226 lines
42 KiB
XML
226 lines
42 KiB
XML
<document id="77B561174A1C8E350C20A4CF45F6CBAC" ID-CLB-Dataset="26155" ID-DOI="10.11646/zootaxa.4646.1.11" ID-GBIF-Dataset="64b27e40-f6d9-403f-8208-2c5058465826" ID-ISSN="1175-5326" ID-Zenodo-Dep="3346299" ID-ZooBank="EBF7F6A3-D6C8-4422-A1A6-6612BAA9ACD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563862654245" checkinUser="plazi" docAuthor="Draney, Michael L., Milne, Marc A., Ulyshen, Michael & Madriz, Gabrielle" docDate="2019" docId="03B587E7735E1870FF23FC75FD9BFE7F" docLanguage="en" docName="zootaxa.4646.1.11.pdf" docOrigin="Zootaxa 4646 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epiceraticelus Crosby & Bishop 1931" docType="treatment" docVersion="10" lastPageNumber="192" masterDocId="FF8CFF9F735C1873FFB4FFA0FFEAFFD0" masterDocTitle="Revision of the spider genus Epiceraticelus (Araneae, Linyphiidae) with a description of a new species" masterLastPageNumber="200" masterPageNumber="189" pageNumber="191" updateTime="1698746982808" updateUser="ExternalLinkService">
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<mods:title id="F6500B5CC9748A6930FAA2A946315DB7">Revision of the spider genus Epiceraticelus (Araneae, Linyphiidae) with a description of a new species</mods:title>
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<mods:namePart id="8435BA1D01007EC1B51B392CA7B25350">Draney, Michael L.</mods:namePart>
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<mods:namePart id="486B06E2664A8810518AE46BEEBFEEDD">Milne, Marc A.</mods:namePart>
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<mods:namePart id="0E3B4EF512D55EDEAE0A930FC97ED6F7">Ulyshen, Michael</mods:namePart>
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<mods:namePart id="341BBCF2AB938C74A4A60802D904E7AA">Madriz, Gabrielle</mods:namePart>
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<mods:title id="22C45AD43E958846D069BBA57FA483F9">Zootaxa</mods:title>
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<treatment id="03B587E7735E1870FF23FC75FD9BFE7F" ID-DOI="http://doi.org/10.5281/zenodo.5922994" ID-GBIF-Taxon="158849545" ID-Zenodo-Dep="5922994" LSID="urn:lsid:plazi:treatment:03B587E7735E1870FF23FC75FD9BFE7F" httpUri="http://treatment.plazi.org/id/03B587E7735E1870FF23FC75FD9BFE7F" lastPageId="3" lastPageNumber="192" pageId="2" pageNumber="191">
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<subSubSection id="C306657A735E1871FF23FC75FD87FC3F" box="[151,621,981,1008]" pageId="2" pageNumber="191" type="nomenclature">
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<paragraph id="8BA336F1735E1871FF23FC75FD87FC3F" blockId="2.[151,621,981,1008]" box="[151,621,981,1008]" pageId="2" pageNumber="191">
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<heading id="D0EB819D735E1871FF23FC75FD87FC3F" bold="true" box="[151,621,981,1008]" fontSize="11" level="1" pageId="2" pageNumber="191" reason="1">
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<taxonomicName id="4C1C4D72735E1871FF23FC75FD87FC3F" ID-CoL="4CXX" authority="Crosby & Bishop, 1931" authorityName="Crosby & Bishop" authorityYear="1931" box="[151,621,981,1008]" class="Arachnida" family="Linyphiidae" genus="Epiceraticelus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAE3735E1871FF23FC75FD87FC3F" bold="true" box="[151,621,981,1008]" pageId="2" pageNumber="191">
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<emphasis id="B968EAE3735E1871FF23FC75FEAEFC20" bold="true" box="[151,324,981,1008]" italics="true" pageId="2" pageNumber="191">Epiceraticelus</emphasis>
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<bibRefCitation id="EF8D4B00735E1871FEFFFC75FD87FC3F" author="Crosby, C. R. & Bishop, S. C." box="[331,621,981,1007]" pageId="2" pageNumber="191" pagination="359 - 403" refId="ref7304" refString="Crosby, C. R. & Bishop, S. C. (1931) Studies in American spiders: genera, Cornicularia, Paracornicularia, Tigellinus, Walckenaera, Epiceraticelus, and Pelecopsis, with descriptions of new genera and species. Journal of the New York Entomological Society, 39 (3), 359 - 403." type="journal article" year="1931">Crosby & Bishop, 1931</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306657A735E1871FF23FBB9FDB1FBE1" box="[151,603,1048,1073]" pageId="2" pageNumber="191" type="reference_group">
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<paragraph id="8BA336F1735E1871FF23FBB9FDB1FBE1" blockId="2.[151,942,1048,1104]" box="[151,603,1048,1073]" pageId="2" pageNumber="191">
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<treatmentCitationGroup id="AB0C11DF735E1871FF23FBB9FDB1FBE1" box="[151,603,1048,1073]" pageId="2" pageNumber="191">
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<taxonomicName id="4C1C4D72735E1871FF23FBB9FDBDFBE1" ID-CoL="4CXX" authority="Crosby & Bishop, 1931: 380" authorityName="Crosby & Bishop" authorityPageNumber="380" authorityYear="1931" box="[151,599,1048,1073]" class="Arachnida" family="Linyphiidae" genus="Epiceraticelus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAE3735E1871FF23FBB9FEC3FBE1" box="[151,297,1049,1073]" italics="true" pageId="2" pageNumber="191">Epiceraticelus</emphasis>
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<treatmentCitation id="0ABD10E0735E1871FE84FBB8FDBDFBE1" author="Crosby, C. R. & Bishop, S. C." box="[304,599,1048,1073]" page="380" pageId="2" pageNumber="191" year="1931">
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<bibRefCitation id="EF8D4B00735E1871FE84FBB8FDBDFBE1" author="Crosby, C. R. & Bishop, S. C." box="[304,599,1048,1073]" pageId="2" pageNumber="191" pagination="359 - 403" refId="ref7304" refString="Crosby, C. R. & Bishop, S. C. (1931) Studies in American spiders: genera, Cornicularia, Paracornicularia, Tigellinus, Walckenaera, Epiceraticelus, and Pelecopsis, with descriptions of new genera and species. Journal of the New York Entomological Society, 39 (3), 359 - 403." type="journal article" year="1931">Crosby & Bishop, 1931: 380</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306657A735E1870FF23FB98FD9BFE7F" lastPageId="3" lastPageNumber="192" pageId="2" pageNumber="191" type="materials_examined">
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<paragraph id="8BA336F1735E1871FF23FB98FC44FB80" blockId="2.[151,942,1048,1104]" box="[151,942,1079,1104]" pageId="2" pageNumber="191">
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<emphasis id="B968EAE3735E1871FF23FB98FEC9FB80" bold="true" box="[151,291,1079,1104]" pageId="2" pageNumber="191">
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<typeStatus id="54A78853735E1871FF23FB98FF26FB80" box="[151,204,1080,1104]" pageId="2" pageNumber="191">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C1C4D72735E1871FE9EFB98FCF2FB80" authority="Crosby and Bishop 1931" authorityName="Crosby and Bishop" authorityYear="1931" box="[298,792,1079,1104]" class="Arachnida" family="Linyphiidae" genus="Epiceraticelus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="fluvialis">
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<emphasis id="B968EAE3735E1871FE9EFB98FDFEFB80" box="[298,532,1080,1104]" italics="true" pageId="2" pageNumber="191">Epiceraticelus fluvialis</emphasis>
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<bibRefCitation id="EF8D4B00735E1871FDAEFB97FCF2FB80" author="Crosby, C. R. & Bishop, S. C." box="[538,792,1079,1104]" pageId="2" pageNumber="191" pagination="359 - 403" refId="ref7304" refString="Crosby, C. R. & Bishop, S. C. (1931) Studies in American spiders: genera, Cornicularia, Paracornicularia, Tigellinus, Walckenaera, Epiceraticelus, and Pelecopsis, with descriptions of new genera and species. Journal of the New York Entomological Society, 39 (3), 359 - 403." type="journal article" year="1931">Crosby and Bishop 1931</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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<paragraph id="8BA336F1735E1871FF23FBDDFD69F9D0" blockId="2.[151,1437,1149,2041]" pageId="2" pageNumber="191">
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<emphasis id="B968EAE3735E1871FF23FBDDFEF8FB48" bold="true" box="[151,274,1149,1176]" pageId="2" pageNumber="191">Diagnosis.</emphasis>
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Males may be separated from similar North American genera (e.g.
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<taxonomicName id="4C1C4D72735E1871FBA5FBDEFB78FB48" authorityName="Simon" authorityYear="1884" box="[1041,1170,1150,1176]" class="Arachnida" family="Linyphiidae" genus="Ceraticelus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAE3735E1871FBA5FBDEFB78FB48" box="[1041,1170,1150,1176]" italics="true" pageId="2" pageNumber="191">Ceraticelus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1C4D72735E1871FB15FBDEFACBFB48" authorityName="Emerton" authorityYear="1882" box="[1185,1313,1150,1176]" class="Arachnida" family="Linyphiidae" genus="Ceratinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAE3735E1871FB15FBDEFACBFB48" box="[1185,1313,1150,1176]" italics="true" pageId="2" pageNumber="191">Ceratinella</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1C4D72735E1871FA9BFBDFFF23FB6C" authorityName="Emerton" authorityYear="1882" class="Arachnida" family="Linyphiidae" genus="Grammonota" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAE3735E1871FA9BFBDFFF23FB6C" italics="true" pageId="2" pageNumber="191">Grammonota</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1C4D72735E1871FF62FB02FEAEFB6C" authorityName="Bishop & Crosby" authorityYear="1938" box="[214,324,1186,1212]" class="Arachnida" family="Linyphiidae" genus="Scylaceus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAE3735E1871FF62FB02FEAEFB6C" box="[214,324,1186,1212]" italics="true" pageId="2" pageNumber="191">Scylaceus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C1C4D72735E1871FE34FB02FDF3FB6C" authorityName="Blackwall" authorityYear="1833" box="[384,537,1186,1212]" class="Arachnida" family="Linyphiidae" genus="Walckenaeria" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAE3735E1871FE34FB02FDF3FB6C" box="[384,537,1186,1212]" italics="true" pageId="2" pageNumber="191">Walckenaeria</emphasis>
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</taxonomicName>
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) by possessing a large palpal tibial apopysis that is serrated on the medial margin (
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<figureCitation id="13272A74735E1871FF2BFB66FEEFFB30" box="[159,261,1222,1248]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1891]" captionTargetBox="[151,1434,211,1836]" captionTargetId="figure@5.[151,1434,211,1836]" captionTargetPageId="5" captionText="FIGURE 2. SEM micrographs of Epiceraticelus fluvialis: A, Retrolateral view of tibial apophysis of left palp of male. B, Ventral view of male left palp. C, Dorsal view of epigynum, cleared. D, Ventral view of epigynum. Abbreviations: CD, copulatory ducts; CG, copulatory grooves; CO, copulatory openings; E, embolus; EA, embolic apophysis; FD, fertilization ducts; MP, median plate; PC, paracymbium; S, spermathecae; ST, subtegulum; T, tegulum; TP, tailpiece. Scale bars, A, B, 0.1 mm; C, D, 0.05 mm." httpUri="https://zenodo.org/record/3346303/files/figure.png" pageId="2" pageNumber="191">Figs. 2A</figureCitation>
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,
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<figureCitation id="13272A74735E1871FEA6FB66FEDFFB30" box="[274,309,1222,1248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1245,1270]" captionTargetBox="[156,1407,186,1217]" captionTargetId="figure@6.[151,1436,181,1226]" captionTargetPageId="6" captionText="FIGURE 3. Free hand Illustrations of Epiceraticelus fluvialis (proporations are approximate): A. Male left palp, ventral view. B, Male left palp, retrolateral view. C, Retrolateral view of tibial apophysis of male left palp. D, Lateral view of male carapace; E, Ventral view of epigynum. F, Dorsal schematic of epigynum. Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece. Proportions are approximate." httpUri="https://zenodo.org/record/3346305/files/figure.png" pageId="2" pageNumber="191">3C</figureCitation>
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,
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<figureCitation id="13272A74735E1871FEF7FB66FE8CFB30" box="[323,358,1222,1248]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1876]" captionTargetBox="[151,1436,191,1818]" captionTargetId="figure@8.[151,1436,191,1818]" captionTargetPageId="8" captionText="FIGURE 4. SEM micrographs of Epiceraticelus mandyae n. sp.: A, Male habitus. B, Ventral view of male left palp. C, Retrolateral view of tibial apophysis of male left palp. D, Dorsal view of epigynum, cleared. E, Ventral view of epigynum. Abbreviations: E, embolus; EA, embolic apophysis; EM, embolic membrane; PC, paracymbium; RTA, retrolateral tibial apophysis; T, tegulum; TP, tailpiece; S, spermathecae; CD, copulatory duct; CO, copulatory opening; MP, median plate. Scale bars, A, 0.5 mm; B, 0.1 mm; C, D, E, 0.05 mm." httpUri="https://zenodo.org/record/3346307/files/figure.png" pageId="2" pageNumber="191">4C</figureCitation>
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,
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<figureCitation id="13272A74735E1871FEC7FB66FE7DFB31" box="[371,407,1222,1249]" captionStart="FIGURE 6" captionStartId="10.[151,250,1118,1143]" captionTargetBox="[170,1436,195,1093]" captionTargetId="figure@10.[151,1436,181,1099]" captionTargetPageId="10" captionText="FIGURE 6. Free hand llustrations of Epiceraticelus mandyae n. sp. (proportions are approximate). A, Male left palp, ventral view. B. Male left palp,, retrolateral view. C, Retrolateral view of tibial apophysis of left palp of male. D, Ventral view of epigynum. E,Dorsal schematic of epigynum. F, Lateral view of male carapace.Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; EM, embolic membrane; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece." httpUri="https://zenodo.org/record/3346311/files/figure.png" pageId="2" pageNumber="191">6C</figureCitation>
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). Females may be separated from other erigonines by having an epigynum split in half by two parallel copulatory grooves with copulatory openings positioned anteriorly and a widened rectangular median plate posteriorly (
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<figureCitation id="13272A74735E1871FEEFFAAEFE57FAF8" box="[347,445,1294,1320]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1891]" captionTargetBox="[151,1434,211,1836]" captionTargetId="figure@5.[151,1434,211,1836]" captionTargetPageId="5" captionText="FIGURE 2. SEM micrographs of Epiceraticelus fluvialis: A, Retrolateral view of tibial apophysis of left palp of male. B, Ventral view of male left palp. C, Dorsal view of epigynum, cleared. D, Ventral view of epigynum. Abbreviations: CD, copulatory ducts; CG, copulatory grooves; CO, copulatory openings; E, embolus; EA, embolic apophysis; FD, fertilization ducts; MP, median plate; PC, paracymbium; S, spermathecae; ST, subtegulum; T, tegulum; TP, tailpiece. Scale bars, A, B, 0.1 mm; C, D, 0.05 mm." httpUri="https://zenodo.org/record/3346303/files/figure.png" pageId="2" pageNumber="191">Figs. 2D</figureCitation>
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,
|
||
<figureCitation id="13272A74735E1871FE7CFAAEFE03FAF8" box="[456,489,1294,1320]" captionStart="FIGURE 3" captionStartId="6.[151,250,1245,1270]" captionTargetBox="[156,1407,186,1217]" captionTargetId="figure@6.[151,1436,181,1226]" captionTargetPageId="6" captionText="FIGURE 3. Free hand Illustrations of Epiceraticelus fluvialis (proporations are approximate): A. Male left palp, ventral view. B, Male left palp, retrolateral view. C, Retrolateral view of tibial apophysis of male left palp. D, Lateral view of male carapace; E, Ventral view of epigynum. F, Dorsal schematic of epigynum. Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece. Proportions are approximate." httpUri="https://zenodo.org/record/3346305/files/figure.png" pageId="2" pageNumber="191">3E</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FE47FAAEFDFFFAF8" box="[499,533,1294,1320]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1876]" captionTargetBox="[151,1436,191,1818]" captionTargetId="figure@8.[151,1436,191,1818]" captionTargetPageId="8" captionText="FIGURE 4. SEM micrographs of Epiceraticelus mandyae n. sp.: A, Male habitus. B, Ventral view of male left palp. C, Retrolateral view of tibial apophysis of male left palp. D, Dorsal view of epigynum, cleared. E, Ventral view of epigynum. Abbreviations: E, embolus; EA, embolic apophysis; EM, embolic membrane; PC, paracymbium; RTA, retrolateral tibial apophysis; T, tegulum; TP, tailpiece; S, spermathecae; CD, copulatory duct; CO, copulatory opening; MP, median plate. Scale bars, A, 0.5 mm; B, 0.1 mm; C, D, E, 0.05 mm." httpUri="https://zenodo.org/record/3346307/files/figure.png" pageId="2" pageNumber="191">4E</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FDABFAAEFDAFFAF9" box="[543,581,1294,1321]" captionStart="FIGURE 6" captionStartId="10.[151,250,1118,1143]" captionTargetBox="[170,1436,195,1093]" captionTargetId="figure@10.[151,1436,181,1099]" captionTargetPageId="10" captionText="FIGURE 6. Free hand llustrations of Epiceraticelus mandyae n. sp. (proportions are approximate). A, Male left palp, ventral view. B. Male left palp,, retrolateral view. C, Retrolateral view of tibial apophysis of left palp of male. D, Ventral view of epigynum. E,Dorsal schematic of epigynum. F, Lateral view of male carapace.Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; EM, embolic membrane; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece." httpUri="https://zenodo.org/record/3346311/files/figure.png" pageId="2" pageNumber="191">6D</figureCitation>
|
||
). Males of
|
||
<taxonomicName id="4C1C4D72735E1871FD0BFAAEFC8BFAF8" authorityName="Crosby & Bishop" authorityYear="1931" box="[703,865,1294,1320]" class="Arachnida" family="Linyphiidae" genus="Epiceraticelus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B968EAE3735E1871FD0BFAAEFC8BFAF8" box="[703,865,1294,1320]" italics="true" pageId="2" pageNumber="191">Epiceraticelus</emphasis>
|
||
</taxonomicName>
|
||
are easily distinguished from
|
||
<taxonomicName id="4C1C4D72735E1871FB18FAAEFA90FAF8" baseAuthorityName="Emerton" baseAuthorityYear="1882" box="[1196,1402,1294,1320]" class="Arachnida" family="Linyphiidae" genus="Scylaceus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="pallidus">
|
||
<emphasis id="B968EAE3735E1871FB18FAAEFA90FAF8" box="[1196,1402,1294,1320]" italics="true" pageId="2" pageNumber="191">Scylaceus pallidus</emphasis>
|
||
</taxonomicName>
|
||
by the long sinuous embolic division and by the fine serrations medially on the retrolateral tibial apophysis. Additionally, the narrow “pistol-shaped” tibial apophysis of
|
||
<taxonomicName id="4C1C4D72735E1871FD63FAF7FCA7FAA0" baseAuthorityName="Emerton" baseAuthorityYear="1882" box="[727,845,1366,1393]" class="Arachnida" family="Linyphiidae" genus="Scylaceus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="pallidus">
|
||
<emphasis id="B968EAE3735E1871FD63FAF7FCA7FAA0" box="[727,845,1366,1393]" italics="true" pageId="2" pageNumber="191">S. pallidus</emphasis>
|
||
</taxonomicName>
|
||
is distinctive (
|
||
<bibRefCitation id="EF8D4B00735E1871FC45FAF6FAFEFAA0" author="Paquin, P. & Duperre, N." box="[1009,1300,1366,1392]" pageId="2" pageNumber="191" pagination="1 - 251" refId="ref7750" refString="Paquin, P. & Duperre, N. (2003) Guide d'identification des araignees de Quebec. Fabreries, Supplement 11, 1 - 251." type="journal article" year="2003">Paquin and Dupérré 2003</bibRefCitation>
|
||
, Figs. 1230, 1231).
|
||
<taxonomicName id="4C1C4D72735E1871FF53FADAFE63FA44" authorityName="Crosby & Bishop" authorityYear="1931" box="[231,393,1402,1428]" class="Arachnida" family="Linyphiidae" genus="Epiceraticelus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B968EAE3735E1871FF53FADAFE63FA44" box="[231,393,1402,1428]" italics="true" pageId="2" pageNumber="191">Epiceraticelus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D72735E1871FE74FADAFD65FA44" baseAuthorityName="Emerton" baseAuthorityYear="1882" box="[448,655,1402,1428]" class="Arachnida" family="Linyphiidae" genus="Scylaceus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="pallidus">
|
||
<emphasis id="B968EAE3735E1871FE74FADAFD65FA44" box="[448,655,1402,1428]" italics="true" pageId="2" pageNumber="191">Scylaceus pallidus</emphasis>
|
||
</taxonomicName>
|
||
females are similar somatically and have a similar posteriorly-placed median plate, but in
|
||
<taxonomicName id="4C1C4D72735E1871FEC9FA3EFDF5FA68" authorityName="Crosby & Bishop" authorityYear="1931" box="[381,543,1438,1464]" class="Arachnida" family="Linyphiidae" genus="Epiceraticelus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B968EAE3735E1871FEC9FA3EFDF5FA68" box="[381,543,1438,1464]" italics="true" pageId="2" pageNumber="191">Epiceraticelus</emphasis>
|
||
</taxonomicName>
|
||
, it is triangular in shape rather than broadly rectangular (
|
||
<figureCitation id="13272A74735E1871FB2BFA3EFAE9FA68" box="[1183,1283,1438,1464]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1891]" captionTargetBox="[151,1434,211,1836]" captionTargetId="figure@5.[151,1434,211,1836]" captionTargetPageId="5" captionText="FIGURE 2. SEM micrographs of Epiceraticelus fluvialis: A, Retrolateral view of tibial apophysis of left palp of male. B, Ventral view of male left palp. C, Dorsal view of epigynum, cleared. D, Ventral view of epigynum. Abbreviations: CD, copulatory ducts; CG, copulatory grooves; CO, copulatory openings; E, embolus; EA, embolic apophysis; FD, fertilization ducts; MP, median plate; PC, paracymbium; S, spermathecae; ST, subtegulum; T, tegulum; TP, tailpiece. Scale bars, A, B, 0.1 mm; C, D, 0.05 mm." httpUri="https://zenodo.org/record/3346303/files/figure.png" pageId="2" pageNumber="191">Figs. 2D</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FABBFA3EFADAFA68" box="[1295,1328,1438,1464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1245,1270]" captionTargetBox="[156,1407,186,1217]" captionTargetId="figure@6.[151,1436,181,1226]" captionTargetPageId="6" captionText="FIGURE 3. Free hand Illustrations of Epiceraticelus fluvialis (proporations are approximate): A. Male left palp, ventral view. B, Male left palp, retrolateral view. C, Retrolateral view of tibial apophysis of male left palp. D, Lateral view of male carapace; E, Ventral view of epigynum. F, Dorsal schematic of epigynum. Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece. Proportions are approximate." httpUri="https://zenodo.org/record/3346305/files/figure.png" pageId="2" pageNumber="191">3E</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FA88FA3EFAB4FA68" box="[1340,1374,1438,1464]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1876]" captionTargetBox="[151,1436,191,1818]" captionTargetId="figure@8.[151,1436,191,1818]" captionTargetPageId="8" captionText="FIGURE 4. SEM micrographs of Epiceraticelus mandyae n. sp.: A, Male habitus. B, Ventral view of male left palp. C, Retrolateral view of tibial apophysis of male left palp. D, Dorsal view of epigynum, cleared. E, Ventral view of epigynum. Abbreviations: E, embolus; EA, embolic apophysis; EM, embolic membrane; PC, paracymbium; RTA, retrolateral tibial apophysis; T, tegulum; TP, tailpiece; S, spermathecae; CD, copulatory duct; CO, copulatory opening; MP, median plate. Scale bars, A, 0.5 mm; B, 0.1 mm; C, D, E, 0.05 mm." httpUri="https://zenodo.org/record/3346307/files/figure.png" pageId="2" pageNumber="191">4E</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FADEFA3EFA7AFA69" box="[1386,1424,1438,1465]" captionStart="FIGURE 6" captionStartId="10.[151,250,1118,1143]" captionTargetBox="[170,1436,195,1093]" captionTargetId="figure@10.[151,1436,181,1099]" captionTargetPageId="10" captionText="FIGURE 6. Free hand llustrations of Epiceraticelus mandyae n. sp. (proportions are approximate). A, Male left palp, ventral view. B. Male left palp,, retrolateral view. C, Retrolateral view of tibial apophysis of left palp of male. D, Ventral view of epigynum. E,Dorsal schematic of epigynum. F, Lateral view of male carapace.Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; EM, embolic membrane; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece." httpUri="https://zenodo.org/record/3346311/files/figure.png" pageId="2" pageNumber="191">6D</figureCitation>
|
||
). Moreover, the epigynum in
|
||
<taxonomicName id="4C1C4D72735E1871FE7AFA63FDAEFA0C" baseAuthorityName="Emerton" baseAuthorityYear="1882" box="[462,580,1474,1501]" class="Arachnida" family="Linyphiidae" genus="Scylaceus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="pallidus">
|
||
<emphasis id="B968EAE3735E1871FE7AFA63FDAEFA0C" box="[462,580,1474,1501]" italics="true" pageId="2" pageNumber="191">S. pallidus</emphasis>
|
||
</taxonomicName>
|
||
has spermathecae that are more posteriolateral in position (
|
||
<figureCitation id="13272A74735E1871FB69FA62FAD4FA0C" box="[1245,1342,1474,1500]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1891]" captionTargetBox="[151,1434,211,1836]" captionTargetId="figure@5.[151,1434,211,1836]" captionTargetPageId="5" captionText="FIGURE 2. SEM micrographs of Epiceraticelus fluvialis: A, Retrolateral view of tibial apophysis of left palp of male. B, Ventral view of male left palp. C, Dorsal view of epigynum, cleared. D, Ventral view of epigynum. Abbreviations: CD, copulatory ducts; CG, copulatory grooves; CO, copulatory openings; E, embolus; EA, embolic apophysis; FD, fertilization ducts; MP, median plate; PC, paracymbium; S, spermathecae; ST, subtegulum; T, tegulum; TP, tailpiece. Scale bars, A, B, 0.1 mm; C, D, 0.05 mm." httpUri="https://zenodo.org/record/3346303/files/figure.png" pageId="2" pageNumber="191">Figs. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FAFEFA62FA82FA0C" box="[1354,1384,1474,1500]" captionStart="FIGURE 3" captionStartId="6.[151,250,1245,1270]" captionTargetBox="[156,1407,186,1217]" captionTargetId="figure@6.[151,1436,181,1226]" captionTargetPageId="6" captionText="FIGURE 3. Free hand Illustrations of Epiceraticelus fluvialis (proporations are approximate): A. Male left palp, ventral view. B, Male left palp, retrolateral view. C, Retrolateral view of tibial apophysis of male left palp. D, Lateral view of male carapace; E, Ventral view of epigynum. F, Dorsal schematic of epigynum. Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece. Proportions are approximate." httpUri="https://zenodo.org/record/3346305/files/figure.png" pageId="2" pageNumber="191">3F</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FAC7FA62FA72FA0C" box="[1395,1432,1474,1500]" captionStart="FIGURE 4" captionStartId="8.[151,250,1851,1876]" captionTargetBox="[151,1436,191,1818]" captionTargetId="figure@8.[151,1436,191,1818]" captionTargetPageId="8" captionText="FIGURE 4. SEM micrographs of Epiceraticelus mandyae n. sp.: A, Male habitus. B, Ventral view of male left palp. C, Retrolateral view of tibial apophysis of male left palp. D, Dorsal view of epigynum, cleared. E, Ventral view of epigynum. Abbreviations: E, embolus; EA, embolic apophysis; EM, embolic membrane; PC, paracymbium; RTA, retrolateral tibial apophysis; T, tegulum; TP, tailpiece; S, spermathecae; CD, copulatory duct; CO, copulatory opening; MP, median plate. Scale bars, A, 0.5 mm; B, 0.1 mm; C, D, E, 0.05 mm." httpUri="https://zenodo.org/record/3346307/files/figure.png" pageId="2" pageNumber="191">4D</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FF23FA46FF5CF9D0" box="[151,182,1510,1536]" captionStart="FIGURE 6" captionStartId="10.[151,250,1118,1143]" captionTargetBox="[170,1436,195,1093]" captionTargetId="figure@10.[151,1436,181,1099]" captionTargetPageId="10" captionText="FIGURE 6. Free hand llustrations of Epiceraticelus mandyae n. sp. (proportions are approximate). A, Male left palp, ventral view. B. Male left palp,, retrolateral view. C, Retrolateral view of tibial apophysis of left palp of male. D, Ventral view of epigynum. E,Dorsal schematic of epigynum. F, Lateral view of male carapace.Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; EM, embolic membrane; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece." httpUri="https://zenodo.org/record/3346311/files/figure.png" pageId="2" pageNumber="191">6E</figureCitation>
|
||
and
|
||
<bibRefCitation id="EF8D4B00735E1871FF59FA46FE16F9D0" author="Paquin, P. & Duperre, N." box="[237,508,1510,1536]" pageId="2" pageNumber="191" pagination="1 - 251" refId="ref7750" refString="Paquin, P. & Duperre, N. (2003) Guide d'identification des araignees de Quebec. Fabreries, Supplement 11, 1 - 251." type="journal article" year="2003">Paquin & Dupérré 2003</bibRefCitation>
|
||
, Fig. 1232).
|
||
</paragraph>
|
||
<paragraph id="8BA336F1735E1870FF73F9A9FD69FE93" blockId="2.[151,1437,1149,2041]" lastBlockId="3.[151,1437,153,431]" lastPageId="3" lastPageNumber="192" pageId="2" pageNumber="191">
|
||
<emphasis id="B968EAE3735E1871FF73F9A9FEB3F9F4" bold="true" box="[199,345,1545,1572]" pageId="2" pageNumber="191">Description.</emphasis>
|
||
Length: 1.1–1.3. Cephalothorax: eyes eight, AME smallest, ALE and PLE equal, PME approximately two-thirds the size of PLE, posterior eye row slightly procurved, anterior eye row slightly recurved; eyes ringed with black. Chelicerae with six or seven promarginal teeth, five retromarginal denticles. Carapace oval, two-thirds as wide as long with one to two medial setae; a pair of small setae between PME and AME of each side; anterior portion of carapace three times as high as section near pedicel, clypeal modifications variable, dusky orange to tan, sternum nearly as long as wide, pointed posteriorly between coxae of leg IV. Abdomen: oval, covered with simple setae, concolorous gray, venter gray with dark gray patches around spinnerets and near epigynum; epiandrous fusules absent; colulus twice as long as wide; anterior spinnerets longer than posterior, cone-shaped. Tracheal system desmitracheate (
|
||
<emphasis id="B968EAE3735E1871FDA1F88BFDB9F894" box="[533,595,1835,1860]" italics="true" pageId="2" pageNumber="191">sensu</emphasis>
|
||
<bibRefCitation id="EF8D4B00735E1871FDE9F88AFCE9F894" author="Millidge, A. F." box="[605,771,1834,1860]" pageId="2" pageNumber="191" pagination="229 - 267" refId="ref7636" refString="Millidge, A. F. (1984) The taxonomy of the Linyphiidae, based chiefly on the epigynal and tracheal characters (Araneae: Linyphiidae). Bulletin of the British Arachnological Society, 6 (6), 229 - 267." type="journal article" year="1984">Millidge 1984</bibRefCitation>
|
||
). Legs: light orange, unmarked; relative length 4, 1, 2, 3; patella–tibia I three-fourths as long as carapace; metatarsi I–III each with a single trichobothrium; paired tarsal claws with eight teeth, median claw with three teeth. Trichobothrium on metatarsus IV lacking. Trichobothrium on metatarsus I at 0.4–0.5. Tibial spines 1111, often 0000. Palp of male complex (
|
||
<figureCitation id="13272A74735E1871FBE4F836FB5CF860" box="[1104,1206,1942,1968]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1891]" captionTargetBox="[151,1434,211,1836]" captionTargetId="figure@5.[151,1434,211,1836]" captionTargetPageId="5" captionText="FIGURE 2. SEM micrographs of Epiceraticelus fluvialis: A, Retrolateral view of tibial apophysis of left palp of male. B, Ventral view of male left palp. C, Dorsal view of epigynum, cleared. D, Ventral view of epigynum. Abbreviations: CD, copulatory ducts; CG, copulatory grooves; CO, copulatory openings; E, embolus; EA, embolic apophysis; FD, fertilization ducts; MP, median plate; PC, paracymbium; S, spermathecae; ST, subtegulum; T, tegulum; TP, tailpiece. Scale bars, A, B, 0.1 mm; C, D, 0.05 mm." httpUri="https://zenodo.org/record/3346303/files/figure.png" pageId="2" pageNumber="191">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="13272A74735E1871FB70F836FB3AF860" box="[1220,1232,1942,1968]" captionStart="FIGURE 3" captionStartId="6.[151,250,1245,1270]" captionTargetBox="[156,1407,186,1217]" captionTargetId="figure@6.[151,1436,181,1226]" captionTargetPageId="6" captionText="FIGURE 3. Free hand Illustrations of Epiceraticelus fluvialis (proporations are approximate): A. Male left palp, ventral view. B, Male left palp, retrolateral view. C, Retrolateral view of tibial apophysis of male left palp. D, Lateral view of male carapace; E, Ventral view of epigynum. F, Dorsal schematic of epigynum. Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece. Proportions are approximate." httpUri="https://zenodo.org/record/3346305/files/figure.png" pageId="2" pageNumber="191">3</figureCitation>
|
||
A–B, 4B, 6A–B), with a well-developed retrolateral tibial apophysis which is finely serrated on medial margin; embolic division with elongate tail-piece oriented with long axis parallel to long axis of palp; tegulum with distal protegulum; distal suprategular apophysis present; long sinuous embolus present (
|
||
<figureCitation id="13272A74735F1870FCD7FF39FC45FF63" box="[867,943,153,179]" captionStart="FIGURE 3" captionStartId="6.[151,250,1245,1270]" captionTargetBox="[156,1407,186,1217]" captionTargetId="figure@6.[151,1436,181,1226]" captionTargetPageId="6" captionText="FIGURE 3. Free hand Illustrations of Epiceraticelus fluvialis (proporations are approximate): A. Male left palp, ventral view. B, Male left palp, retrolateral view. C, Retrolateral view of tibial apophysis of male left palp. D, Lateral view of male carapace; E, Ventral view of epigynum. F, Dorsal schematic of epigynum. Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece. Proportions are approximate." httpUri="https://zenodo.org/record/3346305/files/figure.png" pageId="3" pageNumber="192">Figs. 3</figureCitation>
|
||
A–B, 6A–B). Epigynum (
|
||
<figureCitation id="13272A74735F1870FB62FF39FAC8FF63" box="[1238,1314,153,179]" captionStart="FIGURE 3" captionStartId="6.[151,250,1245,1270]" captionTargetBox="[156,1407,186,1217]" captionTargetId="figure@6.[151,1436,181,1226]" captionTargetPageId="6" captionText="FIGURE 3. Free hand Illustrations of Epiceraticelus fluvialis (proporations are approximate): A. Male left palp, ventral view. B, Male left palp, retrolateral view. C, Retrolateral view of tibial apophysis of male left palp. D, Lateral view of male carapace; E, Ventral view of epigynum. F, Dorsal schematic of epigynum. Abbreviations: C, cymbium; CD, copulatory duct; CO, copulatory opening; DSA, distal suprategular apophysis; E, embolus; EA, embolic apophysis; FD, fertilization duct; MP, median plate; PC, paracymbium; PT, protegulum; RTA, retrolateral tibial apophysis; S, spermathecae; SPT, suprategulum; ST, subtegulum; T, tegulum; TP, tailpiece. Proportions are approximate." httpUri="https://zenodo.org/record/3346305/files/figure.png" pageId="3" pageNumber="192">Figs. 3</figureCitation>
|
||
EF, 6D–E) with dual sinuous median epigynal grooves widening into a posterior median plate (narrower and separated in
|
||
<taxonomicName id="4C1C4D72735F1870FA30FF1EFF1BFF2B" authorityName="Crosby and Bishop" authorityYear="1931" class="Arachnida" family="Linyphiidae" genus="Epiceraticelus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="fluvialis">
|
||
<emphasis id="B968EAE3735F1870FA30FF1EFF1BFF2B" italics="true" pageId="3" pageNumber="192">E. fluvialis</emphasis>
|
||
</taxonomicName>
|
||
, while wider and fused in
|
||
<taxonomicName id="4C1C4D72735F1870FD94FF42FD4FFF2B" authority="Draney & Milne & Ulyshen & Madriz, 2019" authorityName="Draney & Milne & Ulyshen & Madriz" authorityYear="2019" box="[544,677,225,251]" class="Arachnida" family="Linyphiidae" genus="Epiceraticelus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="mandyae" status="sp. nov.">
|
||
<emphasis id="B968EAE3735F1870FD94FF42FD4FFF2B" box="[544,677,225,251]" italics="true" pageId="3" pageNumber="192">E. mandyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B5798735F1870FD19FF41FD07FF2B" box="[685,749,225,251]" pageId="3" pageNumber="192" rank="species">
|
||
<emphasis id="B968EAE3735F1870FD19FF41FD07FF2B" bold="true" box="[685,749,225,251]" pageId="3" pageNumber="192">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
), spermathecae ventrally visible, separated by more than 1.5 times their diameter; internally, spermathecae anterior of copulatory and fertilization ducts, laterally directed and partially covering spermathecae posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BA336F1735F1870FF73FEECFC42FEB7" blockId="3.[151,1437,153,431]" box="[199,936,332,359]" pageId="3" pageNumber="192">
|
||
<emphasis id="B968EAE3735F1870FF73FEECFE8BFEB7" bold="true" box="[199,353,332,359]" pageId="3" pageNumber="192">Distribution.</emphasis>
|
||
New York south to
|
||
<collectingCountry id="F30B7661735F1870FDF4FEEDFD77FEB7" box="[576,669,333,359]" name="Georgia" pageId="3" pageNumber="192">Georgia</collectingCountry>
|
||
, west to Texas (
|
||
<figureCitation id="13272A74735F1870FCE2FEEDFC7DFEB7" box="[854,919,333,359]" captionStart="FIGURE 1" captionStartId="1.[151,250,1652,1677]" captionTargetBox="[151,1436,236,1631]" captionTargetId="figure@1.[151,1436,181,1631]" captionTargetPageId="1" captionText="FIGURE 1. Collection localities of Epiceraticelus fluvialis (filled circles) and Epiceraticelus mandyae n. sp. (filled stars)." httpUri="https://zenodo.org/record/3346301/files/figure.png" pageId="3" pageNumber="192">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA336F1735F1870FF73FED1FD9BFE7F" blockId="3.[151,1437,153,431]" pageId="3" pageNumber="192">
|
||
<emphasis id="B968EAE3735F1870FF73FED1FE61FE5B" bold="true" box="[199,395,368,395]" pageId="3" pageNumber="192">Natural History.</emphasis>
|
||
May be found in the leaf litter of damp forests in winter and early spring. Forests commonly consist of various species of pine and oak.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |