135 lines
16 KiB
XML
135 lines
16 KiB
XML
<document id="CF24EE856E671EEC785DB888BAD00B50" ID="10.11646/zootaxa.4268.2.1SLASH11054" ID-CLB-Dataset="33045" ID-DOI="10.11646/zootaxa.4268.2.1" ID-GBIF-Dataset="e114acb7-3683-4094-a53f-c9bc7ed84353" ID-ISSN="1175-5326" ID-Zenodo-Dep="580173" ID-ZooBank="0742D287-B82C-4014-A6AC-C357F259D5D7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494919993818" checkinUser="plazi" docAuthor="Summers, Mindi M., Messing, Charles G. & Rouse, Greg W." docDate="2017" docId="039EDF70FFA3A17DFF66DF2D13C1F99A" docLanguage="en" docName="zootaxa.4268.2.1SLASH11054.pdf" docOrigin="Zootaxa 4268 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clarkcomanthus albinotus Rowe, Hoggett, Birtles & Vail 1986" docType="treatment" docVersion="8" lastPageNumber="180" masterDocId="FFA7A708FFBEA160FFF1DB31103FFFDD" masterDocTitle="The genera and species of Comatulidae (Comatulida: Crinoidea): taxonomic revisions and a molecular and morphological guide" masterLastPageNumber="190" masterPageNumber="151" pageNumber="180" updateTime="1698439592185" updateUser="ExternalLinkService">
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<mods:title id="9188EC0AECD40BD2D40B2A6880BF1565">The genera and species of Comatulidae (Comatulida: Crinoidea): taxonomic revisions and a molecular and morphological guide</mods:title>
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<mods:namePart id="76A88380B344D0A2EA82B029DADF8099">Summers, Mindi M.</mods:namePart>
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<mods:namePart id="6A36C9904363DCAFD3465381706D7806">Messing, Charles G.</mods:namePart>
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<mods:namePart id="007CFCA52924431F54594F710BD66193">Rouse, Greg W.</mods:namePart>
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<mods:title id="A1CBD95CB01E384DAA72FBB2D3F53BFB">Zootaxa</mods:title>
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<mods:date id="8C2FBA8900B91D09732E763970366DFD">2017</mods:date>
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<mods:number id="136C558A82CF2CE16F14EE685D711181">4268</mods:number>
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<treatment id="039EDF70FFA3A17DFF66DF2D13C1F99A" ID-DOI="http://doi.org/10.5281/zenodo.6009148" ID-GBIF-Taxon="130809852" ID-Zenodo-Dep="6009148" LSID="urn:lsid:plazi:treatment:039EDF70FFA3A17DFF66DF2D13C1F99A" httpUri="http://treatment.plazi.org/id/039EDF70FFA3A17DFF66DF2D13C1F99A" lastPageNumber="180" pageId="29" pageNumber="180">
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<subSubSection id="C32D3DEDFFA3A17DFF66DF2D112CFB8A" pageId="29" pageNumber="180" type="nomenclature">
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<paragraph id="8B886E66FFA3A17DFF66DF2D13A4FBEB" blockId="29.[151,923,1052,1111]" box="[151,923,1052,1078]" pageId="29" pageNumber="180">
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<heading id="D0C0D90AFFA3A17DFF66DF2D13A4FBEB" bold="true" box="[151,923,1052,1078]" fontSize="11" level="1" pageId="29" pageNumber="180" reason="1">
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<taxonomicName id="4C3715E5FFA3A17DFF66DF2D13A4FBEB" authority="Rowe, Hoggett, Birtles & Vail, 1986" authorityName="Rowe, Hoggett, Birtles & Vail" authorityYear="1986" box="[151,923,1052,1078]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="albinotus">
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<emphasis id="B943B274FFA3A17DFF66DF2D13A4FBEB" bold="true" box="[151,923,1052,1078]" pageId="29" pageNumber="180">
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<emphasis id="B943B274FFA3A17DFF66DF2D11E7FBEB" bold="true" box="[151,472,1052,1078]" italics="true" pageId="29" pageNumber="180">Clarkcomanthus albinotus</emphasis>
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<bibRefCitation id="EFA61397FFA3A17DFE11DF2D13A4FBEB" author="Rowe" box="[480,923,1052,1078]" pageId="29" pageNumber="180" refString="Rowe, F. W. E., Hoggett, A. K., Birtles, R. A. & Vail, L. L. (1986) Revision of some comasterid genera from Australia (Echinodermata: Crinoidea), with descriptions of two new genera and nine new species. Zoological Journal of the Linnean Society, 86, 197 - 277." type="journal article" year="1986">Rowe, Hoggett, Birtles & Vail, 1986</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B886E66FFA3A17DFF66DF0F112CFB8A" blockId="29.[151,923,1052,1111]" box="[151,275,1086,1111]" pageId="29" pageNumber="180">
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<figureCitation id="130C72E3FFA3A17DFF66DF0F10DCFB8A" box="[151,227,1086,1111]" captionStart="FIGURE 13" captionStartId="27.[151,250,1675,1697]" captionTargetBox="[252,1326,193,1632]" captionTargetId="figure@27.[252,1326,193,1632]" captionTargetPageId="27" captionText="FIGURE 13. Clarkcomanthus. (A – J) Clarkcomanthus albinotus— (A) live, oral view following extraction from rubble; (B) semicryptic under ledge; (C – H) aboral views; (I – J) short combs tapering near tip, with J showing paired confluent comb teeth. (K – L) Clarkcomanthus luteofuscum, aboral surface showing rounded centrodorsal with row of cirri. (M – N) Clarkcomanthus cf. littoralis — (M) aboral surface showing centrodorsal with no cirri; (N) comb with separated confluent teeth, smaller near tip. (O – R) Clarkcomanthus comanthipinnus— (O, P) live, oral view following extraction from rubble; (Q) aboral surfaces showing rounded centrodorsal with few cirri at interradials; (R) comb tapering to distal point, composed of separated nonconfluent teeth. (S – T) Clarkcomanthus mirus— (S) aboral surface showing centrodorsal with cirrus buds at interradials; (T) comb tapering to blunt tip, composed of separated nonconfluent teeth. Madang, Papua New Guinea: A. MNHN-IE- 2013 - 8060; B, C. MNHN-IE- 2013 - 8122; D. MNHN-IE- 2013 - 8068; K. MNHN-IE- 2013 - 8184; O. MNHN-IE- 2013 - 8082; P. MNHN-IE- 2013 - 8151. Raja Ampat, Indonesia: E. SIO-BIC E 6186; F. SIO-BIC E 6187; G, I. SIO-BIC E 6190; L. SIO-BIC E 6210; M, N. SIO-BIC E 6193. South Island, near Lizard Island, Australia: H, J. AM J 17360; holotype. Lizard Island, Australia: Q-R. AM J 16992, holotype; S, T. AM J 17202, holotype." httpUri="https://zenodo.org/record/580186/files/figure.png" pageId="29" pageNumber="180">Fig. 13</figureCitation>
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A–J
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D3DEDFFA3A17DFF66DFB7128FFB66" pageId="29" pageNumber="180" type="reference_group">
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<paragraph id="8B886E66FFA3A17DFF66DFB7128FFB66" blockId="29.[151,688,1158,1211]" pageId="29" pageNumber="180">
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<treatmentCitationGroup id="AB274948FFA3A17DFF66DFB7128FFB66" pageId="29" pageNumber="180">
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<emphasis id="B943B274FFA3A17DFF66DFB711BEFB40" box="[151,385,1158,1181]" italics="true" pageId="29" pageNumber="180">
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<taxonomicName id="4C3715E5FFA3A17DFF66DFB71144FB40" box="[151,379,1158,1181]" class="Crinoidea" family="Comasteridae" genus="Comanthus" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="parvicirra">Comanthus parvicirra</taxonomicName>
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:
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</emphasis>
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HL
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<treatmentCitation id="0A964877FFA3A17DFE5EDFB71265FB41" author="Clark" box="[431,602,1158,1180]" page="200" pageId="29" pageNumber="180" year="1932">
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<bibRefCitation id="EFA61397FFA3A17DFE5EDFB71215FB41" author="Clark" box="[431,554,1158,1180]" pageId="29" pageNumber="180" refString="Clark, H. L. (1932) Echinodermata (other than Asteroidea) of the Great Barrier Reef Expedition, 1928 - 1929. Scientific Reports of the Great Barrier Reef Expedition, 4, 197 - 239." type="journal article" year="1932">Clark, 1932</bibRefCitation>
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: 200
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</treatmentCitation>
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(part).
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<emphasis id="B943B274FFA3A17DFF66DF9411B4FB66" box="[151,395,1189,1211]" italics="true" pageId="29" pageNumber="180">
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<taxonomicName id="4C3715E5FFA3A17DFF66DF9411BAFB66" box="[151,389,1189,1211]" class="Crinoidea" family="Comasteridae" genus="Comanthus" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="parvicirrus">Comanthus parvicirrus</taxonomicName>
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:
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</emphasis>
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AM
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<treatmentCitation id="0A964877FFA3A17DFE31DF941254FB66" author="Clark" box="[448,619,1189,1211]" page="395" pageId="29" pageNumber="180" year="1975">
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<bibRefCitation id="EFA61397FFA3A17DFE31DF941204FB66" author="Clark" box="[448,571,1189,1211]" pageId="29" pageNumber="180" refString="Clark, A. M. (1975) The Swain Reefs expedition: Crinoidea. Records of the Australian Museum, 29, 391 - 406. https: // doi. org / 10.3853 / j. 0067 - 1975.29.1975.189" type="journal article" year="1975">Clark, 1975</bibRefCitation>
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: 395
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</treatmentCitation>
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(part).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D3DEDFFA3A17DFF66DFD613C1F99A" pageId="29" pageNumber="180" type="discussion">
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<paragraph id="8B886E66FFA3A17DFF66DFD613C1F99A" blockId="29.[151,1436,1255,1608]" pageId="29" pageNumber="180">
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<emphasis id="B943B274FFA3A17DFF66DFD61134FADD" bold="true" box="[151,267,1255,1280]" pageId="29" pageNumber="180">Remarks.</emphasis>
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Molecular data returned six specimens diagnosed morphologically as
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<taxonomicName id="4C3715E5FFA3A17DFBD0DFD91576FADD" box="[1057,1353,1256,1280]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="albinotus">
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<emphasis id="B943B274FFA3A17DFBD0DFD91576FADD" box="[1057,1353,1256,1280]" italics="true" pageId="29" pageNumber="180">Clarkcomanthus albinotus</emphasis>
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</taxonomicName>
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and 14 of
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<taxonomicName id="4C3715E5FFA3A17DFF45DE3D1103FAF9" box="[180,316,1292,1316]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="littoralis">
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<emphasis id="B943B274FFA3A17DFF45DE3D1103FAF9" box="[180,316,1292,1316]" italics="true" pageId="29" pageNumber="180">Cl. littoralis</emphasis>
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</taxonomicName>
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as a single clade with <2.3% COI divergence (
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<figureCitation id="130C72E3FFA3A17DFCB7DE3D13A7FAF9" box="[838,920,1292,1317]" captionStart="FIGURE 14" captionStartId="28.[151,250,1836,1858]" captionTargetBox="[166,1421,211,1805]" captionTargetId="figure@28.[150,1436,193,1815]" captionTargetPageId="28" captionText="FIGURE 14. Maximum likelihood tree of Clarkcomanthus inferred from COI. Symbols, location abbreviations, and labels as in Figure 2. New COI sequences are Genbank KR 010190 - KR 010244." httpUri="https://zenodo.org/record/580187/files/figure.png" pageId="29" pageNumber="180">Fig. 14</figureCitation>
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). The
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<taxonomicName id="4C3715E5FFA3A17DFC2FDE3D1459FAF9" box="[990,1126,1292,1316]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="littoralis">
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<emphasis id="B943B274FFA3A17DFC2FDE3D1459FAF9" box="[990,1126,1292,1316]" italics="true" pageId="29" pageNumber="180">Cl. littoralis</emphasis>
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</taxonomicName>
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specimens ranged from 10- armed with 24 cirri to 38-armed with four cirri. The inclusion of the specimens previously identified as
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<taxonomicName id="4C3715E5FFA3A17DFA8BDE0110C7FAB1" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="littoralis">
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<emphasis id="B943B274FFA3A17DFA8BDE0110C7FAB1" italics="true" pageId="29" pageNumber="180">Cl. littoralis</emphasis>
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</taxonomicName>
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expands the number of cirri found in
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<taxonomicName id="4C3715E5FFA3A17DFD3BDE651360FAB1" box="[714,863,1364,1388]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="albinotus">
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<emphasis id="B943B274FFA3A17DFD3BDE651360FAB1" box="[714,863,1364,1388]" italics="true" pageId="29" pageNumber="180">Cl. albinotus</emphasis>
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</taxonomicName>
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from 0–6 to 0–24 and includes specimens with pinnulars of approximately equal width and length.
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<taxonomicName id="4C3715E5FFA3A17DFD02DE491420FA4D" box="[755,1055,1400,1424]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="albinotus">
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<emphasis id="B943B274FFA3A17DFD02DE491420FA4D" box="[755,1055,1400,1424]" italics="true" pageId="29" pageNumber="180">Clarkcomanthus albinotus</emphasis>
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</taxonomicName>
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was previously distinguished by having middle pinnulars of PII (P D in Rowe
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<emphasis id="B943B274FFA3A17DFD7CDEAC12F9FA69" box="[653,710,1436,1460]" italics="true" pageId="29" pageNumber="180">et al.</emphasis>
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1986) longer than wide, and the following color patterns: khaki brown, mostly green, or paler or orange pinnule tips. The
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<taxonomicName id="4C3715E5FFA3A17DFCC3DEF21382FA06" box="[818,957,1475,1499]" class="Crinoidea" family="Comasteridae" genus="Clarkcomanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="29" pageNumber="180" phylum="Echinodermata" rank="species" species="littoralis">
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<emphasis id="B943B274FFA3A17DFCC3DEF21382FA06" box="[818,957,1475,1499]" italics="true" pageId="29" pageNumber="180">Cl. littoralis</emphasis>
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</taxonomicName>
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specimens recovered in this clade exhibit color patterns found among Rowe
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<emphasis id="B943B274FFA3A17DFDD2DED91263FA22" box="[547,604,1511,1535]" italics="true" pageId="29" pageNumber="180">et al.</emphasis>
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’s (1986) group 1 specimens: green to brown-green, often with yellow or brown markings, but also ranging from uniformly brown to yellow. The species is common on the reef flat and crest, where it often forms dense aggregations among living branching corals.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |