133 lines
17 KiB
XML
133 lines
17 KiB
XML
<document id="E169BEE676462C7798A9195A3C9B317A" ID-CLB-Dataset="21928" ID-DOI="10.11646/zootaxa.4779.4.1" ID-GBIF-Dataset="d48aecc5-2a44-491e-802f-4556f052a7a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3839329" ID-ZooBank="861A7289-DD34-4174-B6B2-783FBEC8DF36" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590137887931" checkinUser="plazi" docAuthor="Kluge, Nikita J. & Suttinun, Chanaporn" docDate="2020" docId="03B587A2F21BFF89ECABFF500823FDE6" docLanguage="en" docName="zootaxa.4779.4.1.pdf" docOrigin="Zootaxa 4779 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Indocloeon (Hindocloeon) longistylus Kluge & Suttinun 2020, comb. n." docType="treatment" docVersion="6" lastPageNumber="479" masterDocId="FF8CFFDAF207FF95EC3CFFC90C6FFFB9" masterDocTitle="Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species" masterLastPageNumber="484" masterPageNumber="451" pageNumber="479" updateTime="1698840818980" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="480F458134C34302BF543B9ED0361AF3">Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species</mods:title>
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<mods:name id="9F7CD85748144A53B01C4B9A530BFB13" type="personal">
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<mods:namePart id="0DF2C2CA51596D44D74306359AE15AA4">Kluge, Nikita J.</mods:namePart>
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<mods:affiliation id="D8507211EFE113E6D0444AEADCC7C6CC">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia Website: http: // www. insecta. bio. spbu. ru. n. kluge @ spbu. ru; https: // orcid. org / 0000 - 0001 - 9741 - 7790</mods:affiliation>
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<mods:namePart id="4CB37ECA18623335180BD5C62BF2E206">Suttinun, Chanaporn</mods:namePart>
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<mods:affiliation id="A46D0EF230D7988DA961F68232D88BA3">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand. chana. suttinun @ gmail. com; https: // orcid. org / 0000 - 0002 - 0380 - 1554</mods:affiliation>
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<mods:title id="47B1522FFD9895E3CE63D6CC85067F7D">Zootaxa</mods:title>
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<mods:date id="B93B01EE9BCE95121E7F2722DD36EA93">2020</mods:date>
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<mods:number id="017AE861867F72475EB7017B1D002AA1">2020-05-21</mods:number>
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<mods:number id="27BBD716C0D47E45E003D765B71B5F50">4779</mods:number>
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<treatment id="03B587A2F21BFF89ECABFF500823FDE6" ID-DOI="http://doi.org/10.5281/zenodo.3853338" ID-GBIF-Taxon="164248614" ID-Zenodo-Dep="3853338" LSID="urn:lsid:plazi:treatment:03B587A2F21BFF89ECABFF500823FDE6" httpUri="http://treatment.plazi.org/id/03B587A2F21BFF89ECABFF500823FDE6" lastPageNumber="479" pageId="28" pageNumber="479">
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<subSubSection id="C306653FF21BFF89ECABFF500FC5FF0D" box="[151,938,153,180]" pageId="28" pageNumber="479" type="nomenclature">
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<paragraph id="8BA336B4F21BFF89ECABFF500FC5FF0D" blockId="28.[151,938,153,216]" box="[151,938,153,180]" pageId="28" pageNumber="479">
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<heading id="D0EB81D8F21BFF89ECABFF500FC5FF0D" bold="true" box="[151,938,153,180]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
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<emphasis id="B968EAA6F21BFF89ECABFF500FC5FF0D" bold="true" box="[151,938,153,180]" pageId="28" pageNumber="479">
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<taxonomicName id="4C1C4D37F21BFF89ECABFF500F57FF0A" authority="(Demoulin 1969)" authorityName="Kluge & Suttinun" authorityYear="2020" baseAuthorityName="Demoulin" baseAuthorityYear="1969" box="[151,824,153,180]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="longistylus" status="comb. nov." subGenus="Hindocloeon">
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<emphasis id="B968EAA6F21BFF89ECABFF500D4FFF0D" bold="true" box="[151,288,153,180]" italics="true" pageId="28" pageNumber="479">Indocloeon</emphasis>
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(
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<emphasis id="B968EAA6F21BFF89ED0DFF500DA2FF0D" bold="true" box="[305,461,153,180]" italics="true" pageId="28" pageNumber="479">Hindocloeon</emphasis>
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)
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<emphasis id="B968EAA6F21BFF89EDE3FF500E0EFF0D" bold="true" box="[479,609,153,180]" italics="true" pageId="28" pageNumber="479">longistylus</emphasis>
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(
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<bibRefCitation id="EF8D4B45F21BFF89EE4CFF500F5FFF0A" author="Demoulin, G." box="[624,816,153,179]" pageId="28" pageNumber="479" pagination="225 - 241" refId="ref14512" refString="Demoulin, G. (1969) Les Ephemeropteres recoltes par la mission danoise du Noona Dan aux iles Philippines et Bismarck. Entomologiske Meddelelser, 37 (3), 225 - 241." type="journal article" year="1969">Demoulin 1969</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A25B57DDF21BFF89EF7CFF530FC5FF0D" box="[832,938,154,180]" pageId="28" pageNumber="479" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653FF21BFF89ECABFF740D68FF61" box="[151,263,189,216]" pageId="28" pageNumber="479" type="description">
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<paragraph id="8BA336B4F21BFF89ECABFF740D68FF61" blockId="28.[151,938,153,216]" box="[151,263,189,216]" pageId="28" pageNumber="479">
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(
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<figureCitation id="13272A31F21BFF89EC9CFF740C90FF61" box="[160,255,189,216]" captionStart="FIGURES 101–105" captionStartId="29.[151,264,1853,1878]" captionTargetBox="[151,1427,281,1828]" captionTargetId="figure@29.[151,1436,273,1828]" captionTargetPageId="29" captionText="FIGURES 101–105. Male genitals of Hindocloeon. 101, imaginal genitals of Indocloeon (Hindocloeon) continentale sp. n. (penis dotted; 1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; gv-d, distal portion of gonovectis; gv-p, proximal portion of gonovectis; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 102, their subimaginal exuviae; 103, subimaginal gonostyli of I. (H.) sp. «Vietnam»; 104, subimaginal gonostyli of I. (H.) sp. «Java»; 105, imaginal genitals of I. longistylus (after Demoulin 1969)." figureDoi="http://doi.org/10.5281/zenodo.3839373" httpUri="https://zenodo.org/record/3839373/files/figure.png" pageId="28" pageNumber="479">Fig. 105</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653FF21BFF89ECABFECB0ECBFEA3" box="[151,676,257,282]" pageId="28" pageNumber="479" type="reference_group">
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<paragraph id="8BA336B4F21BFF89ECABFECB0ECBFEA3" blockId="28.[151,676,257,282]" box="[151,676,257,282]" pageId="28" pageNumber="479">
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<treatmentCitationGroup id="AB0C119AF21BFF89ECABFECB0ECBFEA3" box="[151,676,257,282]" pageId="28" pageNumber="479">
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<treatmentCitation id="0ABD10A5F21BFF89ECABFECB0E42FEA3" author="Demoulin, G." box="[151,557,257,282]" page="231" pageId="28" pageNumber="479" year="1969">
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<taxonomicName id="4C1C4D37F21BFF89ECABFECB0E42FEA3" ID-CoL="5ZC5X" authority="Demoulin 1969: 231" authorityName="Demoulin" authorityPageNumber="231" authorityYear="1969" box="[151,557,257,282]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="longistylus">
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<emphasis id="B968EAA6F21BFF89ECABFECB0D3AFEA3" box="[151,341,258,282]" italics="true" pageId="28" pageNumber="479">Cloeon longistylus</emphasis>
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<bibRefCitation id="EF8D4B45F21BFF89ED67FEC80E42FEA3" author="Demoulin, G." box="[347,557,257,282]" pageId="28" pageNumber="479" pagination="225 - 241" refId="ref14512" refString="Demoulin, G. (1969) Les Ephemeropteres recoltes par la mission danoise du Noona Dan aux iles Philippines et Bismarck. Entomologiske Meddelelser, 37 (3), 225 - 241." type="journal article" year="1969">Demoulin 1969: 231</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(
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<specimenCount id="9D1AFD3DF21BFF89EE07FECB0E21FEA3" box="[571,590,258,282]" pageId="28" pageNumber="479" type="male">♂</specimenCount>
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imago).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653FF21BFF89ECABFE8E0EFBFD83" pageId="28" pageNumber="479" type="discussion">
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<paragraph id="8BA336B4F21BFF89ECABFE8E0EFBFD83" blockId="28.[151,1437,327,607]" pageId="28" pageNumber="479">
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<emphasis id="B968EAA6F21BFF89ECABFE8E0D70FEDB" bold="true" box="[151,287,327,354]" pageId="28" pageNumber="479">Comments.</emphasis>
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This species was briefly described based on
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<specimenCount id="9D1AFD3DF21BFF89EF28FE810F30FEDB" box="[788,863,328,354]" pageId="28" pageNumber="479" type="male">3 male</specimenCount>
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imagines. It was placed in the genus
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<taxonomicName id="4C1C4D37F21BFF89E93CFE810CBDFE3F" authority="Leach 1815" authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EAA6F21BFF89E93CFE81093EFEDB" box="[1280,1361,328,354]" italics="true" pageId="28" pageNumber="479">Cloeon</emphasis>
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<bibRefCitation id="EF8D4B45F21BFF89E964FE810CBDFE3F" author="Leach, W. E." pageId="28" pageNumber="479" pagination="57 - 172" refId="ref14907" refString="Leach, W. E. (1815) Entomology. Brewster's Edinburgh Encyclopedia, 9 (1), 57 - 172." type="journal article" year="1815">Leach 1815</bibRefCitation>
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</taxonomicName>
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, which at that time represented an artificial group and comprised various species with single marginal intercalaries on the fore wing and no hind wings. The structure of genitals figured in the original description (
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<bibRefCitation id="EF8D4B45F21BFF89E916FE590CBCFE77" author="Demoulin, G." pageId="28" pageNumber="479" pagination="225 - 241" refId="ref14512" refString="Demoulin, G. (1969) Les Ephemeropteres recoltes par la mission danoise du Noona Dan aux iles Philippines et Bismarck. Entomologiske Meddelelser, 37 (3), 225 - 241." type="journal article" year="1969">Demoulin 1969</bibRefCitation>
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: fig. 7b) clearly testify that this species belongs to
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<taxonomicName id="4C1C4D37F21BFF89EF37FE7D0FE6FE77" authorityName="Muller-Liebenau" authorityYear="1982" box="[779,905,436,462]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="genus" sensu="lato">
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<emphasis id="B968EAA6F21BFF89EF37FE7D0FE6FE77" box="[779,905,436,462]" italics="true" pageId="28" pageNumber="479">Indocloeon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57DDF21BFF89EFACFE7C0FD7FE77" box="[912,952,436,463]" pageId="28" pageNumber="479" sensu="lato">s. l.</taxonomicNameLabel>
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Judging by this figure, the distal (3rd) segment of the gonostylus is as long as the proximal (1st +2nd) segment (
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<figureCitation id="13272A31F21BFF89EF93FE11087FFE4B" box="[943,1040,471,498]" captionStart="FIGURES 101–105" captionStartId="29.[151,264,1853,1878]" captionTargetBox="[151,1427,281,1828]" captionTargetId="figure@29.[151,1436,273,1828]" captionTargetPageId="29" captionText="FIGURES 101–105. Male genitals of Hindocloeon. 101, imaginal genitals of Indocloeon (Hindocloeon) continentale sp. n. (penis dotted; 1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; gv-d, distal portion of gonovectis; gv-p, proximal portion of gonovectis; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 102, their subimaginal exuviae; 103, subimaginal gonostyli of I. (H.) sp. «Vietnam»; 104, subimaginal gonostyli of I. (H.) sp. «Java»; 105, imaginal genitals of I. longistylus (after Demoulin 1969)." figureDoi="http://doi.org/10.5281/zenodo.3839373" httpUri="https://zenodo.org/record/3839373/files/figure.png" pageId="28" pageNumber="479">Fig. 105</figureCitation>
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), and this distinguishes
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<taxonomicName id="4C1C4D37F21BFF89E922FE110D7FFDAF" authorityName="Kluge & Suttinun" authorityYear="2020" baseAuthorityName="Demoulin" baseAuthorityYear="1969" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="longistylus">
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<emphasis id="B968EAA6F21BFF89E922FE110D7FFDAF" italics="true" pageId="28" pageNumber="479">Indocloeon longistylus</emphasis>
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</taxonomicName>
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from other known species, whose distal segment is shorter than the proximal segment, both in the imaginal and subimaginal stages (
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<figureCitation id="13272A31F21BFF89EDD5FDE90EE7FD83" box="[489,648,544,570]" captionStart="FIGURES 101–105" captionStartId="29.[151,264,1853,1878]" captionTargetBox="[151,1427,281,1828]" captionTargetId="figure@29.[151,1436,273,1828]" captionTargetPageId="29" captionText="FIGURES 101–105. Male genitals of Hindocloeon. 101, imaginal genitals of Indocloeon (Hindocloeon) continentale sp. n. (penis dotted; 1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; gv-d, distal portion of gonovectis; gv-p, proximal portion of gonovectis; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 102, their subimaginal exuviae; 103, subimaginal gonostyli of I. (H.) sp. «Vietnam»; 104, subimaginal gonostyli of I. (H.) sp. «Java»; 105, imaginal genitals of I. longistylus (after Demoulin 1969)." figureDoi="http://doi.org/10.5281/zenodo.3839373" httpUri="https://zenodo.org/record/3839373/files/figure.png" pageId="28" pageNumber="479">Figs 101–104</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653FF21BFF89ECFBFD8A0823FDE6" box="[199,1100,579,607]" pageId="28" pageNumber="479" type="distribution">
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<paragraph id="8BA336B4F21BFF89ECFBFD8A0823FDE6" blockId="28.[151,1437,327,607]" box="[199,1100,579,607]" pageId="28" pageNumber="479">
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<emphasis id="B968EAA6F21BFF89ECFBFD8A0D0CFDE7" bold="true" box="[199,355,579,606]" pageId="28" pageNumber="479">Distribution:</emphasis>
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Balabac and
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<collectingRegion id="49D8F856F21BFF89EDC6FD8D0E36FDE7" box="[506,601,580,606]" country="Philippines" name="Palawan" pageId="28" pageNumber="479">Palawan</collectingRegion>
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Islands of the
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<collectingCountry id="F30B7624F21BFF89EEC2FD8D0F13FDE7" box="[766,892,580,606]" name="Philippines" pageId="28" pageNumber="479">Philippines</collectingCountry>
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(
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<bibRefCitation id="EF8D4B45F21BFF89EFB7FD8D0850FDE6" author="Demoulin, G." box="[907,1087,580,607]" pageId="28" pageNumber="479" pagination="225 - 241" refId="ref14512" refString="Demoulin, G. (1969) Les Ephemeropteres recoltes par la mission danoise du Noona Dan aux iles Philippines et Bismarck. Entomologiske Meddelelser, 37 (3), 225 - 241." type="journal article" year="1969">Demoulin 1969</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |