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<mods:title id="F545F52FAE8E9E7713E75868FF6795E7">Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species</mods:title>
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<mods:namePart id="03D173B471CE39CFE12A818395D55F2B">Chirivi-Joya, Daniel</mods:namePart>
<mods:affiliation id="213125416C9835EA51A3E63E3B25E7A6">Laboratório de Entomología, Departamento de Biología, Facultad de Ciencias, Pontificia Universidad Javeriana. Carrera 7 N °. 40 - 62, Bogotá. &amp; dhipnosodin @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1198 - 782 X</mods:affiliation>
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<mods:namePart id="248258506180C922AFC513786837BA46">Moreno-González, Jairo A.</mods:namePart>
<mods:affiliation id="4B9D34702754E53411F293E023BD964D">Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo. Rua do Matão, trav. 14, n ° 321, Cidade Universitária, São Paulo- SP, CEP: 05508 - 090. hansenochrus @ gmail. com; https: // orcid. org / 0000 - 0001 - 5980 - 1417</mods:affiliation>
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<mods:namePart id="C4E12EB0D95EC2EB18AE335A8241DFD1">Fagua, Giovanny</mods:namePart>
<mods:affiliation id="3DC78C77D35BEA5DAA85ACBB3C884ADD">Laboratório de Entomología, Departamento de Biología, Facultad de Ciencias, Pontificia Universidad Javeriana. Carrera 7 N °. 40 - 62, Bogotá. &amp; fagua @ javeriana. edu. co; https: // orcid. org / 0000 - 0003 - 1253 - 4416</mods:affiliation>
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<taxonomicName id="4C1C4D03B65AFFCCFF7EFD1CFDF074AE" ID-CoL="6LSFJ" authority="Pocock" authorityName="Pocock" authorityYear="1902" box="[151,535,693,720]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="armiger">
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<emphasis id="B968EA92B65AFFCCFF7EFD1CFE5274AE" bold="true" box="[151,437,693,720]" italics="true" pageId="8" pageNumber="9">Heterophrynus armiger</emphasis>
Pocock
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<figureCitation id="13272A05B65AFFCCFF7EFD70FEF87492" box="[151,287,729,755]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="9.[151,250,1937,1962]" captionStartId-1="10.[151,250,1681,1706]" captionStartId-2="11.[151,250,1755,1780]" captionStartId-3="13.[151,250,670,695]" captionTargetBox-0="[161,1427,351,1902]" captionTargetBox-1="[161,1427,355,1644]" captionTargetBox-2="[155,1433,529,1726]" captionTargetBox-3="[152,1435,181,644]" captionTargetId-0="figure@9.[151,1436,345,1912]" captionTargetId-1="figure@10.[151,1436,345,1656]" captionTargetId-2="figure@11.[151,1436,525,1730]" captionTargetId-3="figure@13.[152,1436,181,644]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="13" captionText-0="FIGURE 4. Heterophrynus armiger. Male (IAvH-I 2924): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." captionText-1="FIGURE 5. Heterophrynus armiger. Male (IAvH-I 2924): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." captionText-2="FIGURE 6. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MUSENUV 24303), black arrow indicates the sclerotized area of the seminal receptacle." captionText-3="FIGURE 7. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. Spine over LoL1, dorsal view. B. Detail of spine, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.3907909" figureDoi-1="http://doi.org/10.5281/zenodo.3907911" figureDoi-2="http://doi.org/10.5281/zenodo.3907913" figureDoi-3="http://doi.org/10.5281/zenodo.3907916" httpUri-0="https://zenodo.org/record/3907909/files/figure.png" httpUri-1="https://zenodo.org/record/3907911/files/figure.png" httpUri-2="https://zenodo.org/record/3907913/files/figure.png" httpUri-3="https://zenodo.org/record/3907916/files/figure.png" pageId="8" pageNumber="9">Figures 47</figureCitation>
,
<figureCitation id="13272A05B65AFFCCFEC3FD70FEA17492" box="[298,326,729,755]" captionStart="FIGURE 22" captionStartId="33.[151,250,1654,1679]" captionTargetBox="[185,1402,574,1621]" captionTargetId="figure@33.[177,1410,561,1629]" captionTargetPageId="33" captionText="FIGURE 22. Distribution map. Heterophrynus alces: purple circle. Heterophrynus armiger: green circle. Heterophrynus cheiracanthus: blue circle. Heterophrynus elaphus: orange circle. Heterophrynus awa sp. nov.: gray circle. Heterophrynus origamii sp. nov.: red circle." figureDoi="http://doi.org/10.5281/zenodo.3907947" httpUri="https://zenodo.org/record/3907947/files/figure.png" pageId="8" pageNumber="9">22</figureCitation>
(in part).
<tableCitation id="C69E033BB65AFFCCFE5FFD70FDEF7492" box="[438,520,729,755]" captionStart="TABLE 2" captionStartId="12.[151,239,874,898]" captionText="TABLE 2. Measurements (mm) of the body, pedipalp segments, and femora of the legs in four specimens of Heterophrynus armiger. Empty cells correspond to missing segment or data." pageId="8" pageNumber="9">Table 2</tableCitation>
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<treatmentCitation id="0ABD1091B65AFFCCFF7EFCB4FD677554" author="Pocock, R. I." box="[151,640,796,821]" page="163" pageId="8" pageNumber="9" year="1902">
<taxonomicName id="4C1C4D03B65AFFCCFF7EFCB4FD677554" ID-CoL="6LSFJ" authority="Pocock, 1902: 162 - 163" authorityName="Pocock" authorityPageNumber="162 - 163" authorityYear="1902" box="[151,640,796,821]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="B968EA92B65AFFCCFF7EFCB4FE607554" box="[151,391,797,821]" italics="true" pageId="8" pageNumber="9">Heterophrynus armiger</emphasis>
<bibRefCitation id="EF8D4B71B65AFFCCFE64FCB5FD677554" author="Pocock, R. I." box="[397,640,796,821]" pageId="8" pageNumber="9" pagination="1 - 71" refId="ref23423" refString="Pocock, R. I. (1902) Arachnida: Scorpiones, Pedipalpi &amp; Solifugae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali- Americana. Vol. 3. Taylor and Francis, London, pp. 1 - 71." type="book chapter" year="1902">Pocock, 1902: 162163</bibRefCitation>
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;
<treatmentCitation id="0ABD1091B65AFFCCFD62FCB5FCA27554" author="Pocock, R. I." box="[651,837,796,821]" page="220" pageId="8" pageNumber="9" year="1903">
<bibRefCitation id="EF8D4B71B65AFFCCFD62FCB5FCA27554" author="Pocock, R. I." box="[651,837,796,821]" pageId="8" pageNumber="9" pagination="220 - 226" refId="ref23478" refString="Pocock, R. I. (1903) Descriptions of four new Arachnida of the orders Pedipalpi, Solifugae, and Araneae. Annals and Magazine of Natural History, Series 7, 11, 220 - 226. https: // doi. org / 10.1080 / 00222930308678753" type="journal article" year="1903">Pocock 1903: 220</bibRefCitation>
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;
<treatmentCitation id="0ABD1091B65AFFCCFCB9FCB5FBDF7554" author="Mello-Leitao, C." box="[848,1080,796,821]" page="47" pageId="8" pageNumber="9" year="1931">
<bibRefCitation id="EF8D4B71B65AFFCCFCB9FCB5FBDF7554" author="Mello-Leitao, C." box="[848,1080,796,821]" pageId="8" pageNumber="9" pagination="7 - 72" refId="ref23079" refString="Mello-Leitao, C. (1931) Pedipalpos do Brasil e algumas notas sobre a ordem. Archivos do Museu Nacional, 33, 7 - 72." type="journal article" year="1931">Mello-Leitão 1931: 47</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EF8D4B71B65AFFCCFBABFCB5FB067554" author="Weygoldt, P." box="[1090,1249,796,821]" pageId="8" pageNumber="9" pagination="95 - 107" refId="ref24153" refString="Weygoldt, P. (1972) Geisselskorpione und Geisselspinnen (Uropygi und Amblypygi). Zeitschrift des Kolner Zoo, 15, 95 - 107." type="journal article" year="1972">Weygoldt 1972</bibRefCitation>
: fig. 10;
<bibRefCitation id="EF8D4B71B65AFFCCFAD3FCB5FF1A7535" author="Weygoldt, P." pageId="8" pageNumber="9" pagination="175 - 191" refId="ref24180" refString="Weygoldt, P. (1974) Vergleichende Untersuchungen au zwei Heterophrynus (Admetus) - Arten, H. longicornis Butler und H. batesii (Arachnida, Amblypygi, Tarantulidae). Zoologischer Anzeiger, 192, 175 - 191." type="journal article" year="1974">Weygoldt 1974</bibRefCitation>
: figs 1, 4;
<treatmentCitation id="0ABD1091B65AFFCCFE87FC92FDA77535" author="Giupponi, A." box="[366,576,827,852]" page="173" pageId="8" pageNumber="9" year="2002">
<bibRefCitation id="EF8D4B71B65AFFCCFE87FC92FDA77535" author="Giupponi, A." box="[366,576,827,852]" pageId="8" pageNumber="9" pagination="173 - 175" refId="ref22531" refString="Giupponi, A. (2002) duas novas sinonimias no genero Heterophrynus Pocock, 1894 (Amblypygi, Phrynidae). Revista Iberica de Aracnologia, 6, 173 - 175." type="journal article" year="2002">Giupponi 2002: 173</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD1091B65AFFCCFDA2FC92FCF87535" author="Weygoldt, P." box="[587,799,827,852]" page="298" pageId="8" pageNumber="9" year="2002">
<bibRefCitation id="EF8D4B71B65AFFCCFDA2FC92FCF87535" author="Weygoldt, P." box="[587,799,827,852]" pageId="8" pageNumber="9" pagination="293 - 302" refId="ref24405" refString="Weygoldt, P. (2002) Amblypygi. In: Adis, J. (Ed.), Amazonian Arachnida and Myriapoda. Pensoft Publishers, Sofia, Moscow, pp. 293 - 302." type="book chapter" year="2002">Weygoldt 2002: 298</bibRefCitation>
</treatmentCitation>
, fig 16;
<treatmentCitation id="0ABD1091B65AFFCCFC9EFC95FB457535" author="Viquez, C. &amp; Chirivi, D &amp; Moreno-Gonzalez, J. &amp; Christensen, J." box="[887,1186,827,852]" page="457" pageId="8" pageNumber="9" year="2014">
<bibRefCitation id="EF8D4B71B65AFFCCFC9EFC95FB457535" author="Viquez, C. &amp; Chirivi, D &amp; Moreno-Gonzalez, J. &amp; Christensen, J." box="[887,1186,827,852]" pageId="8" pageNumber="9" pagination="457 - 460" refId="ref24025" refString="Viquez, C., Chirivi, D, Moreno-Gonzalez, J. &amp; Christensen, J. (2014) Heterophrynus armiger Pocock, 1902 (Amblypygi: Phrynidae): First record from Colombia, with notes on its historic distribution records and natural history. Check List, 10 (2), 457 - 460. https: // doi. org / 10.15560 / 10.2.457" type="journal article" year="2014">
Víquez
<emphasis id="B968EA92B65AFFCCFC21FC94FC1A7535" box="[968,1021,828,852]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2014: 457459
</bibRefCitation>
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, figs: 15;
<treatmentCitation id="0ABD1091B65AFFCCFAFFFC95FEC47512" author="Armas, L. F. &amp; Torres, R. &amp; Alvarez, D." page="69" pageId="8" pageNumber="9" year="2015">
<bibRefCitation id="EF8D4B71B65AFFCCFAFFFC95FEC47512" author="Armas, L. F. &amp; Torres, R. &amp; Alvarez, D." pageId="8" pageNumber="9" pagination="69 - 73" refId="ref22084" refString="Armas, L. F., Torres, R. &amp; Alvarez, D. (2015) Nueva especie de Heterophrynus (Amblypygi: Phrynidae) del Caribe Colombiano. Revista Iberica de Aracnologia, 26, 69 - 73." type="journal article" year="2015">
Armas
<emphasis id="B968EA92B65AFFCCFA8BFC94FA707535" box="[1378,1431,828,852]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
, 2015: 69
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ABD1091B65AFFCCFEC7FCF3FDC77512" author="Alvarez, D. &amp; Armas, L. F. &amp; Diaz, J." box="[302,544,858,883]" page="45" pageId="8" pageNumber="9" year="2015">
<bibRefCitation id="EF8D4B71B65AFFCCFEC7FCF3FDC77512" author="Alvarez, D. &amp; Armas, L. F. &amp; Diaz, J." box="[302,544,858,883]" pageId="8" pageNumber="9" pagination="45 - 49" refId="ref21803" refString="Alvarez, D., Armas, L. F. &amp; Diaz, J. (2015) Una especie nueva de Heterophrynus (Amblypygi: Phrynidae) del nordeste de Colombia. Revista Iberica de Aracnologia, 27, 45 - 49." type="journal article" year="2015">
Álvarez
<emphasis id="B968EA92B65AFFCCFE6DFCF2FE5F7513" box="[388,440,858,882]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
, 2015: 45
</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EF8D4B71B65AFFCCFDC3FCF3FC997512" author="Miranda, G. &amp; Giupponi, A. &amp; Kury, A." box="[554,894,858,883]" pageId="8" pageNumber="9" pagination="33 - 55" refId="ref23171" refString="Miranda, G., Giupponi, A. &amp; Kury, A. (2018) Review of Trichodamon Mello-Leitao 1935 and phylogenetic placement of the genus in Phrynichidae (Arachnida, Amblypygi). Zoologischer Anzeiger - A Journal of Comparative Zoology, 273, 33 - 55. https: // doi. org / 10.1016 / j. jcz. 2018.02.006" type="journal article" year="2018">
<collectingRegion id="49D8F862B65AFFCCFDC3FCF3FD657512" box="[554,642,858,883]" country="Venezuela" name="Miranda" pageId="8" pageNumber="9">Miranda</collectingRegion>
, Kury &amp; Giupponi 2018
</bibRefCitation>
: table 1;
<treatmentCitation id="0ABD1091B65AFFCCFC30FCF2FB2C7512" author="Vasquez, S. &amp; Chirivi, D. &amp; Garcia, A. &amp; Matilla-Meluk, H. &amp; Torres, J." box="[985,1227,858,883]" page="33" pageId="8" pageNumber="9" year="2019">
<bibRefCitation id="EF8D4B71B65AFFCCFC30FCF2FB2C7512" author="Vasquez, S. &amp; Chirivi, D. &amp; Garcia, A. &amp; Matilla-Meluk, H. &amp; Torres, J." box="[985,1227,858,883]" pageId="8" pageNumber="9" pagination="32 - 45" refId="ref23955" refString="Vasquez, S., Chirivi, D., Garcia, A., Matilla-Meluk, H. &amp; Torres, J. (2019) Variacion morfologica en Heterophrynus boterorum (Arachnida: Amblypygi: Phrynidae). Biota Colombiana, 20 (2), 32 - 45. https: // doi. org / 10.21068 / c 2019. v 20 n 02 a 03" type="journal article" year="2019">
Vásquez
<emphasis id="B968EA92B65AFFCCFBDFFCF2FB8D7513" box="[1078,1130,858,882]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2019: 33
</bibRefCitation>
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.
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<subSubSection id="C306650BB65AFFCCFF7EFC08FF097361" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BA33680B65AFFCCFF7EFC08FDF175DA" blockId="8.[151,1437,928,2035]" box="[151,534,928,955]" pageId="8" pageNumber="9">
<materialsCitation id="3B743CDDB65AFFCCFF7EFC08FDF575DA" box="[151,530,928,955]" country="Ecuador" location="Pambilar" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
<emphasis id="B968EA92B65AFFCCFF7EFC08FEDE75DA" bold="true" box="[151,313,928,955]" pageId="8" pageNumber="9">
<typeStatus id="54A78822B65AFFCCFF7EFC08FF3575DA" box="[151,210,929,955]" pageId="8" pageNumber="9">Type</typeStatus>
locality:
</emphasis>
<location id="8EC3605BB65AFFCCFEA9FC08FE4075DA" LSID="urn:lsid:plazi:treatment:03B58796B65AFFC8FF7EFD1CFC53755A:8EC3605BB65AFFCCFEA9FC08FE4075DA" box="[320,423,929,955]" country="Ecuador" name="Pambilar" pageId="8" pageNumber="9">Pambilar</location>
,
<collectingCountry id="F30B7610B65AFFCCFE5BFC08FDF575DA" box="[434,530,929,955]" name="Ecuador" pageId="8" pageNumber="9">Ecuador</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA33680B65AFFCCFF2EFC6DFF097361" blockId="8.[151,1437,928,2035]" pageId="8" pageNumber="9">
<emphasis id="B968EA92B65AFFCCFF2EFC6DFD6075BE" bold="true" box="[199,647,964,991]" pageId="8" pageNumber="9">
Revised material (
<specimenCount id="9D1AFD09B65AFFCCFE76FC6DFDF775BE" box="[415,528,964,991]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
,
<specimenCount id="9D1AFD09B65AFFCCFDF2FC6CFD9D75BE" box="[539,634,964,991]" pageId="8" pageNumber="9" type="male">7 males</specimenCount>
):
</emphasis>
<materialsCitation id="3B743CDDB65AFFCCFD64FC6CFF1C7262" country="Colombia" location="Natural National Park" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Cauca">
<collectingCountry id="F30B7610B65AFFCCFD64FC6CFCE775BE" box="[653,768,965,991]" name="Colombia" pageId="8" pageNumber="9">Colombia</collectingCountry>
:
<emphasis id="B968EA92B65AFFCCFCE3FC6CFCB475BE" box="[778,851,965,991]" italics="true" pageId="8" pageNumber="9">
<collectingRegion id="49D8F862B65AFFCCFCE3FC6CFCB475BE" box="[778,851,965,991]" country="Colombia" name="Cauca" pageId="8" pageNumber="9">Cauca</collectingRegion>
</emphasis>
: Guapi municipality,
<location id="8EC3605BB65AFFCCFBACFC6CFADE75BE" LSID="urn:lsid:plazi:treatment:03B58796B65AFFC8FF7EFD1CFC53755A:8EC3605BB65AFFCCFBACFC6CFADE75BE" box="[1093,1337,965,991]" country="Colombia" name="Natural National Park" pageId="8" pageNumber="9" stateProvince="Cauca">Natural National Park</location>
(N.N.P.) Gorgona
</materialsCitation>
,
<materialsCitation id="3B743CDDB65AFFCCFEE1FC40FF0D7361" collectingDate="1987-12-02" collectingDateMax="2011-02-24" collectingDateMin="1987-12-02" collectorName="M. L. Baena &amp; J. Portilla &amp; E. Solis &amp; El Poblado &amp; de Insectos &amp; M. Baena" country="Iceland" county="Gorgona" elevation="5" latitude="2.9543889" location="Grupo de Insectos" longLatPrecision="1" longitude="-78.168495" municipality="Village" pageId="8" pageNumber="9" specimenCount="4" specimenCount-male="4">
<collectingCountry id="F30B7610B65AFFCCFEE1FC40FED77262" box="[264,304,1001,1027]" name="Iceland" pageId="8" pageNumber="9">Isla</collectingCountry>
<collectingCounty id="62C24E0CB65AFFCCFED0FC40FE7D7262" box="[313,410,1001,1027]" pageId="8" pageNumber="9">Gorgona</collectingCounty>
(
<geoCoordinate id="EE285047B65AFFCCFE42FC41FDC87262" box="[427,559,1000,1027]" degrees="2" direction="north" minutes="57" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="53" value="2.9647222">2°5753″ N</geoCoordinate>
,
<geoCoordinate id="EE285047B65AFFCCFDD5FC41FD337262" box="[572,724,1000,1027]" degrees="78" direction="west" minutes="10" orientation="longitude" pageId="8" pageNumber="9" precision="15" seconds="30" value="-78.174995">78°1030″ W</geoCoordinate>
):
<collectingMunicipality id="6BC7ACFAB65AFFCCFD01FC40FCDE7262" box="[744,825,1001,1027]" pageId="8" pageNumber="9">Village</collectingMunicipality>
<quantity id="4CE49B65B65AFFCCFCABFC41FC8F7262" box="[834,872,1000,1027]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="5.0">
<elevation id="0031D1B3B65AFFCCFCABFC41FC8F7262" box="[834,872,1000,1027]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="5.0">5m</elevation>
</quantity>
, at
<quantity id="4CE49B65B65AFFCCFC7BFC40FC097262" box="[914,1006,1001,1027]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="9" unit="m" value="13.0">
<elevation id="0031D1B3B65AFFCCFC7BFC40FC097262" box="[914,1006,1001,1027]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="9" unit="m" value="13.0">13 masl</elevation>
</quantity>
, inside a house,
<date id="FFA21040B65AFFCCFB45FC41FAA87262" box="[1196,1359,1000,1027]" pageId="8" pageNumber="9" value="1990-03-05">
<collectingDate id="EFE6E9A8B65AFFCCFB45FC41FAA87262" box="[1196,1359,1000,1027]" pageId="8" pageNumber="9" value="1990-03-05">5 March 1990</collectingDate>
</date>
,
<collectorName id="26E95356B65AFFCCFAB5FC40FF077246" pageId="8" pageNumber="9">M. L. Baena</collectorName>
, male (IAvH without number); in a house, April 23 of 1991,
<collectorName id="26E95356B65AFFCCFC42FBA4FBFC7246" box="[939,1051,1037,1063]" pageId="8" pageNumber="9">J. Portilla</collectorName>
, manual collection, male (IAvH-I 2923); between bricks and construction material,
<date id="FFA21040B65AFFCCFD27FB98FC77722A" box="[718,912,1073,1099]" pageId="8" pageNumber="9" value="1991-10-20">
<collectingDate id="EFE6E9A8B65AFFCCFD27FB98FC77722A" box="[718,912,1073,1099]" pageId="8" pageNumber="9" value="1991-10-20">20 October 1991</collectingDate>
</date>
,
<collectorName id="26E95356B65AFFCCFC77FB98FC1D722A" box="[926,1018,1073,1099]" pageId="8" pageNumber="9">E. Solis</collectorName>
, manual collection,
<specimenCount id="9D1AFD09B65AFFCCFB08FB98FADE722A" box="[1249,1337,1073,1099]" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
(IAvH-I 2925, IAvH-I 2924). Sendero Cerro Los Micos (
<geoCoordinate id="EE285047B65AFFCCFD5AFBFDFCB3720E" box="[691,852,1108,1135]" degrees="2" direction="north" minutes="58" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="20,5" value="2.972361">2°58´20,5´´ N</geoCoordinate>
,
<geoCoordinate id="EE285047B65AFFCCFCB6FBFCFBFB720E" box="[863,1052,1109,1135]" degrees="78" direction="west" minutes="10" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="38,8" value="-78.177444">78° 10´38,8´´ W</geoCoordinate>
):
<date id="FFA21040B65AFFCCFBC6FBFCFB09720E" box="[1071,1262,1109,1135]" pageId="8" pageNumber="9" value="2010-10-21">
<collectingDate id="EFE6E9A8B65AFFCCFBC6FBFCFB09720E" box="[1071,1262,1109,1135]" pageId="8" pageNumber="9" value="2010-10-21">21 October 2010</collectingDate>
</date>
, Zoology team, female (MUSENUV 24303). Playa Palmeras (
<geoCoordinate id="EE285047B65AFFCCFD77FBD1FCD872F2" box="[670,831,1144,1171]" degrees="2" direction="north" minutes="56" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="28,6" value="2.9412777">2°56´28,6´´ N</geoCoordinate>
,
<geoCoordinate id="EE285047B65AFFCCFCA2FBD0FBE072F2" box="[843,1031,1145,1171]" degrees="78" direction="west" minutes="12" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="21,4" value="-78.20594">78° 12´21,4´´ W</geoCoordinate>
): Forest behind of a house,
<quantity id="4CE49B65B65AFFCCFAD4FBD0FA7F72F2" box="[1341,1432,1145,1171]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="9" unit="m" value="28.0">
<elevation id="0031D1B3B65AFFCCFAD4FBD0FA7F72F2" box="[1341,1432,1145,1171]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="9" unit="m" value="28.0">28 masl</elevation>
</quantity>
,
<date id="FFA21040B65AFFCCFF7EFB34FE8172D6" box="[151,358,1181,1207]" pageId="8" pageNumber="9" value="2011-02-24">
<collectingDate id="EFE6E9A8B65AFFCCFF7EFB34FE8172D6" box="[151,358,1181,1207]" pageId="8" pageNumber="9" value="2011-02-24">24 February 2011</collectingDate>
</date>
,
<specimenCount id="9D1AFD09B65AFFCCFE9AFB34FE2372D6" box="[371,452,1181,1207]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
,
<specimenCount id="9D1AFD09B65AFFCCFE3BFB34FDC772D6" box="[466,544,1181,1207]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
juvenile (MUSENUV 24304).
<collectorName id="26E95356B65AFFCCFC62FB34FBED72D6" box="[907,1034,1181,1207]" pageId="8" pageNumber="9">El Poblado</collectorName>
(
<geoCoordinate id="EE285047B65AFFCCFBF5FB35FB2772D6" box="[1052,1216,1180,1207]" degrees="2" direction="north" minutes="57" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="15,8" value="2.9543889">2°57´15,8´´ N</geoCoordinate>
,
<geoCoordinate id="EE285047B65AFFCCFB27FB34FA9F72D6" box="[1230,1400,1181,1207]" degrees="78" direction="west" minutes="10" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="6,6" value="-78.168495">78°10´6,6´´ W</geoCoordinate>
):
<date id="FFA21040B65AFFCCFA67FB34FEAA72BA" pageId="8" pageNumber="9" value="1987-12-02">
<collectingDate id="EFE6E9A8B65AFFCCFA67FB34FEAA72BA" pageId="8" pageNumber="9" value="1987-12-02">2 December 1987</collectingDate>
</date>
,
<location id="8EC3605BB65AFFCCFEB3FB68FDCC72BA" LSID="urn:lsid:plazi:treatment:03B58796B65AFFC8FF7EFD1CFC53755A:8EC3605BB65AFFCCFEB3FB68FDCC72BA" box="[346,555,1217,1243]" country="Iceland" county="Gorgona" latitude="2.9543889" longLatPrecision="1" longitude="-78.168495" municipality="Village" name="Grupo de Insectos" pageId="8" pageNumber="9">
Grupo
<collectorName id="26E95356B65AFFCCFE40FB68FDCC72BA" box="[425,555,1217,1243]" pageId="8" pageNumber="9">de Insectos</collectorName>
</location>
, male (MUSENUV 24307);
<date id="FFA21040B65AFFCCFC99FB68FBCC72BA" box="[880,1067,1217,1243]" pageId="8" pageNumber="9" value="1989-09">
<collectingDate id="EFE6E9A8B65AFFCCFC99FB68FBCC72BA" box="[880,1067,1217,1243]" pageId="8" pageNumber="9" value="1989-09">September 1989</collectingDate>
</date>
,
<collectorName id="26E95356B65AFFCCFBDEFB68FB4F72BA" box="[1079,1192,1217,1243]" pageId="8" pageNumber="9">M. Baena</collectorName>
, female (MUSENUV 24306)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C306650BB65AFFCCFF2EFAA1FDC67326" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BA33680B65AFFCCFF2EFAA1FDC67326" blockId="8.[151,1437,928,2035]" pageId="8" pageNumber="9">
<emphasis id="B968EA92B65AFFCCFF2EFAA1FE867342" bold="true" box="[199,353,1288,1315]" pageId="8" pageNumber="9">Distribution.</emphasis>
Countries:
<collectingCountry id="F30B7610B65AFFCCFE0EFAA0FDBF7342" box="[487,600,1289,1315]" name="Colombia" pageId="8" pageNumber="9">Colombia</collectingCountry>
,
<collectingCountry id="F30B7610B65AFFCCFD8FFAA0FD217342" box="[614,710,1289,1315]" name="Ecuador" pageId="8" pageNumber="9">Ecuador</collectingCountry>
. Other records from:
<bibRefCitation id="EF8D4B71B65AFFCCFC55FAA0FB4F7342" author="Viquez, C. &amp; Chirivi, D &amp; Moreno-Gonzalez, J. &amp; Christensen, J." box="[956,1192,1289,1315]" pageId="8" pageNumber="9" pagination="457 - 460" refId="ref24025" refString="Viquez, C., Chirivi, D, Moreno-Gonzalez, J. &amp; Christensen, J. (2014) Heterophrynus armiger Pocock, 1902 (Amblypygi: Phrynidae): First record from Colombia, with notes on its historic distribution records and natural history. Check List, 10 (2), 457 - 460. https: // doi. org / 10.15560 / 10.2.457" type="journal article" year="2014">
Víquez
<emphasis id="B968EA92B65AFFCCFBFFFAA3FBB77342" box="[1046,1104,1289,1315]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2014)
</bibRefCitation>
:
<collectingCountry id="F30B7610B65AFFCCFB5CFAA0FAC17342" box="[1205,1318,1289,1315]" name="Colombia" pageId="8" pageNumber="9">Colombia</collectingCountry>
, Gorgona;
<collectingCountry id="F30B7610B65AFFCCFF7EFA84FF117326" box="[151,246,1325,1351]" name="Ecuador" pageId="8" pageNumber="9">Ecuador</collectingCountry>
, Pambilar, Durango River.
</paragraph>
</subSubSection>
<subSubSection id="C306650BB65AFFC8FF2EFAF9FEE07741" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BA33680B65AFFCCFF2EFAF9FC5B73D2" blockId="8.[151,1437,928,2035]" pageId="8" pageNumber="9">
<emphasis id="B968EA92B65AFFCCFF2EFAF9FEA5730A" bold="true" box="[199,322,1360,1387]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
<taxonomicName id="4C1C4D03B65AFFCCFEA1FAF8FDA8730A" authorityName="Pocock" authorityYear="1902" box="[328,591,1361,1387]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="B968EA92B65AFFCCFEA1FAF8FDA8730A" box="[328,591,1361,1387]" italics="true" pageId="8" pageNumber="9">Heterophrynus armiger</emphasis>
</taxonomicName>
can be differentiated from other species by present the following characters together: four teeth in the ectal portion of the basal segment of the chelicerae. Five long spines in the ventral femur, spine FIII is smaller than FII and FV, FIV is absent or like as a tubercle.
</paragraph>
<paragraph id="8BA33680B65AFFCCFF2EFA14FCA7739D" blockId="8.[151,1437,928,2035]" pageId="8" pageNumber="9">
Female genitalia with gonopod sclerites elongated and with laminar shape, and an acuminated apex. Seminal receptacles oval with a sclerotized area at the base (
<figureCitation id="13272A05B65AFFCCFD30FA48FCD3739D" box="[729,820,1505,1532]" captionStart="FIGURE 6" captionStartId="11.[151,250,1755,1780]" captionTargetBox="[155,1433,529,1726]" captionTargetId="figure@11.[151,1436,525,1730]" captionTargetPageId="11" captionText="FIGURE 6. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MUSENUV 24303), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907913" httpUri="https://zenodo.org/record/3907913/files/figure.png" pageId="8" pageNumber="9">Fig. 6D</figureCitation>
).
</paragraph>
<paragraph id="8BA33680B65AFFCCFF2EF9ACFE3070CE" blockId="8.[151,1437,928,2035]" pageId="8" pageNumber="9">
Similar species:
<taxonomicName id="4C1C4D03B65AFFCCFE96F9AFFE1C707E" authorityName="Pocock" authorityYear="1902" box="[383,507,1542,1567]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="B968EA92B65AFFCCFE96F9AFFE1C707E" box="[383,507,1542,1567]" italics="true" pageId="8" pageNumber="9">H. armiger</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C1C4D03B65AFFCCFD65F9AFFD00707E" authorityName="Pocock" authorityYear="1902" box="[652,743,1541,1567]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alces">
<emphasis id="B968EA92B65AFFCCFD65F9AFFD00707E" box="[652,743,1541,1567]" italics="true" pageId="8" pageNumber="9">H. alces</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D03B65AFFCCFCF5F9AFFC8A707E" authority="Chirivi-Joya &amp; Moreno-González &amp; Fagua, 2020" authorityName="Chirivi-Joya &amp; Moreno-González &amp; Fagua" authorityYear="2020" box="[796,877,1542,1567]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
<emphasis id="B968EA92B65AFFCCFCF5F9AFFC8A707E" box="[796,877,1542,1567]" italics="true" pageId="8" pageNumber="9">H. awa</emphasis>
</taxonomicName>
<emphasis id="B968EA92B65AFFCCFC9AF9ACFC2A707E" bold="true" box="[883,973,1541,1567]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A25B57E9B65AFFCCFC9AF9ACFC2A707E" box="[883,973,1541,1567]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, however,
<taxonomicName id="4C1C4D03B65AFFCCFBAEF9AFFB44707E" authorityName="Pocock" authorityYear="1902" box="[1095,1187,1541,1567]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alces">
<emphasis id="B968EA92B65AFFCCFBAEF9AFFB44707E" box="[1095,1187,1541,1567]" italics="true" pageId="8" pageNumber="9">H. alces</emphasis>
</taxonomicName>
presents a small spine TIII and T3, which are absent in
<emphasis id="B968EA92B65AFFCCFDF9F983FD747022" box="[528,659,1578,1603]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C1C4D03B65AFFCCFDF9F983FD687022" authorityName="Pocock" authorityYear="1902" box="[528,655,1578,1603]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="armiger">H. armiger</taxonomicName>
.
</emphasis>
The female genital is completely different from the one of
<taxonomicName id="4C1C4D03B65AFFCCFAD1F983FA717022" authorityName="Pocock" authorityYear="1902" box="[1336,1430,1577,1603]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alces">
<emphasis id="B968EA92B65AFFCCFAD1F983FA717022" box="[1336,1430,1577,1603]" italics="true" pageId="8" pageNumber="9">H. alces</emphasis>
</taxonomicName>
, which presents the sclerites of the gonopods with a rounded shape, and
<taxonomicName id="4C1C4D03B65AFFCCFC5CF9E7FBE17006" authority="Chirivi-Joya &amp; Moreno-González &amp; Fagua, 2020" authorityName="Chirivi-Joya &amp; Moreno-González &amp; Fagua" authorityYear="2020" box="[949,1030,1614,1639]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
<emphasis id="B968EA92B65AFFCCFC5CF9E7FBE17006" box="[949,1030,1614,1639]" italics="true" pageId="8" pageNumber="9">H. awa</emphasis>
</taxonomicName>
<emphasis id="B968EA92B65AFFCCFBE5F9E4FB827006" bold="true" box="[1036,1125,1613,1639]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A25B57E9B65AFFCCFBE5F9E4FB827006" box="[1036,1125,1613,1639]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which have sclerites of the gonopods wider at the middle region. The male genital of
<taxonomicName id="4C1C4D03B65AFFCCFCFEF9DBFC7570EA" authorityName="Pocock" authorityYear="1902" box="[791,914,1650,1675]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="B968EA92B65AFFCCFCFEF9DBFC7570EA" box="[791,914,1650,1675]" italics="true" pageId="8" pageNumber="9">H. armiger</emphasis>
</taxonomicName>
presents a spine in LOL 1, which is not present in any other species (
<figureCitation id="13272A05B65AFFCCFE6DF93CFE2F70CE" box="[388,456,1685,1711]" captionStart="FIGURE 7" captionStartId="13.[151,250,670,695]" captionTargetBox="[152,1435,181,644]" captionTargetId="figure@13.[152,1436,181,644]" captionTargetPageId="13" captionText="FIGURE 7. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. Spine over LoL1, dorsal view. B. Detail of spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3907916" httpUri="https://zenodo.org/record/3907916/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BA33680B65AFFCCFF2EF911FAD17096" blockId="8.[151,1437,928,2035]" pageId="8" pageNumber="9">
<emphasis id="B968EA92B65AFFCCFF2EF911FCC770B2" bold="true" box="[199,800,1720,1747]" pageId="8" pageNumber="9">Complementary description. Male (IAvH-I 2924):</emphasis>
Total length
<quantity id="4CE49B65B65AFFCCFC52F910FBFB70B2" box="[955,1052,1721,1747]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" pageId="8" pageNumber="9" unit="mm" value="40.2">40.2mm</quantity>
. Carapace, opisthosoma, and legs have a black coloration, with brown areas, pedipalps black. Measures of the male are provided in
<tableCitation id="C69E033BB65AFFCCFB34F974FAD57096" box="[1245,1330,1757,1783]" captionStart="TABLE 2" captionStartId="12.[151,239,874,898]" captionText="TABLE 2. Measurements (mm) of the body, pedipalp segments, and femora of the legs in four specimens of Heterophrynus armiger. Empty cells correspond to missing segment or data." pageId="8" pageNumber="9">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BA33680B65AFFCCFF2EF8A8FE247102" blockId="8.[151,1437,928,2035]" pageId="8" pageNumber="9">
<emphasis id="B968EA92B65AFFCCFF2EF8A8FEDB717A" bold="true" box="[199,316,1793,1819]" italics="true" pageId="8" pageNumber="9">Carapace.</emphasis>
Anterior margin straight without lobes, carapace surface with a widely spaced granulation, posterior margin of carapace with two lobes. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (
<figureCitation id="13272A05B65AFFCCFEB5F8E0FE517102" box="[348,438,1865,1891]" captionStart="FIGURE 4" captionStartId="9.[151,250,1937,1962]" captionTargetBox="[161,1427,351,1902]" captionTargetId="figure@9.[151,1436,345,1912]" captionTargetPageId="9" captionText="FIGURE 4. Heterophrynus armiger. Male (IAvH-I 2924): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907909" httpUri="https://zenodo.org/record/3907909/files/figure.png" pageId="8" pageNumber="9">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="8BA33680B65AFFCDFF2EF8C4FCE476B6" blockId="8.[151,1437,928,2035]" lastBlockId="9.[151,1437,153,287]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B968EA92B65AFFCCFF2EF8C4FED671E6" bold="true" box="[199,305,1901,1927]" italics="true" pageId="8" pageNumber="9">Sternum.</emphasis>
Tri-segmented, segments are well sclerotized; the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with 10 setae in the basal region, four in the medial region two in distal region and two more in the apex. The second segment (tetrasternum) rounded and with four setae; two in the basal region and two in the apical region. The third segment (pentasternum) rounded, smaller than the second segment, has four setae; one on each side of the apical region, one on each side of the basal region. Metasternum longitudinally divided with two setae in each segment (
<figureCitation id="13272A05B65BFFCDFD77FF14FD1176B6" box="[670,758,189,215]" captionStart="FIGURE 4" captionStartId="9.[151,250,1937,1962]" captionTargetBox="[161,1427,351,1902]" captionTargetId="figure@9.[151,1436,345,1912]" captionTargetPageId="9" captionText="FIGURE 4. Heterophrynus armiger. Male (IAvH-I 2924): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907909" httpUri="https://zenodo.org/record/3907909/files/figure.png" pageId="9" pageNumber="10">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="8BA33680B65BFFCDFF2EFF49FF2F777E" blockId="9.[151,1437,153,287]" pageId="9" pageNumber="10">
<emphasis id="B968EA92B65BFFCDFF2EFF49FEDB769A" bold="true" box="[199,316,224,251]" italics="true" pageId="9" pageNumber="10">Abdomen.</emphasis>
Oblong, color black and brown, with soft punctuations. Carapace wider than the Abdomen (
<figureCitation id="13272A05B65BFFCDFA86FF48FF5B777E" captionStart="FIGURE 4" captionStartId="9.[151,250,1937,1962]" captionTargetBox="[161,1427,351,1902]" captionTargetId="figure@9.[151,1436,345,1912]" captionTargetPageId="9" captionText="FIGURE 4. Heterophrynus armiger. Male (IAvH-I 2924): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907909" httpUri="https://zenodo.org/record/3907909/files/figure.png" pageId="9" pageNumber="10">Fig. 4C</figureCitation>
).
</paragraph>
<caption id="DF636608B65BFFCDFF7EF838FD2A71AF" ID-DOI="http://doi.org/10.5281/zenodo.3907909" ID-Zenodo-Dep="3907909" httpUri="https://zenodo.org/record/3907909/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1937,1962]" targetBox="[161,1427,351,1902]" targetPageId="9">
<paragraph id="8BA33680B65BFFCDFF7EF838FD2A71AF" blockId="9.[151,1437,1937,1998]" pageId="9" pageNumber="10">
<emphasis id="B968EA92B65BFFCDFF7EF838FEF171CB" bold="true" box="[151,278,1937,1962]" pageId="9" pageNumber="10">FIGURE 4.</emphasis>
<taxonomicName id="4C1C4D03B65BFFCDFEF6F838FDF671C8" authorityName="Pocock" authorityYear="1902" box="[287,529,1937,1961]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="armiger">
<emphasis id="B968EA92B65BFFCDFEF6F838FDF671C8" box="[287,529,1937,1961]" italics="true" pageId="9" pageNumber="10">Heterophrynus armiger</emphasis>
</taxonomicName>
. Male (IAvH-I 2924): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view.
</paragraph>
</caption>
<paragraph id="8BA33680B658FFCEFF2EFF31FA6F7722" blockId="10.[151,1436,152,323]" pageId="10" pageNumber="11">
<emphasis id="B968EA92B658FFCEFF2EFF31FED976D2" bold="true" box="[199,318,152,179]" italics="true" pageId="10" pageNumber="11">Chelicera.</emphasis>
Presents a mesal row of three teeth on the basal segment of the chelicerae with the first in the proximal region, with two cuspids, follow by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (
<figureCitation id="13272A05B658FFCEFE39FF48FDCB769A" box="[464,556,225,251]" captionStart="FIGURE 4" captionStartId="9.[151,250,1937,1962]" captionTargetBox="[161,1427,351,1902]" captionTargetId="figure@9.[151,1436,345,1912]" captionTargetPageId="9" captionText="FIGURE 4. Heterophrynus armiger. Male (IAvH-I 2924): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907909" httpUri="https://zenodo.org/record/3907909/files/figure.png" pageId="10" pageNumber="11">Fig. 4D</figureCitation>
). The ectal row of the basal segment of the chelicerae have four conspicuous teeth, two with acuminated apex placed in the proximal and medial portions (tooth 1 and 2), and two with acuminated apex placed on the medial portion (tooth 3 and 4) (
<figureCitation id="13272A05B658FFCEFCFAFE80FC8E7722" box="[787,873,297,323]" captionStart="FIGURE 4" captionStartId="9.[151,250,1937,1962]" captionTargetBox="[161,1427,351,1902]" captionTargetId="figure@9.[151,1436,345,1912]" captionTargetPageId="9" captionText="FIGURE 4. Heterophrynus armiger. Male (IAvH-I 2924): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907909" httpUri="https://zenodo.org/record/3907909/files/figure.png" pageId="10" pageNumber="11">Fig. 4F</figureCitation>
). Mobile finger of the chelicerae with five teeth.
</paragraph>
<caption id="DF636608B658FFCEFF7EF938FCC370AF" ID-DOI="http://doi.org/10.5281/zenodo.3907911" ID-Zenodo-Dep="3907911" httpUri="https://zenodo.org/record/3907911/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1681,1706]" targetBox="[161,1427,355,1644]" targetPageId="10">
<paragraph id="8BA33680B658FFCEFF7EF938FCC370AF" blockId="10.[151,1437,1681,1742]" pageId="10" pageNumber="11">
<emphasis id="B968EA92B658FFCEFF7EF938FEF470CB" bold="true" box="[151,275,1681,1706]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
<emphasis id="B968EA92B658FFCEFEF0F93BFDEB70CB" box="[281,524,1682,1706]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C1C4D03B658FFCEFEF0F93BFDEF70CB" authorityName="Pocock" authorityYear="1902" box="[281,520,1682,1706]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="armiger">Heterophrynus armiger</taxonomicName>
.
</emphasis>
Male (IAvH-I 2924): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral
<taxonomicName id="4C1C4D03B658FFCEFAC8F938FA7E70CB" authority="(Pv)" baseAuthorityName="Pv" box="[1313,1433,1681,1706]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="10" pageNumber="34" phylum="Mollusca" rank="species" species="patella">patella (Pv)</taxonomicName>
. D. Dorsal
<taxonomicName id="4C1C4D03B658FFCEFEE9F91CFE9E70AF" authority="(Pd)" baseAuthorityName="Pd" box="[256,377,1717,1742]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="10" pageNumber="34" phylum="Mollusca" rank="species" species="patella">patella (Pd)</taxonomicName>
. E. Ventral tibia (Tv). F. Dorsal tibia (Td).
</paragraph>
</caption>
<paragraph id="8BA33680B658FFCFFF2EF952FDA17796" blockId="10.[151,1436,1787,2031]" lastBlockId="11.[151,1437,152,503]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B968EA92B658FFCEFF2EF952FE5F7177" box="[199,440,1787,1814]" italics="true" pageId="10" pageNumber="11">
<emphasis id="B968EA92B658FFCEFF2EF952FED47177" bold="true" box="[199,307,1787,1814]" italics="true" pageId="10" pageNumber="11">Pedipalp.</emphasis>
Trochanter
</emphasis>
: Prolateral face with four spines; spines Tr1 and Tr3 placed in the medial region, Tr2 placed near to ventral margin. Tr3 and Tr4 with subequal lengths. Spines lengths: Tr2&gt;Tr1&gt;Tr4≥Tr3. Dorsal oblique series of six setal tubercles. The dorsomedial area without spines but with one tubercle (
<figureCitation id="13272A05B658FFCEFB6DF8EDFB3A713F" box="[1156,1245,1860,1886]" captionStart="FIGURE 4" captionStartId="9.[151,250,1937,1962]" captionTargetBox="[161,1427,351,1902]" captionTargetId="figure@9.[151,1436,345,1912]" captionTargetPageId="9" captionText="FIGURE 4. Heterophrynus armiger. Male (IAvH-I 2924): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907909" httpUri="https://zenodo.org/record/3907909/files/figure.png" pageId="10" pageNumber="11">Fig. 4E</figureCitation>
).
<emphasis id="B968EA92B658FFCEFB18F8ECFADB713F" box="[1265,1340,1861,1886]" italics="true" pageId="10" pageNumber="11">Femur</emphasis>
: Ventral face with five major spines, there are three tubercles between FIIFIII, and FIIIFV, a small spine and 12 tubercles, between FVFVI, ten tubercles more distally than FVI, FIV absent. Spine lengths: FI&gt;FII&gt;FV&gt;FVI&gt;FIII (
<figureCitation id="13272A05B658FFCEFADFF825FA7771C7" box="[1334,1424,1931,1958]" captionStart="FIGURE 5" captionStartId="10.[151,250,1681,1706]" captionTargetBox="[161,1427,355,1644]" captionTargetId="figure@10.[151,1436,345,1656]" captionTargetPageId="10" captionText="FIGURE 5. Heterophrynus armiger. Male (IAvH-I 2924): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907911" httpUri="https://zenodo.org/record/3907911/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Dorsal face with five major spines, there are two tubercles between F2F3, a very small spine with two tubercles at each side between F3F4, a very small spine with three tubercles more proximal and two more distally between F4F5, and more than ten tubercles more distally than F5. F1 and F2 are joined at the base, F3 clearly separated from F2. Spine lengths: F3&gt;F4&gt;F1=F2&gt;F5 (
<figureCitation id="13272A05B659FFCFFD66FF14FD0F76B6" box="[655,744,188,215]" captionStart="FIGURE 5" captionStartId="10.[151,250,1681,1706]" captionTargetBox="[161,1427,355,1644]" captionTargetId="figure@10.[151,1436,345,1656]" captionTargetPageId="10" captionText="FIGURE 5. Heterophrynus armiger. Male (IAvH-I 2924): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907911" httpUri="https://zenodo.org/record/3907911/files/figure.png" pageId="11" pageNumber="12">Fig. 5B</figureCitation>
).
<taxonomicName id="4C1C4D03B659FFCFFD14FF14FC4876B6" authority=": Ventral" authorityName="Ventral" box="[765,943,189,215]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="B968EA92B659FFCFFD14FF14FCA976B6" box="[765,846,189,215]" italics="true" pageId="11" pageNumber="12">Patella</emphasis>
: Ventral
</taxonomicName>
face with five major spines, there are three tubercles between PVPIV, three between PIVPIII, four between PIIIPII and between PIIPI, PI wider than the other spines, PVI is absent. Spine lengths: PIV&gt;PII&gt;PV&gt;PI&gt;PIII (
<figureCitation id="13272A05B659FFCFFC7AFEACFC08777E" box="[915,1007,260,287]" captionStart="FIGURE 5" captionStartId="10.[151,250,1681,1706]" captionTargetBox="[161,1427,355,1644]" captionTargetId="figure@10.[151,1436,345,1656]" captionTargetPageId="10" captionText="FIGURE 5. Heterophrynus armiger. Male (IAvH-I 2924): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907911" httpUri="https://zenodo.org/record/3907911/files/figure.png" pageId="11" pageNumber="12">Fig. 5C</figureCitation>
). Dorsal face with five major spines, seven tubercles between P6P5; P7, and P2 are absent. Spine lengths: P3=P4≥P5&gt;P6&gt;P1 (
<figureCitation id="13272A05B659FFCFFB7DFE80FB097722" box="[1172,1262,296,323]" captionStart="FIGURE 5" captionStartId="10.[151,250,1681,1706]" captionTargetBox="[161,1427,355,1644]" captionTargetId="figure@10.[151,1436,345,1656]" captionTargetPageId="10" captionText="FIGURE 5. Heterophrynus armiger. Male (IAvH-I 2924): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907911" httpUri="https://zenodo.org/record/3907911/files/figure.png" pageId="11" pageNumber="12">Fig. 5D</figureCitation>
).
<emphasis id="B968EA92B659FFCFFAEAFE80FADA7722" box="[1283,1341,297,323]" italics="true" pageId="11" pageNumber="12">Tibia</emphasis>
: Ventral face with two major spines. Spine lengths: TII&gt;TI (
<figureCitation id="13272A05B659FFCFFD32FEE4FCD57706" box="[731,818,332,359]" captionStart="FIGURE 5" captionStartId="10.[151,250,1681,1706]" captionTargetBox="[161,1427,355,1644]" captionTargetId="figure@10.[151,1436,345,1656]" captionTargetPageId="10" captionText="FIGURE 5. Heterophrynus armiger. Male (IAvH-I 2924): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907911" httpUri="https://zenodo.org/record/3907911/files/figure.png" pageId="11" pageNumber="12">Fig. 5E</figureCitation>
). Dorsal face with two major spines, with one tubercle between T2T3. Spine lengths: T2&gt;T1 (
<figureCitation id="13272A05B659FFCFFDBFFED8FD4C77EA" box="[598,683,368,395]" captionStart="FIGURE 5" captionStartId="10.[151,250,1681,1706]" captionTargetBox="[161,1427,355,1644]" captionTargetId="figure@10.[151,1436,345,1656]" captionTargetPageId="10" captionText="FIGURE 5. Heterophrynus armiger. Male (IAvH-I 2924): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907911" httpUri="https://zenodo.org/record/3907911/files/figure.png" pageId="11" pageNumber="12">Fig. 5F</figureCitation>
).
<emphasis id="B968EA92B659FFCFFD57FEDBFC7277EA" box="[702,917,370,395]" italics="true" pageId="11" pageNumber="12">Tarsus-metatarsus:</emphasis>
Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible.
<emphasis id="B968EA92B659FFCFFBB9FE3CFB6177CE" bold="true" box="[1104,1158,405,431]" italics="true" pageId="11" pageNumber="12">Legs</emphasis>
. Femora brown. Femora lengths: I&gt;III&gt;II&gt;IV (
<tableCitation id="C69E033BB659FFCFFE63FE10FE0577B2" box="[394,482,441,467]" captionStart="TABLE 2" captionStartId="12.[151,239,874,898]" captionText="TABLE 2. Measurements (mm) of the body, pedipalp segments, and femora of the legs in four specimens of Heterophrynus armiger. Empty cells correspond to missing segment or data." pageId="11" pageNumber="12">Table 2</tableCitation>
). Leg I: Tibia with 31 segments and tarsus with 70 segments in the right and left leg. Leg IV: Basitibia with three segments.
</paragraph>
<caption id="DF636608B659FFCFFF7EF972FDA77101" ID-DOI="http://doi.org/10.5281/zenodo.3907913" ID-Zenodo-Dep="3907913" httpUri="https://zenodo.org/record/3907913/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1755,1780]" targetBox="[155,1433,529,1726]" targetPageId="11">
<paragraph id="8BA33680B659FFCFFF7EF972FDA77101" blockId="11.[151,1437,1755,1888]" pageId="11" pageNumber="12">
<emphasis id="B968EA92B659FFCFFF7EF972FEF37095" bold="true" box="[151,276,1755,1780]" pageId="11" pageNumber="12">FIGURE 6.</emphasis>
Male genital:
<emphasis id="B968EA92B659FFCFFE44F972FD457092" box="[429,674,1755,1779]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C1C4D03B659FFCFFE44F972FD7A7092" authorityName="Pocock" authorityYear="1902" box="[429,669,1755,1779]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="armiger">Heterophrynus armiger</taxonomicName>
.
</emphasis>
(IAvH-I 2924): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi =
<emphasis id="B968EA92B659FFCFFE2BF956FDE67176" box="[450,513,1791,1815]" italics="true" pageId="11" pageNumber="12">fistula</emphasis>
; LaM =
<emphasis id="B968EA92B659FFCFFD89F956FCE37176" box="[608,772,1791,1815]" italics="true" pageId="11" pageNumber="12">lamina medialis</emphasis>
; LoD =
<emphasis id="B968EA92B659FFCFFC89F956FC167176" box="[864,1009,1791,1815]" italics="true" pageId="11" pageNumber="12">lobus dorsalis</emphasis>
; LoL 1=
<emphasis id="B968EA92B659FFCFFBBCF956FAD07176" box="[1109,1335,1791,1815]" italics="true" pageId="11" pageNumber="12">lobus lateralis primus</emphasis>
; LoL 2 =
<emphasis id="B968EA92B659FFCFFF7EF88AFE6C715A" box="[151,395,1827,1851]" italics="true" pageId="11" pageNumber="12">lobus lateralis secundus</emphasis>
; Pi =
<emphasis id="B968EA92B659FFCFFE2EF88DFD65715A" box="[455,642,1828,1851]" italics="true" pageId="11" pageNumber="12">processus internus</emphasis>
of gonopod. D. Female genital (MUSENUV 24303), black arrow indicates the sclerotized area of the seminal receptacle.
</paragraph>
</caption>
<paragraph id="8BA33680B659FFCFFF2EF827FAA871C9" blockId="11.[151,1436,1934,2032]" box="[199,1359,1934,1960]" pageId="11" pageNumber="12">Basitibia-distitibia lengths: BT1&gt;DT&gt;BT3&gt;BT2. Basitarsus shorter than telotarsus. Tarsus tetramerous.</paragraph>
<paragraph id="8BA33680B659FFC8FF2EF818FEE07741" blockId="11.[151,1436,1934,2032]" lastBlockId="12.[151,1437,153,827]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B968EA92B659FFCFFF2EF818FD7271AD" bold="true" box="[199,661,1969,1996]" pageId="11" pageNumber="12">
<emphasis id="B968EA92B659FFCFFF2EF818FE7471AD" bold="true" box="[199,403,1969,1996]" italics="true" pageId="11" pageNumber="12">Female genitalia.</emphasis>
(MUSENUV 24303):
</emphasis>
Genital operculum pentagonal, reaching the second segment of the opisthosoma, seminal receptacles with an oval shape, completely separated at the medial portion, dorsal surface sclerotized at the proximal region, with brown coloration, medial and distal portion white. Sclerites with laminar shape, distal portion a little curved and thinner than the basal portion, with acuminated apex, sclerites placed parallel, longer than the width of the seminal receptacle, sclerites cover almost the total length of the seminal receptacle (
<figureCitation id="13272A05B65EFFC8FF49FEACFF1D7741" box="[160,250,261,288]" captionStart="FIGURE 6" captionStartId="11.[151,250,1755,1780]" captionTargetBox="[155,1433,529,1726]" captionTargetId="figure@11.[151,1436,525,1730]" captionTargetPageId="11" captionText="FIGURE 6. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MUSENUV 24303), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907913" httpUri="https://zenodo.org/record/3907913/files/figure.png" pageId="12" pageNumber="13">Fig. 6D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306650BB65EFFC8FF2EFE81FC53755A" pageId="12" pageNumber="13" type="description">
<paragraph id="8BA33680B65EFFC8FF2EFE81FA7B7722" blockId="12.[151,1437,153,827]" box="[199,1436,296,323]" pageId="12" pageNumber="13">
<emphasis id="B968EA92B65EFFC8FF2EFE81FED87722" bold="true" box="[199,319,296,323]" pageId="12" pageNumber="13">Variation.</emphasis>
Variation in measures and segments of the basitibia IV, tibia I and tarsus I, are provided in
<tableCitation id="C69E033BB65EFFC8FAABFE80FA7F7722" box="[1346,1432,297,323]" captionStart="TABLE 2" captionStartId="12.[151,239,874,898]" captionText="TABLE 2. Measurements (mm) of the body, pedipalp segments, and femora of the legs in four specimens of Heterophrynus armiger. Empty cells correspond to missing segment or data." pageId="12" pageNumber="13">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BA33680B65EFFC8FF7EFEE4FDB27402" blockId="12.[151,1437,153,827]" pageId="12" pageNumber="13">
<emphasis id="B968EA92B65EFFC8FF7EFEE4FEE57706" bold="true" box="[151,258,333,359]" pageId="12" pageNumber="13">
<emphasis id="B968EA92B65EFFC8FF7EFEE4FF1C7706" bold="true" box="[151,251,333,359]" italics="true" pageId="12" pageNumber="13">Sternum</emphasis>
.
</emphasis>
The number of setae over all segments of the sternum was variable.
<emphasis id="B968EA92B65EFFC8FBFFFEE5FB9C7706" bold="true" box="[1046,1147,332,359]" italics="true" pageId="12" pageNumber="13">Pedipalp</emphasis>
.
<emphasis id="B968EA92B65EFFC8FB62FEE4FAE17706" box="[1163,1286,333,359]" italics="true" pageId="12" pageNumber="13">Trochanter</emphasis>
: line of setal tubercles vary between five and six.
<emphasis id="B968EA92B65EFFC8FDC3FEDBFD9277EA" box="[554,629,370,395]" italics="true" pageId="12" pageNumber="13">Femur</emphasis>
: Ventral face: number of tubercles between spines are variable, the small spine between FV and FVI, and tubercles can be more conspicuous. In the specimen MUSENUV 24306, there is an FIV with the same size of FIII. Dorsal face: tubercles between spines are variable, can be present a very small spine between F4 and F5.
<taxonomicName id="4C1C4D03B65EFFC8FE2BFE74FDF47796" box="[450,531,477,503]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="B968EA92B65EFFC8FE2BFE74FDF47796" box="[450,531,477,503]" italics="true" pageId="12" pageNumber="13">Patella</emphasis>
</taxonomicName>
. Ventral face: number and size of the tubercles are variable, PVI can be present as a small spine. Dorsal face: number of tubercles are variable, P2 can be present.
<emphasis id="B968EA92B65EFFC8FBA8FDA8FB65747A" box="[1089,1154,513,539]" italics="true" pageId="12" pageNumber="13">Tibia.</emphasis>
Ventral and Dorsal face: number of tubercles between spines are variable.
<emphasis id="B968EA92B65EFFC8FD2BFD8CFCE7745E" bold="true" box="[706,768,549,575]" italics="true" pageId="12" pageNumber="13">Legs.</emphasis>
leg I: Number of segments of tibia vary between 31 and 32 and tarsus between 66 and 71 (
<tableCitation id="C69E033BB65EFFC8FE19FDE0FDAF7402" box="[496,584,585,611]" captionStart="TABLE 2" captionStartId="12.[151,239,874,898]" captionText="TABLE 2. Measurements (mm) of the body, pedipalp segments, and femora of the legs in four specimens of Heterophrynus armiger. Empty cells correspond to missing segment or data." pageId="12" pageNumber="13">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BA33680B65EFFC8FF2EFDC4FBAC74CA" blockId="12.[151,1437,153,827]" pageId="12" pageNumber="13">
<emphasis id="B968EA92B65EFFC8FF2EFDC4FEEC74E6" bold="true" box="[199,267,621,647]" pageId="12" pageNumber="13">Male.</emphasis>
Observed Males were bigger than females (
<tableCitation id="C69E033BB65EFFC8FD1EFDC4FCAA74E6" box="[759,845,621,647]" captionStart="TABLE 2" captionStartId="12.[151,239,874,898]" captionText="TABLE 2. Measurements (mm) of the body, pedipalp segments, and femora of the legs in four specimens of Heterophrynus armiger. Empty cells correspond to missing segment or data." pageId="12" pageNumber="13">Table 2</tableCitation>
); Spination pattern same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma.
</paragraph>
<paragraph id="8BA33680B65EFFC8FF2EFD1DFC53755A" blockId="12.[151,1437,153,827]" pageId="12" pageNumber="13">
<emphasis id="B968EA92B65EFFC8FF2EFD1DFDCC74AE" bold="true" box="[199,555,692,719]" pageId="12" pageNumber="13">
<emphasis id="B968EA92B65EFFC8FF2EFD1DFE8974AE" bold="true" box="[199,366,692,719]" italics="true" pageId="12" pageNumber="13">Male genitalia</emphasis>
. (IAvH-I 2924):
</emphasis>
LaM longer than Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute conical projections, placed close together with the base not visible; LoL 2 is wrinkled and does not present projections. In both LoL 1, there is present a spine sticking out from the projections. In dorsal view, the LoD are elongated and do not cover LoL1 and LoL 2 (
<figureCitation id="13272A05B65EFFC8FD56FC88FCBD755A" box="[703,858,801,827]" captionStart="FIGURE 6" captionStartId="11.[151,250,1755,1780]" captionTargetBox="[155,1433,529,1726]" captionTargetId="figure@11.[151,1436,525,1730]" captionTargetPageId="11" captionText="FIGURE 6. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MUSENUV 24303), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907913" httpUri="https://zenodo.org/record/3907913/files/figure.png" pageId="12" pageNumber="13">Fig. 6A, B, C</figureCitation>
;
<figureCitation id="13272A05B65EFFC8FC8CFC88FC94755A" box="[869,883,801,827]" captionStart="FIGURE 7" captionStartId="13.[151,250,670,695]" captionTargetBox="[152,1435,181,644]" captionTargetId="figure@13.[152,1436,181,644]" captionTargetPageId="13" captionText="FIGURE 7. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. Spine over LoL1, dorsal view. B. Detail of spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3907916" httpUri="https://zenodo.org/record/3907916/files/figure.png" pageId="12" pageNumber="13">7</figureCitation>
A-B).
</paragraph>
</subSubSection>
</treatment>
</document>