341 lines
40 KiB
XML
341 lines
40 KiB
XML
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<mods:title id="799C71FCFBE74CDEACCAAA1197AA149F">A review of the New Guinea species of Chimarra Stephens (Trichoptera: Philopotamidae)</mods:title>
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<mods:namePart id="E8132ADF26B17F058D7465304A0F89A4">Cartwright, David</mods:namePart>
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<mods:title id="D56A2473F36FE36F5FECFB1442153940">Memoirs of Museum Victoria</mods:title>
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<mods:date id="F12A5EEDDC17D9B4C36A084F354C65EC">2020</mods:date>
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<mods:number id="9413CC7CD2C89EC2A057B5BAD2DA70CB">79</mods:number>
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<treatment id="03B5879CB017FFBAF0CCB76AFB8DF884" ID-DOI="http://doi.org/10.5281/zenodo.8082615" ID-GBIF-Taxon="210477123" ID-Zenodo-Dep="8082615" LSID="urn:lsid:plazi:treatment:03B5879CB017FFBAF0CCB76AFB8DF884" httpUri="http://treatment.plazi.org/id/03B5879CB017FFBAF0CCB76AFB8DF884" lastPageNumber="5" pageId="4" pageNumber="5">
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<subSubSection id="C3066501B017FFBAF0CCB76AFEAAFEDC" box="[140,341,288,311]" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="8BA3368AB017FFBAF0CCB76AFEAAFEDC" blockId="4.[140,341,288,311]" box="[140,341,288,311]" pageId="4" pageNumber="5">
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<heading id="D0EB81E6B017FFBAF0CCB76AFEAAFEDC" box="[140,341,288,311]" fontSize="9" level="2" pageId="4" pageNumber="5" reason="5">
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<taxonomicName id="4C1C4D09B017FFBAF0CCB76AFEAAFEDC" ID-CoL="3N68" ID-ENA="177910" authority="Stephens" authorityName="Stephens" box="[140,341,288,311]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EA98B017FFBAF0CCB76AFF0AFEDD" bold="true" box="[140,245,288,310]" italics="true" pageId="4" pageNumber="5">Chimarra</emphasis>
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Stephens
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066501B017FFBAF0F2B707FB5DFA68" pageId="4" pageNumber="5" type="materials_examined">
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<paragraph id="8BA3368AB017FFBAF0F2B707FD8EFE8B" blockId="4.[178,625,332,352]" box="[178,625,332,352]" pageId="4" pageNumber="5">
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<emphasis id="B968EA98B017FFBAF0F2B707FEDBFEB4" box="[178,292,333,351]" italics="true" pageId="4" pageNumber="5">
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<typeStatus id="54A78828B017FFBAF0F2B707FF22FEB4" box="[178,221,333,351]" pageId="4" pageNumber="5">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="4C1C4D09B017FFBAF16FB706FE10FEB4" box="[303,495,332,351]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="marginata">
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<emphasis id="B968EA98B017FFBAF16FB706FE10FEB4" box="[303,495,332,351]" italics="true" pageId="4" pageNumber="5">Chimarra marginata</emphasis>
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</taxonomicName>
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by monotypy.
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</paragraph>
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<paragraph id="8BA3368AB017FFBAF0CCB73DFD62FE67" blockId="4.[140,669,374,397]" box="[140,669,374,397]" pageId="4" pageNumber="5">
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General features of males of New
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<collectingCountry id="F30B761AB017FFBAF1A5B73DFDD1FE66" box="[485,558,375,397]" name="Guinea-Bissau" pageId="4" pageNumber="5">Guinea</collectingCountry>
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<taxonomicName id="4C1C4D09B017FFBAF274B73CFD62FE67" box="[564,669,374,396]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B968EA98B017FFBAF274B73CFD62FE67" bold="true" box="[564,669,374,396]" italics="true" pageId="4" pageNumber="5">Chimarra</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA3368AB017FFBAF0CCB7EEFE79FD48" blockId="4.[139,773,419,1903]" pageId="4" pageNumber="5">No formal key is provided for several reasons – primarily because many characters varied considerably between species (e.g. Rs sinuous or curved, thickened ranging to straight and not thickened basad of discoidal cell in forewing, ventral process on segment IX ranged from elongate and pole-like to absent). Because of the poor condition of many of the specimens (including the wings) and at times inadequate clearing of the genitalia, some features were difficult to discern with confidence.</paragraph>
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<paragraph id="8BA3368AB017FFBAF0F2B4E0FF00FCB9" blockId="4.[139,773,419,1903]" pageId="4" pageNumber="5">Species are progressively separated throughout this paper on the basis of the L/W ratio of the ventral process on segment IX. Species were identified secondly by small differences in the shape of the inferior appendages (especially in lateral view) and the shape and position of the pair of lateral lobes on segment X.</paragraph>
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<paragraph id="8BA3368AB017FFBAF0F2B510FD92F9A0" blockId="4.[139,773,419,1903]" pageId="4" pageNumber="5">
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The first feature used was the ventral process on segment IX. The relative length and the L/W ratio are used as a basic means of separating the New
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<collectingCountry id="F30B761AB017FFBAF184B5DEFDF1FC41" box="[452,526,916,938]" name="Guinea-Bissau" pageId="4" pageNumber="5">Guinea</collectingCountry>
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<taxonomicName id="4C1C4D09B017FFBAF257B5DEFD2CFC41" box="[535,723,916,938]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B968EA98B017FFBAF257B5DEFD83FC42" box="[535,636,916,937]" italics="true" pageId="4" pageNumber="5">Chimarra</emphasis>
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species.
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</taxonomicName>
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The L/W ratio varies from
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<quantity id="4CE49B6FB017FFBAF13BB5F8FE4FFC2C" box="[379,432,945,967]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="5" unit="in" value="20.0">20 in</quantity>
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<taxonomicName id="4C1C4D09B017FFBAF1F8B5F8FDE0FC2C" authorityName="Cartwright" authorityYear="2020" box="[440,543,946,967]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pertica">
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<emphasis id="B968EA98B017FFBAF1F8B5F8FDE0FC2C" box="[440,543,946,967]" italics="true" pageId="4" pageNumber="5">C. pertica</emphasis>
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</taxonomicName>
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to 0 (absent) in many species. Having a similar ratio does not necessarily imply a relationship between the species. Only a few species (mostly from the
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<taxonomicName id="4C1C4D09B017FFBAF0AEB243FE97FBF4" authorityName="Kimmins" authorityYear="1962" box="[238,360,1033,1055]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="papuana">
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<emphasis id="B968EA98B017FFBAF0AEB243FE97FBF4" box="[238,360,1033,1055]" italics="true" pageId="4" pageNumber="5">C. papuana</emphasis>
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</taxonomicName>
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group) such as
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<taxonomicName id="4C1C4D09B017FFBAF24CB243FD8CFBF4" authorityName="Cartwright" authorityYear="2020" box="[524,627,1033,1055]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pertica">
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<emphasis id="B968EA98B017FFBAF24CB243FD8CFBF4" box="[524,627,1033,1055]" italics="true" pageId="4" pageNumber="5">C. pertica</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1C4D09B017FFBAF2C1B243FCFCFBF5" authorityName="Mey" authorityYear="2006" box="[641,771,1033,1054]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="guentheri">
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<emphasis id="B968EA98B017FFBAF2C1B243FCFCFBF5" box="[641,771,1033,1054]" italics="true" pageId="4" pageNumber="5">C. guentheri</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1C4D09B017FFBAF0FEB26DFE99FBD0" authorityName="Cartwright" authorityYear="2020" box="[190,358,1062,1084]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ukarumpana">
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<emphasis id="B968EA98B017FFBAF0FEB26DFE99FBD0" box="[190,358,1062,1084]" italics="true" pageId="4" pageNumber="5">C. ukarumpana</emphasis>
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</taxonomicName>
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have an elongate and slender ventral process. Most species have a triangular or rounded keel or no obvious process. In the region, most Australian species have no obvious keel or only a small triangular one (
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<bibRefCitation id="EF8D4B7BB017FFBAF2CBB234FF3CFB5A" author="Cartwright, D. I." pageId="4" pageNumber="5" pagination="393 - 437" refId="ref37070" refString="Cartwright, D. I. 2002. The Australian species of Chimarra Stephens (Trichoptera: Philopotamidae). Memoirs of Museum Victoria. 59: 393 - 437." type="journal article" year="2002">Cartwright, 2002</bibRefCitation>
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). Of the nearby
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<collectingCountry id="F30B761AB017FFBAF12EB2D1FDE7FB5A" box="[366,536,1179,1201]" name="Solomon Islands" pageId="4" pageNumber="5">Solomon Islands</collectingCountry>
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species, most have no obvious keel but
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<taxonomicName id="4C1C4D09B017FFBAF17CB2F2FCFDFB25" authority="Johanson and Espeland" authorityName="Johanson and Espeland" box="[316,770,1208,1230]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="talinensis">
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<emphasis id="B968EA98B017FFBAF17CB2F2FDF3FB26" box="[316,524,1208,1229]" italics="true" pageId="4" pageNumber="5">Chimarra talinensis</emphasis>
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Johanson and Espeland
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</taxonomicName>
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has an elongate and robust ventral process (
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<bibRefCitation id="EF8D4B7BB017FFBAF231B29CFED4FAE2" author="Johanson, K. A. & Espeland, M." pageId="4" pageNumber="5" pagination="25 - 43" refId="ref37190" refString="Johanson, K. A., and Espeland, M. 2010. Description of new Chimarra (Trichoptera: Philopotamidae) species from the Solomon Islands. Zootaxa 2638: 25 - 43." type="journal article" year="2010">Johanson and Espeland, 2010</bibRefCitation>
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), while in 24 species from
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<collectingCountry id="F30B761AB017FFBAF205B2B9FD98FAE2" box="[581,615,1267,1289]" name="Fiji" pageId="4" pageNumber="5">Fiji</collectingCountry>
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(
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<bibRefCitation id="EF8D4B7BB017FFBAF236B2B9FF02FACD" author="Johanson, K. A. & Olah, J." pageId="4" pageNumber="5" pagination="1 - 58" refId="ref37226" refString="Johanson, K. A., and Olah, J. 2012. Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species. Zootaxa 3354: 1 - 58." type="journal article" year="2012">Johanson and Oláh, 2012</bibRefCitation>
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) and three species from
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<collectingCountry id="F30B761AB017FFBAF1BFB35BFDABFACD" box="[511,596,1297,1318]" name="Vanuatu" pageId="4" pageNumber="5">Vanuatu</collectingCountry>
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(
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<bibRefCitation id="EF8D4B7BB017FFBAF222B35AFF41FAAF" author="Johanson, K. A. & Wells, A. & Malm, T. & Espeland, M." pageId="4" pageNumber="5" pagination="279 - 320" refId="ref37264" refString="Johanson, K. A., Wells, A., Malm, T., and Espeland, M. 2011. The Trichoptera of Vanuatu. Deutsche Entomologische Zeitschrift 58: 279 - 320." type="journal article" year="2011">Johanson et al., 2011</bibRefCitation>
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), none has an obvious ventral process. Of the south-east Asian species, most have no obvious process or only a small keel; a few have a more pointed and elongate process, including
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<taxonomicName id="4C1C4D09B017FFBAF0B4B3CCFD01FA70" authority="Chantaramongkol and Malicky" authorityName="Chantaramongkol and Malicky" box="[244,766,1413,1435]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chiangmaiensis">
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<emphasis id="B968EA98B017FFBAF0B4B3CCFE4CFA71" box="[244,435,1413,1435]" italics="true" pageId="4" pageNumber="5">C. chiangmaiensis</emphasis>
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Chantaramongkol and Malicky
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</taxonomicName>
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,
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<taxonomicName id="4C1C4D09B017FFBAF0CEB3E9FD81FA53" authority="Chantaramongkol and Malicky" authorityName="Chantaramongkol and Malicky" box="[142,638,1442,1464]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="khamuorum">
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<emphasis id="B968EA98B017FFBAF0CEB3E9FED3FA5C" box="[142,300,1442,1464]" italics="true" pageId="4" pageNumber="5">C. khamuorum</emphasis>
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Chantaramongkol and Malicky
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</taxonomicName>
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,
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<taxonomicName id="4C1C4D09B017FFBAF2CDB3E9FE57FA3D" authority="Malicky (Malicky, 2010)" authorityName="Malicky (Malicky" authorityYear="2010" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="demeter">
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<emphasis id="B968EA98B017FFBAF2CDB3E9FCFCFA5C" box="[653,771,1442,1464]" italics="true" pageId="4" pageNumber="5">C. demeter</emphasis>
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Malicky (
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<bibRefCitation id="EF8D4B7BB017FFBAF0BDB38AFE5EFA3D" author="Malicky, H." box="[253,417,1472,1494]" pageId="4" pageNumber="5" refId="ref37500" refString="Malicky, H. 2010. Atlas of southeast Asian Trichoptera. Biology Department, Faculty of Science, Chiang Mai University: Chiang Mai, Thailand. 346 pp." type="book" year="2010">Malicky, 2010</bibRefCitation>
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)
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</taxonomicName>
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. In Borneo, species in
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<collectingRegion id="49D8F868B017FFBAF296B38AFEFAFA18" country="Indonesia" name="Kalimantan Timur" pageId="4" pageNumber="5">East Kalimantan</collectingRegion>
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(
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<collectingCountry id="F30B761AB017FFBAF155B397FE85FA18" box="[277,378,1501,1523]" name="Indonesia" pageId="4" pageNumber="5">Indonesia</collectingCountry>
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) and
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<collectingRegion id="49D8F868B017FFBAF1F9B397FE08FA18" box="[441,503,1501,1523]" country="Malaysia" name="Sabah" pageId="4" pageNumber="5">Sabah</collectingRegion>
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(
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<collectingCountry id="F30B761AB017FFBAF247B397FD99FA18" box="[519,614,1501,1523]" name="Malaysia" pageId="4" pageNumber="5">Malaysia</collectingCountry>
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) have elongate ventral processes, including
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<taxonomicName id="4C1C4D09B017FFBAF1ECB3B0FD02F9FB" authority="Blahnik et al." authorityName="Blahnik" box="[428,765,1530,1552]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="devogeli">
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<emphasis id="B968EA98B017FFBAF1ECB3B0FD92F9E4" box="[428,621,1530,1551]" italics="true" pageId="4" pageNumber="5">Chimarra devogeli</emphasis>
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Blahnik et al.
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</taxonomicName>
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,
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<taxonomicName id="4C1C4D09B017FFBAF0CCB052FE69F9C6" authority="Blahnik et al." authorityName="Blahnik" box="[140,406,1559,1581]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="drepane">
|
||
<emphasis id="B968EA98B017FFBAF0CCB052FF00F9C7" box="[140,255,1559,1581]" italics="true" pageId="4" pageNumber="5">C. drepane</emphasis>
|
||
Blahnik et al.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF1E5B052FD49F9C6" authority="Blahnik et al." authorityName="Blahnik" box="[421,694,1559,1581]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fuilianae">
|
||
<emphasis id="B968EA98B017FFBAF1E5B052FDDDF9C7" box="[421,546,1559,1581]" italics="true" pageId="4" pageNumber="5">C. fuilianae</emphasis>
|
||
Blahnik et al.
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D09B017FFBAF2ABB052FD96F9A0" authority="Blahnik et al. (Blahnik et al., 2009)" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="xiphosella">
|
||
<emphasis id="B968EA98B017FFBAF2ABB052FF09F9A1" italics="true" pageId="4" pageNumber="5">C. xiphosella</emphasis>
|
||
Blahnik et al. (
|
||
<bibRefCitation id="EF8D4B7BB017FFBAF1D3B07FFD9EF9A0" author="Blahnik, R. J. & Holzenthal, R. W. & Huisman, J." box="[403,609,1589,1611]" pageId="4" pageNumber="5" pagination="109 - 166" refId="ref36936" refString="Blahnik, R. J., Holzenthal, R. W., and Huisman, J. 2009. Chimarra of Sabah and Sarawak, northern Borneo (Trichoptera: Philopotamidae). Tijdschrift voor Entomologie 152: 109 - 166." type="journal article" year="2009">Blahnik et al., 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA3368AB017FFBAF0F2B018FBE2FEDE" blockId="4.[139,773,419,1903]" lastBlockId="4.[809,1440,229,454]" pageId="4" pageNumber="5">
|
||
The second character used here for species separation is the shape of the inferior appendages (predominantly as viewed laterally). The worldwide range of variation in this structure is remarkable among
|
||
<taxonomicName id="4C1C4D09B017FFBAF198B0E0FD66F92B" box="[472,665,1706,1728]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B968EA98B017FFBAF198B0E0FDC2F954" box="[472,573,1706,1727]" italics="true" pageId="4" pageNumber="5">Chimarra</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Although most New
|
||
<collectingCountry id="F30B761AB017FFBAF0BFB08DFEB6F936" box="[255,329,1735,1757]" name="Guinea-Bissau" pageId="4" pageNumber="5">Guinea</collectingCountry>
|
||
species have distinctively shaped inferior appendages, several exhibit slight variations on a pattern of otherwise almost uniformly elongate sub-triangular inferior appendages (in lateral view), making it difficult to distinguish them from each other using this character. Many of the species have inferior appendages with acute apices that are inflexed or directed posteromesally and so are hidden in the lateral view; instead, these appendages may appear slightly truncate or abbreviated.
|
||
</paragraph>
|
||
<paragraph id="8BA3368AB017FFBAF30FB776FA90FE2D" blockId="4.[809,1440,229,454]" pageId="4" pageNumber="5">
|
||
Other characteristics of interest include the form of the lateral lobes of segment X, the shape of preanal appendages on segment IX, the
|
||
<taxonomicName id="4C1C4D09B017FFBAF3B0B73CFB9DFE67" box="[1008,1122,374,396]" pageId="4" pageNumber="5" rank="variety" variety="and">variety and</taxonomicName>
|
||
shape of phallic structures, the presence or absence of obvious sensilla on the lateral processes of segment X, and variations in wing venation.
|
||
</paragraph>
|
||
<paragraph id="8BA3368AB017FFBAF369B796FA5FFCF8" blockId="4.[809,1441,476,787]" pageId="4" pageNumber="5">
|
||
<emphasis id="B968EA98B017FFBAF369B796FABCFE1A" box="[809,1347,476,497]" italics="true" pageId="4" pageNumber="5">
|
||
General characteristics of the New
|
||
<collectingCountry id="F30B761AB017FFBAF4CFB796FB25FE1A" box="[1167,1242,476,497]" name="Guinea-Bissau" pageId="4" pageNumber="5">Guinea</collectingCountry>
|
||
<taxonomicName id="4C1C4D09B017FFBAF49FB796FABCFE1A" box="[1247,1347,476,497]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Chimarra</taxonomicName>
|
||
</emphasis>
|
||
. General body colour and wings brownish (unless faded with time in alcohol). Spur formula 1:4:4. Small to medium-sized adults. Forewing length range, males: 3.5–
|
||
<quantity id="4CE49B6FB017FFBAF4C9B478FB3BFDA3" box="[1161,1220,562,584]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="5" unit="mm" value="7.0">7 mm</quantity>
|
||
, more commonly
|
||
<quantity id="4CE49B6FB017FFBAF538B479FCABFD8E" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="4" pageNumber="5" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
|
||
. Forewing often with Rs sinuous or curved, thickened (sometimes straight and not thickened) basad of discoidal cell, occasionally with small, clear, depressed window (fig. 63), forks 1, 2, 3 and 5 present; hind wing with forks 1 (usually), 2, 3 and 5 present (fig. 7). At least two species –
|
||
<taxonomicName id="4C1C4D09B017FFBAF4B6B48EFA8AFD33" authorityName="Kimmins" authorityYear="1962" box="[1270,1397,707,729]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cyclopica">
|
||
<emphasis id="B968EA98B017FFBAF4B6B48EFA8AFD33" box="[1270,1397,707,729]" italics="true" pageId="4" pageNumber="5">C. cyclopica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D09B017FFBAF369B4ABFC6EFD1E" authorityName="Cartwright" authorityYear="2020" box="[809,913,736,758]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aliceae">
|
||
<emphasis id="B968EA98B017FFBAF369B4ABFC6EFD1E" box="[809,913,736,758]" italics="true" pageId="4" pageNumber="5">C. aliceae</emphasis>
|
||
</taxonomicName>
|
||
– appear to have fork 1 absent on the hind wing, as originally noted by Kimmins for
|
||
<taxonomicName id="4C1C4D09B017FFBAF432B4B7FA63FCF8" authority="(Kimmins 1962)" baseAuthorityName="Kimmins" baseAuthorityYear="1962" box="[1138,1436,765,787]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cyclopica">
|
||
<emphasis id="B968EA98B017FFBAF432B4B7FB10FCF9" box="[1138,1263,765,786]" italics="true" pageId="4" pageNumber="5">C. cyclopica</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8D4B7BB017FFBAF4BAB4B7FA6AFCF8" author="Kimmins, D. E." box="[1274,1429,765,787]" pageId="4" pageNumber="5" pagination="99 - 187" refId="ref37347" refString="Kimmins, D. E. 1962. Miss L. E. Cheeseman's expeditions to New Guinea. Trichoptera. Bulletin of the British Museum (Natural History), Entomology 11: 99 - 187." type="journal article" year="1962">Kimmins 1962</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA3368AB017FFBAF368B563FAE2FB96" blockId="4.[808,1441,809,1149]" pageId="4" pageNumber="5">
|
||
<emphasis id="B968EA98B017FFBAF368B563FCA3FCD5" box="[808,860,809,830]" italics="true" pageId="4" pageNumber="5">Male</emphasis>
|
||
. Segment IX anterior margin ventral basally usually welldeveloped, often rounded or V-shaped; ventral process usually present, rarely very long (figs 1, 4), more usually short, keel-like (figs 70, 78, 132) or without obvious process (figs 155, 158). Preanal appendages usually short, rounded. Segment X with mesal lobe membranous, reduced and pair of more heavily sclerotised lateral lobes, mostly adpressed to phallus and without short hair-like sensilla discerned. Phallus generally tubular, phallobase expanded, rounded, usually with a pair of short, slender, straight or slightly curved spines included subapically, more rarely with only one or without obvious spines. Inferior appendages one-segmented, shape highly variable.
|
||
</paragraph>
|
||
<paragraph id="8BA3368AB017FFBAF369B2D9FA96FB2D" blockId="4.[809,1440,1171,1222]" pageId="4" pageNumber="5">
|
||
<emphasis id="B968EA98B017FFBAF369B2D9FC8BFB43" box="[809,884,1171,1192]" italics="true" pageId="4" pageNumber="5">Female</emphasis>
|
||
. Females have rarely been associated for any New
|
||
<collectingCountry id="F30B761AB017FFBAF369B2FAFC8DFB2D" box="[809,882,1200,1222]" name="Guinea-Bissau" pageId="4" pageNumber="5">Guinea</collectingCountry>
|
||
species and were not examined during this study.
|
||
</paragraph>
|
||
<paragraph id="8BA3368AB017FFBAF369B296FB5DFA68" blockId="4.[809,1441,1244,1903]" pageId="4" pageNumber="5">
|
||
<emphasis id="B968EA98B017FFBAF369B296FC7DFB1A" box="[809,898,1244,1265]" italics="true" pageId="4" pageNumber="5">Remarks</emphasis>
|
||
. No new material was seen for most of the previously described New
|
||
<collectingCountry id="F30B761AB017FFBAF384B2B3FBF2FAE4" box="[964,1037,1273,1295]" name="Guinea-Bissau" pageId="4" pageNumber="5">Guinea</collectingCountry>
|
||
species. During this study, new material was examined for the following nine species:
|
||
<taxonomicName id="4C1C4D09B017FFBAF572B35DFA65FAC7" authorityName="Korboot" authorityYear="1965" box="[1330,1434,1303,1324]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aiyura">
|
||
<emphasis id="B968EA98B017FFBAF572B35DFA65FAC7" box="[1330,1434,1303,1324]" italics="true" pageId="4" pageNumber="5">C. aiyura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF369B37EFC59FAA3" authorityName="Kimmins" authorityYear="1957" box="[809,934,1331,1353]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="biramosa">
|
||
<emphasis id="B968EA98B017FFBAF369B37EFC59FAA3" box="[809,934,1331,1353]" italics="true" pageId="4" pageNumber="5">C. biramosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF3F1B37EFBD0FAA3" baseAuthorityName="Kimmins" baseAuthorityYear="1962" box="[945,1071,1331,1353]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cyclopica">
|
||
<emphasis id="B968EA98B017FFBAF3F1B37EFBD0FAA3" box="[945,1071,1331,1353]" italics="true" pageId="4" pageNumber="5">C. cyclopica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF47BB37EFB61FAA3" authorityName="Kimmins" authorityYear="1962" box="[1083,1182,1331,1353]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="falcata">
|
||
<emphasis id="B968EA98B017FFBAF47BB37EFB61FAA3" box="[1083,1182,1331,1353]" italics="true" pageId="4" pageNumber="5">C. falcata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF4EAB37EFAF2FAA2" authorityName="Sykora" authorityYear="1967" box="[1194,1293,1332,1353]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="goroca">
|
||
<emphasis id="B968EA98B017FFBAF4EAB37EFAF2FAA2" box="[1194,1293,1332,1353]" italics="true" pageId="4" pageNumber="5">C. goroca</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF559B37EFA66FAA3" authorityName="Kimmins" authorityYear="1962" box="[1305,1433,1331,1353]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kokodana">
|
||
<emphasis id="B968EA98B017FFBAF559B37EFA66FAA3" box="[1305,1433,1331,1353]" italics="true" pageId="4" pageNumber="5">C. kokodana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B968EA98B017FFBAF369B31BFBE2FA8D" box="[809,1053,1360,1382]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C1C4D09B017FFBAF369B31BFC56FA8E" authorityName="Sykora" authorityYear="1967" box="[809,937,1360,1382]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sedlaceki">C. sedlaceki</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF3F2B31BFBE2FA8D" authorityName="Kimmins" authorityYear="1962" box="[946,1053,1361,1382]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sinuosa">C. sinuosa</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C1C4D09B017FFBAF40EB31BFB51FA8E" authorityName="Kimmins" authorityYear="1962" box="[1102,1198,1360,1382]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ulmeri">
|
||
<emphasis id="B968EA98B017FFBAF40EB31BFB51FA8E" box="[1102,1198,1360,1382]" italics="true" pageId="4" pageNumber="5">C. ulmeri</emphasis>
|
||
</taxonomicName>
|
||
. These are re-described and significant parts are figured here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066501B017FFBAF30FB3C0FB8DF884" pageId="4" pageNumber="5" type="reference_group">
|
||
<paragraph id="8BA3368AB017FFBAF30FB3C0FB8DF884" blockId="4.[809,1441,1244,1903]" pageId="4" pageNumber="5">
|
||
A
|
||
<emphasis id="B968EA98B017FFBAF330B3C1FC34FA4B" box="[880,971,1419,1440]" italics="true" pageId="4" pageNumber="5">papuana</emphasis>
|
||
species group comprising eight species –
|
||
<taxonomicName id="4C1C4D09B017FFBAF369B3EDFC41FA56" authority="Olah" authorityName="Olah" authorityYear="2013" box="[809,958,1447,1469]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="agasa">
|
||
<emphasis id="B968EA98B017FFBAF369B3EDFC7BFA56" box="[809,900,1447,1469]" italics="true" pageId="4" pageNumber="5">C. agasa</emphasis>
|
||
Oláh
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF388B3EDFB9DFA56" authority="Olah" authorityName="Olah" authorityYear="2012" box="[968,1122,1447,1469]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bobita">
|
||
<emphasis id="B968EA98B017FFBAF388B3EDFBD7FA57" box="[968,1064,1447,1468]" italics="true" pageId="4" pageNumber="5">C. bobita</emphasis>
|
||
Oláh
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF42CB3EDFB00FA56" authority="Olah" authorityName="Olah" authorityYear="1993" box="[1132,1279,1447,1469]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kalija">
|
||
<emphasis id="B968EA98B017FFBAF42CB3EDFB3AFA57" box="[1132,1221,1447,1468]" italics="true" pageId="4" pageNumber="5">C. kalija</emphasis>
|
||
Oláh
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF549B3EDFC78FA32" authority="Olah and Mey" authorityName="Olah and Mey" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kozela">
|
||
<emphasis id="B968EA98B017FFBAF549B3EDFA97FA57" box="[1289,1384,1447,1468]" italics="true" pageId="4" pageNumber="5">C. kozela</emphasis>
|
||
Oláh and Mey
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF3D5B38EFB80FA31" ID-CoL="7S7XR" authority="Kimmins" authorityName="Kimmins" authorityYear="1962" box="[917,1151,1476,1498]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis id="B968EA98B017FFBAF3D5B38EFBEEFA31" box="[917,1041,1476,1498]" italics="true" pageId="4" pageNumber="5">C. papuana</emphasis>
|
||
Kimmins
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF4CCB38EFACEFA31" authority="Olah" authorityName="Olah" box="[1164,1329,1476,1498]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="porsen">
|
||
<emphasis id="B968EA98B017FFBAF4CCB38EFB0DFA31" box="[1164,1266,1476,1498]" italics="true" pageId="4" pageNumber="5">C. porsen</emphasis>
|
||
Oláh
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF57EB38EFCA5FA1C" authority="Olah" authorityName="Olah" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tompa">
|
||
<emphasis id="B968EA98B017FFBAF57EB38EFA5FFA31" box="[1342,1440,1476,1498]" italics="true" pageId="4" pageNumber="5">C. tompa</emphasis>
|
||
Oláh
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D09B017FFBAF3CFB3ABFBE6FA1C" authority="Olah" authorityName="Olah" box="[911,1049,1505,1527]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tulok">
|
||
<emphasis id="B968EA98B017FFBAF3CFB3ABFC1EFA1D" box="[911,993,1505,1526]" italics="true" pageId="4" pageNumber="5">C. tulok</emphasis>
|
||
Oláh
|
||
</taxonomicName>
|
||
– was recognised by
|
||
<bibRefCitation id="EF8D4B7BB017FFBAF4BBB3ABFA8BFA1C" author="Mey, W." box="[1275,1396,1505,1527]" pageId="4" pageNumber="5" pagination="261 - 263" refId="ref37604" refString="Mey, W. 2006. Chimarra guentheri sp. nov. - a new species from New Guinea (Insecta, Trichoptera) Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe 82: 261 - 263." type="journal article" year="2006">Mey (2006)</bibRefCitation>
|
||
and further refined by
|
||
<bibRefCitation id="EF8D4B7BB017FFBAF3A5B3B4FB9FF9FF" author="Olah, J." box="[997,1120,1534,1556]" pageId="4" pageNumber="5" pagination="91 - 131" refId="ref38109" refString="Olah, J. 2014. On the Trichoptera of Batanta Island (Indonesia, West Papua, Raja Ampat Archipelago). III. Folio Entomologica Hungarica Rovartani Kozlemenyek 75: 91 - 131." type="journal article" year="2014">Oláh (2014)</bibRefCitation>
|
||
. In this study, four new species (
|
||
<taxonomicName id="4C1C4D09B017FFBAF371B051FC5EF9DB" authority="Cartwright, 2020" authorityName="Cartwright" authorityYear="2020" box="[817,929,1563,1584]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bintang" status="sp. nov.">
|
||
<emphasis id="B968EA98B017FFBAF371B051FC5EF9DB" box="[817,929,1563,1584]" italics="true" pageId="4" pageNumber="5">C. bintang</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57E3B017FFBAF3EAB051FC03F9DB" box="[938,1020,1563,1584]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF449B051FB6FF9DB" authority="Cartwright, 2020" authorityName="Cartwright" authorityYear="2020" box="[1033,1168,1563,1584]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mendiana" status="sp. nov.">
|
||
<emphasis id="B968EA98B017FFBAF449B051FB6FF9DB" box="[1033,1168,1563,1584]" italics="true" pageId="4" pageNumber="5">C. mendiana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57E3B017FFBAF4DAB051FB13F9DB" box="[1178,1260,1563,1584]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C1C4D09B017FFBAF4B9B051FA85F9DB" authority="Cartwright, 2020" authorityName="Cartwright" authorityYear="2020" box="[1273,1402,1563,1584]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="milneana" status="sp. nov.">
|
||
<emphasis id="B968EA98B017FFBAF4B9B051FA85F9DB" box="[1273,1402,1563,1584]" italics="true" pageId="4" pageNumber="5">C. milneana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57E3B017FFBAF5C4B051FCABF9A6" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C1C4D09B017FFBAF3C7B072FBD6F9A6" authority="Cartwright, 2020" authorityName="Cartwright" authorityYear="2020" box="[903,1065,1592,1613]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ukarumpana" status="sp. nov.">
|
||
<emphasis id="B968EA98B017FFBAF3C7B072FBD6F9A6" box="[903,1065,1592,1613]" italics="true" pageId="4" pageNumber="5">C. ukarumpana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57E3B017FFBAF46FB072FB82F9A6" box="[1071,1149,1592,1613]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) are aligned with the group, primarily on the basis of the unique characteristic of the long and curved filiform dorsoapical process on the inferior appendages, supported by the generally elongate ventral process on segment IX. Due to the poor condition of many of the dried BPBM specimens examined during this study, it is not possible to check and confirm all of the group characters, especially some characters discussed by
|
||
<bibRefCitation id="EF8D4B7BB017FFBAF486B149FAC1F8F3" author="Mey, W." box="[1222,1342,1794,1816]" pageId="4" pageNumber="5" pagination="261 - 263" refId="ref37604" refString="Mey, W. 2006. Chimarra guentheri sp. nov. - a new species from New Guinea (Insecta, Trichoptera) Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe 82: 261 - 263." type="journal article" year="2006">Mey (2006)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF8D4B7BB017FFBAF52FB148FC93F8DE" author="Olah, J." pageId="4" pageNumber="5" pagination="91 - 131" refId="ref38109" refString="Olah, J. 2014. On the Trichoptera of Batanta Island (Indonesia, West Papua, Raja Ampat Archipelago). III. Folio Entomologica Hungarica Rovartani Kozlemenyek 75: 91 - 131." type="journal article" year="2014">Oláh (2014)</bibRefCitation>
|
||
. No clear groupings were discerned among most of the New
|
||
<collectingCountry id="F30B761AB017FFBAF320B176FC55F8B9" box="[864,938,1852,1874]" name="Guinea-Bissau" pageId="4" pageNumber="5">Guinea</collectingCountry>
|
||
species; therefore, none of the species below are formally or informally grouped.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |