">
<emphasis id="B9688855FFE6FFA6FF06E4ACFE06FA0F" bold="true" box="[199,398,1497,1524]" pageId="4" pageNumber="141">Ecological notes.</emphasis>
On
<date id="FFA27287FFE6FFA6FE7FE4AFFDD6FA0F" box="[446,606,1498,1524]" pageId="4" pageNumber="141" value="2017-06-20">June 20, 2017</date>
, a female individual (raised in Xiuning Institute of Rare Aquatic Animals) laid
<specimenCount id="9D1A9FCEFFE6FFA6FF0AE488FEBBF9E3" box="[203,307,1533,1560]" pageId="4" pageNumber="141" type="egg">five eggs</specimenCount>
with a postpartum weight of
<quantity id="4CE4F9A2FFE6FFA6FD4FE48BFD5BF9E3" box="[654,723,1534,1560]" metricMagnitude="-1" metricUnit="kg" metricValue="1.34" pageId="4" pageNumber="141" unit="g" value="134.0">134 g</quantity>
. The eggs are white, nearly spherical; average length 15.95 ± 0.11 (15.7916.01) mm; average short diameter 15.6 ± 0.16 (15.5615.89) mm; and average weight 2.12 ± 0.23 (1.842.4) g. Hatching occurs at natural temperatures after 6065 days. The average weight of neonates was 1.30 ± 0.08 (1.201.40) g. Three months old horse-hoof soft-shelled turtles are shown in
<figureCitation id="132748C2FFE6FFA6FBEFE71CFBF9F97F" box="[1070,1137,1641,1668]" captionStart="FIGURE 5" captionStartId="6.[151,250,689,713]" captionTargetBox="[152,1436,181,665]" captionTargetId="figure-23@6.[151,1436,181,665]" captionTargetPageId="6" captionText="FIGURE 5. The three months old Huangshan soft-shelled turtles." figureDoi="http://doi.org/10.5281/zenodo.5606866" httpUri="https://zenodo.org/record/5606866/files/figure.png" pageId="4" pageNumber="141">Fig. 5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30607CCFFE6FFA6FF06E7FBFAD6F90B" pageId="4" pageNumber="141" type="etymology">
<paragraph id="8BA35447FFE6FFA6FF06E7FBFAD6F90B" blockId="4.[151,1437,1497,2028]" pageId="4" pageNumber="141">
<emphasis id="B9688855FFE6FFA6FF06E7FBFEC2F953" bold="true" box="[199,330,1678,1704]" pageId="4" pageNumber="141">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="54A7EAE5FFE6FFA6FD63E7FBFD59F953" box="[674,721,1678,1704]" pageId="4" pageNumber="141">type</typeStatus>
locality of this species, Huangshan City,
<collectingRegion id="49D89AA5FFE6FFA6FB4CE7F8FAB3F95C" box="[1165,1339,1677,1703]" country="China" name="Anhui" pageId="4" pageNumber="141">Anhui Province</collectingRegion>
. We suggest the following common names: horse-hoof soft-shelled turtle in English and “Huáng Shān Mǎ tí Biē (
<emphasis id="B9688855FFE6FFA6FB3EE7C7FA0FF932" bold="true" box="[1279,1415,1714,1737]" pageId="4" pageNumber="141">•山ƼÞ×</emphasis>
)” in Chinese. The native people of Huangshan have called it “Mǎ tí Biē (
<emphasis id="B9688855FFE6FFA6FC58E7A3FC63F916" bold="true" box="[921,1003,1750,1773]" pageId="4" pageNumber="141">ƼÞ×</emphasis>
)” for more than a thousand years.
</paragraph>
</subSubSection>
<subSubSection id="C30607CCFFE6FFA6FF06E78CFC11F817" pageId="4" pageNumber="141" type="distribution">
<paragraph id="8BA35447FFE6FFA6FF06E78CFE28F8CC" blockId="4.[151,1437,1497,2028]" pageId="4" pageNumber="141">
<emphasis id="B9688855FFE6FFA6FF06E78CFEE9F8E8" bold="true" box="[199,353,1785,1811]" pageId="4" pageNumber="141">Distribution.</emphasis>
This species is currently known only from the
<typeStatus id="54A7EAE5FFE6FFA6FC41E78FFC38F8EF" box="[896,944,1786,1812]" pageId="4" pageNumber="141">type</typeStatus>
locality, Xiuning County, Huangshan City,
<collectingRegion id="49D89AA5FFE6FFA6FF56E668FEC2F8CC" box="[151,330,1821,1847]" country="China" name="Anhui" pageId="4" pageNumber="141">Anhui Province</collectingRegion>
,
<collectingCountry id="F30B14D7FFE6FFA6FE97E668FE15F8CC" box="[342,413,1821,1847]" name="China" pageId="4" pageNumber="141">China</collectingCountry>
.
</paragraph>
<paragraph id="8BA35447FFE6FFA6FF06E634FC11F817" blockId="4.[151,1437,1497,2028]" pageId="4" pageNumber="141">
<emphasis id="B9688855FFE6FFA6FF06E634FEE5F8A0" bold="true" box="[199,365,1857,1883]" pageId="4" pageNumber="141">Comparisons.</emphasis>
<taxonomicName id="4C1C2FC4FFE6FFA6FEB6E634FD25F8A0" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[375,685,1857,1883]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="141" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE6FFA6FEB6E634FD25F8A0" box="[375,685,1857,1883]" italics="true" pageId="4" pageNumber="141">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE6FFA6FD76E637FC9BF8A7" bold="true" box="[695,787,1858,1884]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="A25B352EFFE6FFA6FD76E637FC9BF8A7" box="[695,787,1858,1884]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C1C2FC4FFE6FFA6FB93E637FB5CF8A0" box="[1106,1236,1858,1883]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="141" phylum="Chordata" rank="species" species="axenaria">
<emphasis id="B9688855FFE6FFA6FB93E637FB5CF8A0" box="[1106,1236,1858,1883]" italics="true" pageId="4" pageNumber="141">P. axenaria</emphasis>
</taxonomicName>
by the following morphological characters: (1) Many tubercles on the dorsal surface, but indistinct in the center (vs. smooth carapace, no tubercles); (2) plastron yellowish-white, and only a dark patch on each side of the armpit (vs. a single dark-gray central figure enclosed by the hypoplastra and xiphiplastra); and (3) white longitudinal bands on both sides of the neck in juveniles, absent in adults (vs. absence of spots on the neck).
</paragraph>
</subSubSection>
<caption id="DF6304CFFFE7FFA7FF56E54DFA11FBAB" ID-DOI="http://doi.org/10.5281/zenodo.5606860" ID-Zenodo-Dep="5606860" box="[151,1433,1080,1104]" httpUri="https://zenodo.org/record/5606860/files/figure.png" pageId="5" pageNumber="142" startId="5.[151,250,1080,1104]" targetBox="[165,1420,196,1044]" targetPageId="5">
<paragraph id="8BA35447FFE7FFA7FF56E54DFA11FBAB" blockId="5.[151,1433,1080,1104]" box="[151,1433,1080,1104]" pageId="5" pageNumber="142">
<emphasis id="B9688855FFE7FFA7FF56E54DFE9BFBAB" bold="true" box="[151,275,1080,1104]" pageId="5" pageNumber="142">FIGURE 3.</emphasis>
Dorsal and ventral aspects of the paratypes of
<taxonomicName id="4C1C2FC4FFE7FFA7FD2FE54DFB8CFBAB" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[750,1028,1080,1104]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="142" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE7FFA7FD2FE54DFB8CFBAB" box="[750,1028,1080,1104]" italics="true" pageId="5" pageNumber="142">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE7FFA7FBCBE54DFBD3FBAB" bold="true" box="[1034,1115,1080,1104]" pageId="5" pageNumber="142">
<taxonomicNameLabel id="A25B352EFFE7FFA7FBCBE54DFBD3FBAB" box="[1034,1115,1080,1104]" pageId="5" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ANU20210002, adult female)
</paragraph>
</caption>
<caption id="DF6304CFFFE7FFA7FF56E67AFA15F8D3" ID-DOI="http://doi.org/10.5281/zenodo.5606864" ID-Zenodo-Dep="5606864" box="[151,1437,1807,1832]" httpUri="https://zenodo.org/record/5606864/files/figure.png" pageId="5" pageNumber="142" startId="5.[151,250,1807,1831]" targetBox="[151,1436,1128,1784]" targetPageId="5">
<paragraph id="8BA35447FFE7FFA7FF56E67AFA15F8D3" blockId="5.[151,1437,1807,1832]" box="[151,1437,1807,1832]" pageId="5" pageNumber="142">
<emphasis id="B9688855FFE7FFA7FF56E67AFE9BF8D3" bold="true" box="[151,275,1807,1832]" pageId="5" pageNumber="142">FIGURE 4.</emphasis>
Head and neck of
<taxonomicName id="4C1C2FC4FFE7FFA7FE10E665FD60F8D3" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[465,744,1808,1832]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="142" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE7FFA7FE10E665FD60F8D3" box="[465,744,1808,1832]" italics="true" pageId="5" pageNumber="142">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE7FFA7FD2FE665FCB6F8D3" bold="true" box="[750,830,1808,1832]" pageId="5" pageNumber="142">
<taxonomicNameLabel id="A25B352EFFE7FFA7FD2FE665FCB6F8D3" box="[750,830,1808,1832]" pageId="5" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; A, B ANU20210001, male; C, D ANU20210004, juvenile.
</paragraph>
</caption>
<caption id="DF6304CFFFE4FFA4FF56E3C4FCBBFD32" ID-DOI="http://doi.org/10.5281/zenodo.5606866" ID-Zenodo-Dep="5606866" box="[151,819,689,713]" httpUri="https://zenodo.org/record/5606866/files/figure.png" pageId="6" pageNumber="143" startId="6.[151,250,689,713]" targetBox="[152,1436,181,665]" targetPageId="6">
<paragraph id="8BA35447FFE4FFA4FF56E3C4FCBBFD32" blockId="6.[151,819,689,713]" box="[151,819,689,713]" pageId="6" pageNumber="143">
<emphasis id="B9688855FFE4FFA4FF56E3C4FE9BFD32" bold="true" box="[151,275,689,713]" pageId="6" pageNumber="143">FIGURE 5.</emphasis>
The three months old Huangshan soft-shelled turtles.
</paragraph>
</caption>
<subSubSection id="C30607CCFFE4FFA4FF06E382FE12F9B5" pageId="6" pageNumber="143" type="discussion">
<paragraph id="8BA35447FFE4FFA4FF06E382FC79FC3D" blockId="6.[151,1436,759,1614]" pageId="6" pageNumber="143">
<taxonomicName id="4C1C2FC4FFE4FFA4FF06E382FE72FCEA" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[199,506,759,785]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE4FFA4FF06E382FE72FCEA" box="[199,506,759,785]" italics="true" pageId="6" pageNumber="143">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE4FFA4FDC1E38DFDD1FCE9" bold="true" box="[512,601,760,786]" pageId="6" pageNumber="143">
<taxonomicNameLabel id="A25B352EFFE4FFA4FDC1E38DFDD1FCE9" box="[512,601,760,786]" pageId="6" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C1C2FC4FFE4FFA4FC19E38DFBC4FCEA" baseAuthorityName="Stuckas &amp; Fritz" baseAuthorityYear="2011" box="[984,1100,760,785]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="B9688855FFE4FFA4FC19E38DFBC4FCEA" box="[984,1100,760,785]" italics="true" pageId="6" pageNumber="143">P. sinensis</emphasis>
</taxonomicName>
by the following morphological characters: (1) Small size; maximum carapace length
<quantity id="4CE4F9A2FFE4FFA4FCE4E26EFC1FFCCD" box="[805,919,795,822]" metricMagnitude="-1" metricUnit="m" metricValue="1.156" pageId="6" pageNumber="143" unit="mm" value="115.6">115.6 mm</quantity>
(vs. medium size; maximum carapace length
<quantity id="4CE4F9A2FFE4FFA4FF56E235FF72FCA1" box="[151,250,832,858]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="143" unit="mm" value="230.0">230 mm</quantity>
); (2) plastron yellowish-white, and only a dark patch on each side of the armpit (vs. plastron yellowishwhite with many dark spots); (3) keel high (vs. keel low); (4) no black pinstripes around the eyes (vs. black pinstripes around the eyes); (5) the throat has many tiny yellowish-white spots (vs. large yellowish-white spots with dark edges); and (6) entoplastron “⌒” shaped (vs. entoplastron “˄” shaped).
</paragraph>
<paragraph id="8BA35447FFE4FFA4FF06E2BAFC02FB65" blockId="6.[151,1436,759,1614]" pageId="6" pageNumber="143">
<taxonomicName id="4C1C2FC4FFE4FFA4FF06E2BAFE72FC12" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[199,506,975,1001]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE4FFA4FF06E2BAFE72FC12" box="[199,506,975,1001]" italics="true" pageId="6" pageNumber="143">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE4FFA4FDC1E2A5FDD1FC11" bold="true" box="[512,601,976,1002]" pageId="6" pageNumber="143">
<taxonomicNameLabel id="A25B352EFFE4FFA4FDC1E2A5FDD1FC11" box="[512,601,976,1002]" pageId="6" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C1C2FC4FFE4FFA4FC19E2A5FBC4FC12" baseAuthorityName="Brandt" baseAuthorityYear="1857" box="[984,1100,975,1001]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="maackii">
<emphasis id="B9688855FFE4FFA4FC19E2A5FBC4FC12" box="[984,1100,975,1001]" italics="true" pageId="6" pageNumber="143">P. maackii</emphasis>
</taxonomicName>
by the following morphological characters: (1) Small size; maximum carapace length of
<quantity id="4CE4F9A2FFE4FFA4FC83E286FC3CFBF5" box="[834,948,1011,1038]" metricMagnitude="-1" metricUnit="m" metricValue="1.156" pageId="6" pageNumber="143" unit="mm" value="115.6">115.6 mm</quantity>
(vs. larger size; maximum carapace length more than
<quantity id="4CE4F9A2FFE4FFA4FED3E562FEFDFBC9" box="[274,373,1047,1074]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="143" unit="mm" value="350.0">350 mm</quantity>
); (2) plastron yellowish-white, and only a dark patch on each side of the armpit (vs. plastron yellowish-white with no markings or relatively small, faint round to oval dark markings); (3) the throat has many tiny yellowish-white spots (vs. large yellowish-white spots with dark edges); (4) keel high (vs. keel low); and (5) no black stripes around the eyes (vs. black pinstripes around the eyes).
</paragraph>
<paragraph id="8BA35447FFE4FFA4FF06E5D2FCE8FAA9" blockId="6.[151,1436,759,1614]" pageId="6" pageNumber="143">
<taxonomicName id="4C1C2FC4FFE4FFA4FF06E5D2FE75FB3A" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[199,509,1191,1217]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE4FFA4FF06E5D2FE75FB3A" box="[199,509,1191,1217]" italics="true" pageId="6" pageNumber="143">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE4FFA4FDC7E5DDFDEAFB39" bold="true" box="[518,610,1192,1218]" pageId="6" pageNumber="143">
<taxonomicNameLabel id="A25B352EFFE4FFA4FDC7E5DDFDEAFB39" box="[518,610,1192,1218]" pageId="6" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C1C2FC4FFE4FFA4FC34E5DDFB1EFB3A" authorityName="Tang" authorityYear="1997" box="[1013,1174,1191,1217]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="parviformis">
<emphasis id="B9688855FFE4FFA4FC34E5DDFB1EFB3A" box="[1013,1174,1191,1217]" italics="true" pageId="6" pageNumber="143">P. parviformis</emphasis>
</taxonomicName>
by the following morphological characters: (1) Plastron yellowish-white, and only a dark patch on each side of the armpit (vs. plastron yellowish-white with no markings); (2) many tubercles on the dorsal surface, but indistinct in the center (vs. dorsal tubercles in longitudinal series more or less discrete; central tubercle in front of marginal ridge of carapace small); and (3) entoplastron “⌒” shaped (vs. entoplastron “˄” shaped).
</paragraph>
<paragraph id="8BA35447FFE4FFA4FF06E42EFE12F9B5" blockId="6.[151,1436,759,1614]" pageId="6" pageNumber="143">
<taxonomicName id="4C1C2FC4FFE4FFA4FF06E42EFE73FA8E" authority="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie, 2021" authorityName="Gong &amp; Peng &amp; Huang &amp; Lin &amp; Huang &amp; Xu &amp; Yang &amp; Nie" authorityYear="2021" box="[199,507,1371,1397]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="huangshanensis" status="sp. nov.">
<emphasis id="B9688855FFE4FFA4FF06E42EFE73FA8E" box="[199,507,1371,1397]" italics="true" pageId="6" pageNumber="143">Pelodiscus huangshanensis</emphasis>
</taxonomicName>
<emphasis id="B9688855FFE4FFA4FDC3E429FDD4FA8D" bold="true" box="[514,604,1372,1398]" pageId="6" pageNumber="143">
<taxonomicNameLabel id="A25B352EFFE4FFA4FDC3E429FDD4FA8D" box="[514,604,1372,1398]" pageId="6" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C1C2FC4FFE4FFA4FC20E429FBFBFA8E" authorityName="Farkas, Ziegler, Pham, Ong &amp; Fritz" authorityYear="2019" box="[993,1139,1372,1397]" class="Reptilia" family="Trionychidae" genus="Pelodiscus" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="variegatus">
<emphasis id="B9688855FFE4FFA4FC20E429FBFBFA8E" box="[993,1139,1372,1397]" italics="true" pageId="6" pageNumber="143">P. variegatus</emphasis>
</taxonomicName>
by the following morphological characters: (1) Many tubercles on the dorsal surface, but indistinct in the center (vs. dorsal surface roughened by longitudinal ridging and smaller protuberances spread over the leathery margin); (2) plastron yellowish-white, and only a dark patch on each side of the armpit (vs. plastron pinkish white to pale reddish orange distinct, large dark blotches, underside of leathery margin of carapace pigmented); (3) no black pinstripes around the eyes (vs. black pinstripes around the eyes); and (4) the throat has many tiny yellowish-white spots (vs. large yellowish-white spots with dark edges).
</paragraph>
</subSubSection>
</treatment>
</document>