173 lines
18 KiB
XML
173 lines
18 KiB
XML
<document id="0249E33871CA2D5877D3E2DF32586EE8" ID-CLB-Dataset="36820" ID-DOI="10.11646/zootaxa.3829.1.1" ID-GBIF-Dataset="a4b69269-5a4c-4b5b-9f89-6b314d3ba284" ID-ISSN="1175-5326" ID-Zenodo-Dep="286556" ID-ZooBank="75210CDC-AC6A-4624-A6F1-1BC969BC7CAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452709399" checkinUser="plazi" docAuthor="Pyron, Robert Alexander & Wallach, Van" docDate="2014" docId="03B587DDC106B171CFD7CC5EFA2EF874" docLanguage="en" docName="zt03829p081.pdf" docOrigin="Zootaxa 3829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Typhlopidae" docType="treatment" docVersion="8" lastPageNumber="62" masterDocId="FF8CFFA5C13AB14CCF40CA0EFFB9FFA2" masterDocTitle="Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence" masterLastPageNumber="81" masterPageNumber="1" pageNumber="61" updateTime="1698338434491" updateUser="ExternalLinkService">
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<mods:mods id="03020E7FF21F9A773066DF7410B0839A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A978BC17005ED5906005F04657578D34">
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<mods:title id="A6C88DF0A44F35162F942514AC974FA8">Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence</mods:title>
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<mods:name id="27C372419467207BFA01ADAF412BE18B" type="personal">
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<mods:roleTerm id="958EC98EA858A764F3B851D788714E3B">Author</mods:roleTerm>
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<mods:namePart id="2001B4C8E30AE1B987F62BF92712AE06">Pyron, Robert Alexander</mods:namePart>
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<mods:name id="B491F455EB1E5BE3860E0EB7484A9C89" type="personal">
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<mods:roleTerm id="5E104C7DAD8F15469BD1410BB34F5BBB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="30BBEC644A8CBC73A1AB1ECAFBB37417">Wallach, Van</mods:namePart>
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<mods:typeOfResource id="30EBB1D0C5C98EDFE1C5D55F09B7DB4A">text</mods:typeOfResource>
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<mods:title id="F9FBED8D85BEDCA0C5932C540F300FD0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A5458363E079782BE2D3024212F37D7C">
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<mods:date id="FFFEAAEF6A1F2C0A899ABE9435D3E3EF">2014</mods:date>
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<mods:detail id="008E5B583F5FC3DB54A7B1B2667C5BB6" type="volume">
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<mods:number id="F410E6435274422DD8B312553C6E6D25">3829</mods:number>
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<mods:detail id="63AA69687D5E2A9533C0419E4B88B3D6" type="issue">
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<mods:number id="7FB407ED3BF81AEB35593BB26B97AE7B">1</mods:number>
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<mods:classification id="53C0E55A5188165C7B8E39662438D772">journal article</mods:classification>
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<mods:identifier id="E3F301F8D96E753D5EB69585B560A879" type="DOI">10.11646/zootaxa.3829.1.1</mods:identifier>
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<treatment id="03B587DDC106B171CFD7CC5EFA2EF874" ID-DOI="http://doi.org/10.5281/zenodo.6128010" ID-GBIF-Taxon="127668684" ID-Zenodo-Dep="6128010" LSID="urn:lsid:plazi:treatment:03B587DDC106B171CFD7CC5EFA2EF874" httpUri="http://treatment.plazi.org/id/03B587DDC106B171CFD7CC5EFA2EF874" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
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<subSubSection id="C3066540C106B171CFD7CC5EFA2EF874" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="nomenclature">
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<paragraph id="8BA336CBC106B170CFD7CC5EFE67F9C8" blockId="60.[151,478,1616,1642]" box="[151,478,1616,1642]" pageId="60" pageNumber="61">
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<heading id="D0EB81A7C106B170CFD7CC5EFE67F9C8" bold="true" box="[151,478,1616,1642]" fontSize="11" level="1" pageId="60" pageNumber="61" reason="1">
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<emphasis id="B968EAD9C106B170CFD7CC5EFE67F9C8" bold="true" box="[151,478,1616,1642]" pageId="60" pageNumber="61">
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<taxonomicName id="4C1C4D48C106B170CFD7CC5EFE8AF9C8" box="[151,307,1616,1642]" class="Reptilia" family="Typhlopidae" kingdom="Animalia" order="Squamata" pageId="60" pageNumber="61" phylum="Chordata" rank="family">Typhlopidae</taxonomicName>
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<emphasis id="B968EAD9C106B170CE7ACC5EFE67F9C8" bold="true" box="[314,478,1616,1642]" italics="true" pageId="60" pageNumber="61">incertae sedis</emphasis>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA336CBC106B170CFD7CC98FEDBF851" blockId="60.[151,1436,1686,2035]" pageId="60" pageNumber="61">
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The species “
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<taxonomicName id="4C1C4D48C106B170CE6DCC98FE28F90D" box="[301,401,1686,1711]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C106B170CE6DCC98FE28F90D" box="[301,401,1686,1711]" italics="true" pageId="60" pageNumber="61">Typhlops</emphasis>
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</taxonomicName>
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”
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<taxonomicName id="4C1C4D48C106B170CEE4CC98FD91F90D" box="[420,552,1686,1711]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="longissimus">
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<emphasis id="B968EAD9C106B170CEE4CC98FD91F90D" box="[420,552,1686,1711]" italics="true" pageId="60" pageNumber="61">longissimus</emphasis>
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</taxonomicName>
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was described from North
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<collectingCountry id="F30B765BC106B170CC18CC98FC00F90D" box="[856,953,1686,1711]" name="United States of America" pageId="60" pageNumber="61">America</collectingCountry>
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by
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<bibRefCitation id="EF8D4B3AC106B170CCA2CC98FABDF90D" author="Dumeril" box="[994,1284,1686,1711]" pageId="60" pageNumber="61" refString="Dumeril, A. M. C. & Bibron, G. (1844) Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Tome sixieme. Librairie Encyclopedique de Roret, Paris, xii + 609 pp." type="book" year="1844">Duméril & Bibron (1844)</bibRefCitation>
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, but
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<bibRefCitation id="EF8D4B3AC106B170CA7ACC98FEDBF971" author="Dixon" pageId="60" pageNumber="61" refString="Dixon, J. R. & Hendricks, F. S. (1979) The Wormsnakes (Family Typhlopidae) of the Neotropics, exclusive of the Antilles. Zoologische Verhandelingen, 173, 1 - 39." type="journal article" year="1979">Dixon & Hendricks (1979)</bibRefCitation>
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,
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<bibRefCitation id="EF8D4B3AC106B170CE33CCB4FDB0F971" author="Hahn" box="[371,521,1722,1747]" pageId="60" pageNumber="61" refString="Hahn, D. E. (1980) Liste der rezenten Amphibien und Reptilien: Anomalepididae, Leptotyphlopidae, Typhlopidae. Das Tierreich, Berlin, 101, i - xii + 1 - 93. http: // dx. doi. org / 10.1515 / 9783111385556. fm" type="journal article" year="1980">Hahn (1980)</bibRefCitation>
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, and
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<bibRefCitation id="EF8D4B3AC106B170CD0ECCB4FCC9F971" box="[590,880,1722,1747]" pageId="60" pageNumber="61" refString="McDiarmid, R. W., Campbell, J. A. & Toure, T. S. A. (1999) Snake Species of the World: a Taxonomic and Geographic Reference (Volume 1). Herpetologists' League, Washington, DC, 512 pp." type="book">
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McDiarmid
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<emphasis id="B968EAD9C106B170CD9CCCB2FCA3F971" box="[732,794,1722,1747]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
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(1999)
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</bibRefCitation>
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all considered it
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<emphasis id="B968EAD9C106B170CB04CCB2FB5DF971" box="[1092,1252,1722,1747]" italics="true" pageId="60" pageNumber="61">incertae sedis</emphasis>
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with erroneous locality data. The
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<typeStatus id="54A78869C106B170CE2ACCD1FE23F955" box="[362,410,1759,1783]" pageId="60" pageNumber="61">type</typeStatus>
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(MNHN 1061) has been re-examined and shows a possible relationship to
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<taxonomicName id="4C1C4D48C106B170CA4ACCD0FEB5F8B9" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="tenuicollis">
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<emphasis id="B968EAD9C106B170CA4ACCD0FEB5F8B9" italics="true" pageId="60" pageNumber="61">Indotyphlops tenuicollis</emphasis>
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</taxonomicName>
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of northern
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<collectingCountry id="F30B765BC106B170CEE5CD0CFE66F8B9" box="[421,479,1794,1819]" name="India" pageId="60" pageNumber="61">India</collectingCountry>
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based on scale counts and proportion of head shields (Table 2). We continue to consider it
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<taxonomicName id="4C1C4D48C106B170CE55CD28FE18F89D" box="[277,417,1830,1855]" class="Reptilia" family="Typhlopidae" kingdom="Animalia" order="Squamata" pageId="60" pageNumber="61" phylum="Chordata" rank="family">Typhlopidae</taxonomicName>
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<emphasis id="B968EAD9C106B170CEE9CD29FDFCF89D" box="[425,581,1830,1855]" italics="true" pageId="60" pageNumber="61">incertae sedis</emphasis>
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, though it is considered a valid species (Wallach
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<emphasis id="B968EAD9C106B170CB37CD29FB08F89D" box="[1143,1201,1830,1855]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
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2014). This taxon is also the
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<typeStatus id="54A78869C106B170CFB8CD45FE91F8C1" box="[248,296,1867,1891]" pageId="60" pageNumber="61">type</typeStatus>
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species of the genus
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<taxonomicName id="4C1C4D48C106B170CD61CD44FC4CF8C1" authority="Dumeril & Bibron, 1844" authorityName="Dumeril & Bibron" authorityYear="1844" box="[545,1013,1866,1891]" class="Reptilia" family="Typhlopidae" genus="Opthalmidion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C106B170CD61CD44FD04F8C1" box="[545,701,1866,1891]" italics="true" pageId="60" pageNumber="61">Opthalmidion</emphasis>
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(
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<bibRefCitation id="EF8D4B3AC106B170CD8DCD44FC54F8C1" author="Dumeril" box="[717,1005,1866,1891]" pageId="60" pageNumber="61" refString="Dumeril, A. M. C. & Bibron, G. (1844) Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Tome sixieme. Librairie Encyclopedique de Roret, Paris, xii + 609 pp." type="book" year="1844">Duméril & Bibron, 1844</bibRefCitation>
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)
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</taxonomicName>
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, by subsequent designation of
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<bibRefCitation id="EF8D4B3AC106B170CA17CD45FF5CF825" author="Peters" pageId="60" pageNumber="61" refString="Peters, W. C. H. (1881) Einige herpetologische Mittheilungen. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin, 1881, 69 - 72." type="journal article" year="1881">Peters (1881)</bibRefCitation>
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. Thus, if this species is found to be placed in
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<taxonomicName id="4C1C4D48C106B170CC60CD60FC0BF825" box="[800,946,1902,1927]" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C106B170CC60CD60FC0BF825" box="[800,946,1902,1927]" italics="true" pageId="60" pageNumber="61">Indotyphlops</emphasis>
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</taxonomicName>
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(or any other genus erected after 1844),
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<taxonomicName id="4C1C4D48C106B170CFD7CD9CFEF8F809" box="[151,321,1938,1963]" class="Squamata" family="Typhlopidae" genus="Ophthalmidion" higherTaxonomySource="CoL" kingdom="Animalia" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C106B170CFD7CD9CFEF8F809" box="[151,321,1938,1963]" italics="true" pageId="60" pageNumber="61">Ophthalmidion</emphasis>
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</taxonomicName>
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will assume priority for that taxon. A number of other species of indeterminate or ambiguous origin have been described in
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<taxonomicName id="4C1C4D48C106B170CEE7CDB8FD8AF86D" box="[423,563,1974,1999]" class="Reptilia" family="Typhlopidae" kingdom="Animalia" order="Squamata" pageId="60" pageNumber="61" phylum="Chordata" rank="family">Typhlopidae</taxonomicName>
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(see Wallach
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<emphasis id="B968EAD9C106B170CD97CDB9FCABF86D" box="[727,786,1974,1999]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
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2014), but we have no further evidence to bear on their status at this time.
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</paragraph>
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<caption id="DF636643C107B171CFD7CA96FA2CFE8D" pageId="61" pageNumber="62">
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<paragraph id="8BA336CBC107B171CFD7CA96FA2CFE8D" blockId="61.[151,1436,152,303]" pageId="61" pageNumber="62">
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<emphasis id="B968EAD9C107B171CFD7CA96FEB4FF0D" bold="true" box="[151,269,152,175]" pageId="61" pageNumber="62">TABLE 9.</emphasis>
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Visceral character states separating
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<emphasis id="B968EAD9C107B171CDDACA96FBB7FF0D" box="[666,1038,152,175]" italics="true" pageId="61" pageNumber="62">
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<taxonomicName id="4C1C4D48C107B171CDDACA96FC2BFF0D" box="[666,914,152,175]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="polygrammicus">Anilios polygrammicus</taxonomicName>
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(
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<taxonomicName id="4C1C4D48C107B171CCE4CA96FBBAFF0D" box="[932,1027,152,175]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="poly">An. poly</taxonomicName>
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.)
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</emphasis>
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from
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<emphasis id="B968EAD9C107B171CB13CA96FA22FF0D" box="[1107,1435,152,175]" italics="true" pageId="61" pageNumber="62">
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<taxonomicName id="4C1C4D48C107B171CB13CA96FA99FF0D" box="[1107,1312,152,175]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="torresianus">Anilios torresianus</taxonomicName>
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(
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<taxonomicName id="4C1C4D48C107B171CA71CA96FA35FF0D" box="[1329,1420,152,175]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="torr">An. torr</taxonomicName>
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.),
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</emphasis>
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based on measurement of the specimens in Appendix II (part). Characters are as follows: data in sections
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<emphasis id="B968EAD9C107B171CA09CAB6FA25FF6D" bold="true" box="[1353,1436,184,207]" pageId="61" pageNumber="62">(A)-(D)</emphasis>
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represent sample means as % SVL; organ lengths (PT = posterior tip) are included in section
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<emphasis id="B968EAD9C107B171CB3CCAD6FB18FF4D" bold="true" box="[1148,1185,216,239]" pageId="61" pageNumber="62">(A)</emphasis>
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; organ midpoints (MP) are listed in section
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<emphasis id="B968EAD9C107B171CE35CAF6FE20FEAD" bold="true" box="[373,409,248,271]" pageId="61" pageNumber="62">(B)</emphasis>
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; organ gaps (GAP) and intervals (INT) are compiled in section
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<emphasis id="B968EAD9C107B171CB1CCAF6FB38FEAD" bold="true" box="[1116,1153,248,271]" pageId="61" pageNumber="62">(C)</emphasis>
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; organ midpoint intervals (MP-MP INT) are included in section
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<emphasis id="B968EAD9C107B171CD6FCB16FDEDFE8D" bold="true" box="[559,596,280,303]" pageId="61" pageNumber="62">(D)</emphasis>
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; meristic values are listed in section
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<emphasis id="B968EAD9C107B171CC9FCB16FBBBFE8D" bold="true" box="[991,1026,280,303]" pageId="61" pageNumber="62">(E)</emphasis>
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; ratios of two characters listed in
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<emphasis id="B968EAD9C107B171CA2CCB16FA37FE8D" bold="true" box="[1388,1422,280,303]" pageId="61" pageNumber="62">(F)</emphasis>
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.
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</paragraph>
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</caption>
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<paragraph id="8BA336CBC107B171CFDFCB45FF7AFE11" blockId="61.[159,1431,330,2006]" pageId="61" pageNumber="62">
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Taxon
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<emphasis id="B968EAD9C107B171CDF9CB44FB68FEC2" box="[697,1233,330,352]" italics="true" pageId="61" pageNumber="62">
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<taxonomicName id="4C1C4D48C107B171CDF9CB44FCECFEC2" box="[697,853,330,352]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="polygrammicus">polygrammicus</taxonomicName>
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<taxonomicName id="4C1C4D48C107B171CB1ECB45FB68FEC2" authority="Sample" authorityName="Sample" box="[1118,1233,331,352]" class="Squamata" family="Typhlopidae" genus="Anilios" higherTaxonomySource="CoL" kingdom="Animalia" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="torresianus">torresianus</taxonomicName>
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</emphasis>
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Sample size
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<emphasis id="B968EAD9C107B171CDF9CB7BFD64FE28" box="[697,733,373,394]" italics="true" pageId="61" pageNumber="62">n =</emphasis>
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2
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<emphasis id="B968EAD9C107B171CB1ECB7BFB3BFE28" box="[1118,1154,373,394]" italics="true" pageId="61" pageNumber="62">n =</emphasis>
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3 (A)
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</paragraph>
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<paragraph id="8BA336CBC107B171CFDFCBC8FF78FC6E" blockId="61.[159,1431,330,2006]" pageId="61" pageNumber="62">Right liver 25.4 23.5 Left liver 25.1 23.5 Total (left + right) liver 50.4 47 Right kidney 3.3 4.8 Left kidney 3.3 4.5 Total (left + right) kidney 6.6 9.3 Rectal caecum 2.5 4.9 Trachea 31.5 36 Tracheal lung 19.1 24.3 Right lung 17.4 20.6 Right lung PT 49.8 57.5 Right bronchus 11.6 7.3 (B)</paragraph>
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<paragraph id="8BA336CBC107B171CFDFC9EFFF78F9C0" blockId="61.[159,1431,330,2006]" pageId="61" pageNumber="62">Heart MP 30.2 34.8 Right liver MP 47.7 50.8 Left liver MP 44.3 48.9 Total (left + right) liver MP 46.1 49.9 Right gonad MP 78.7 76.9 Left gonad MP 83.4 81.4 Total (left + right) gonad MP 81 79.1 Left adrenal MP 86 84.2 Total (left + right) adrenal MP 84.6 82.3 Right kidney MP 89.7 86.4 Left kidney MP 92.2 89.9 Total (left + right) kidney MP 90.9 88.1 Trachea MP 16.7 18.9 Tracheal lung MP 18.6 20.6 Right lung MP 41.1 47.2 (C)</paragraph>
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<paragraph id="8BA336CBC107B171CFDFCC7BFB33F80E" blockId="61.[159,1431,330,2006]" pageId="61" pageNumber="62">Snout-heart INT 32.4 36.9 Heart-liver INT 32.3 29.9 Heart-gall bladder GAP 34.6 31.3 Liver-gall bladder INT 33.4 30.7 Liver-kidney GAP 27.7 21.4 Liver-kidney INT 58.8 53 Gall bladder-kidney GAP 19.6 14.2 Gall bladder-kidney INT 26.9 23.9</paragraph>
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<paragraph id="8BA336CBC107B171CB21CDCFFA2EF874" blockId="61.[159,1431,330,2006]" box="[1121,1431,1984,2006]" pageId="61" pageNumber="62">
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..
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<emphasis id="B968EAD9C107B171CB2DCDCEFA2EF874" box="[1133,1431,1984,2006]" italics="true" pageId="61" pageNumber="62">....continued on the next page</emphasis>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |