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<document id="CEAEDFBCB5E77FF0718E243BA867F480" ID-DOI="10.11646/zootaxa.4039.4.5" ID-GBIF-Dataset="5d48e0d6-4af9-422e-af66-81503dd42524" ID-ISSN="1175-5326" ID-Zenodo-Dep="241183" ID-ZooBank="C6FD1263-3CAB-47F3-8661-700612D0B59A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461333100469" checkinUser="plazi" docAuthor="Huang, Hao, Bartolozzi, Luca &amp; Chen, Chang-Chin" docDate="2015" docId="0393D10F5A697550D39FAF26F150F8CC" docLanguage="en" docName="zt04039p565.pdf" docOrigin="Zootaxa 4039 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Echinoaesalus hidakai" docType="treatment" docVersion="9" lastPageNumber="554" masterDocId="FFAAA9775A687551D308AE67F265FC59" masterDocTitle="Notes on the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae), with the description of a new species from the Malay Peninsula" masterLastPageNumber="565" masterPageNumber="553" pageNumber="554" updateTime="1698618809578" updateUser="plazi">
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<mods:title id="04759673A684D9A40D5DBCA854445E73">Notes on the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae), with the description of a new species from the Malay Peninsula</mods:title>
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<mods:namePart id="1568C67039BB45AAF967E8462C1C09C2">Huang, Hao</mods:namePart>
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<mods:namePart id="D0CD4F10C479E3E1027640EE91A15089">Bartolozzi, Luca</mods:namePart>
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<mods:namePart id="A99CDECD25F1B9E2C97C00CEF88FC942">Chen, Chang-Chin</mods:namePart>
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<mods:date id="E74BC8CCB4C67420878311434459FA9F">2015</mods:date>
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<mods:number id="A7D3D8A52BF2CE5E2ADE676BDD3C667C">4039</mods:number>
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<mods:identifier id="B8A21D305D058A1F3F22FBE9FC744642" type="DOI">10.11646/zootaxa.4039.4.5</mods:identifier>
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<treatment id="0393D10F5A697550D39FAF26F150F8CC" ID-DOI="http://doi.org/10.5281/zenodo.6097382" ID-GBIF-Taxon="119644247" ID-Zenodo-Dep="6097382" LSID="urn:lsid:plazi:treatment:0393D10F5A697550D39FAF26F150F8CC" httpUri="http://treatment.plazi.org/id/0393D10F5A697550D39FAF26F150F8CC" lastPageNumber="554" pageId="1" pageNumber="554">
<subSubSection id="C32033925A697550D39FAF26F0A6FD25" pageId="1" pageNumber="554" type="nomenclature">
<paragraph id="8B8560195A697550D39FAF26F170FD02" blockId="1.[151,789,321,381]" box="[151,789,321,347]" pageId="1" pageNumber="554">
<heading id="D0CDD7755A697550D39FAF26F170FD02" bold="true" box="[151,789,321,347]" fontSize="11" level="1" pageId="1" pageNumber="554" reason="1">
<emphasis id="B94EBC0B5A697550D39FAF26F170FD02" bold="true" box="[151,789,321,347]" pageId="1" pageNumber="554">
<taxonomicName id="4C3A1B9A5A697550D39FAF26F3CEFD02" ID-CoL="7GQN4" box="[151,427,321,347]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="hidakai">
<emphasis id="B94EBC0B5A697550D39FAF26F3CEFD02" bold="true" box="[151,427,321,347]" italics="true" pageId="1" pageNumber="554">Echinoaesalus hidakai</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAB1DE85A697550D2B2AF26F168FD02" author="Araya" box="[442,781,321,347]" pageId="1" pageNumber="554" refString="Araya, K., Kon, M. &amp; Johki, Y. (1993) A new species of the genus Aesalus (Coleoptera, Lucanidae) from the Malay Peninsula. Elytra, 21, 93 - 107." type="journal article" year="1993">Araya, Kon, &amp; Johki, 1993</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8560195A697550D39FAF03F0A6FD25" blockId="1.[151,789,321,381]" box="[151,707,356,381]" pageId="1" pageNumber="554">
(
<figureCitation id="13017C9C5A697550D397AF03F289FD25" box="[159,236,356,381]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1750,1773]" captionTargetBox="[183,1395,205,1699]" captionTargetId="figure@2.[183,1395,193,1713]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Antenna. 1, Echinoaesalus hidakai, L 12, western Malaysia; 2, E. borneoensis, L 11, Borneo; 3, E. timidus, L 19, western Malaysia; 4, E. sabahensis, L 14, Borneo; 5, E. gedeensis, holotype, Java; 6, E. cechovskyi, holotype, western Malaysia; 7, E. cechovskyi, paratype, Singapore. Arrow directed to end of third antennomere." httpUri="https://zenodo.org/record/241184/files/figure.png" pageId="1" pageNumber="554">Figs. 1</figureCitation>
,
<figureCitation id="13017C9C5A697550D3F2AF03F351FD25" box="[250,308,356,380]" captionStart="FIGURES 8 – 30" captionStartId="2.[151,264,1842,1865]" captionText="FIGURES 8 – 30. Mentum under same scale (scale bar 0.5 mm). 8, Echinoaesalus hidakai, L 12, western Malaysia; 9, E. borneoensis, L 01, Borneo; 10, E. hidakai, L 10, western Malaysia; 11, E. borneoensis, L 07, Borneo; 12, E. sabahensis, after Krikken (2008), Borneo; 13, E. sabahensis, L 14, Borneo; 14, E. cechovskyi, holotype, western Malaysia; 15, E. cechovskyi, paratype, Singapore; 16 – 18, E. timidus; 16, L 21; 17, L 22; 18, L 18; 19, E. javanus, after Krikken (2008), holotype, Java; 20, E. gedeensis, holotype, Java; 21, E. timidus, after Krikken (2008), holotype, probably male; 22 – 30, E. timidus; 22, L 08; 23, L 02; 24, L 17; 25, L 23; 26, L 09; 27, L 06; 28, L 20; 29, L 13; 30, L 29." pageId="1" pageNumber="554">8, 10</figureCitation>
,
<figureCitation id="13017C9C5A697550D248AF03F3ECFD25" box="[320,393,356,380]" captionStart="FIGURES 31 – 34" captionStartId="3.[151,264,1761,1784]" captionText="FIGURES 31 – 34. Labium (left half, with ligula) and maxilla (right half) under same scale (scale bar 0.2 mm). 31, Echinoaesalus hidakai, L 12, western Malaysia; 32, E. hidakai, L 10, western Malaysia; 33, E. timidus, L 17, western Malaysia; 34, E. timidus, L 06, Borneo." pageId="1" pageNumber="554">31–32</figureCitation>
,
<figureCitation id="13017C9C5A697550D29CAF03F3B8FD24" box="[404,477,356,381]" captionStart="FIGURES 35 – 52" captionStartId="3.[151,264,1853,1876]" captionTargetBox="[202,1395,193,1742]" captionTargetId="figure@3.[202,1396,193,1744]" captionTargetPageId="3" captionText="FIGURES 35 – 52. Protibia (left), mesotibia (middle), and metatibia (right), or just protibia. 35, Echinoaesalus hidakai, L 12; 36, E. hidakai, L 10; 37, E. borneoensis, L 11; 38, E. borneoensis, L 07; 39 – 48, E. timidus; 39, L 17; 40, L 13; 41, L 08; 42, L 02; 43, L 23; 44, L 21; 45, L 22; 46, L 18; 47, L 09; 48, L 06; 49, E. sabahensis, L 14; 50, E. cechovskyi, holotype; 51, E. cechovskyi, paratype; 52, E. gedeensis, holotype." httpUri="https://zenodo.org/record/241185/files/figure.png" pageId="1" pageNumber="554">35–36</figureCitation>
,
<figureCitation id="13017C9C5A697550D2E0AF03F057FD24" box="[488,562,356,381]" captionStart="FIGURES 57 – 76" captionStartId="4.[151,264,1567,1590]" captionTargetBox="[151,1436,203,1545]" captionTargetId="figure@4.[151,1436,193,1546]" captionTargetPageId="4" captionText="FIGURES 57 – 76. Prosternal process in ventral view. 57, Echinoaesalus hidakai, L 12; 58, E. hidakai, L 10; 59 – 62, E. borneoensis; 59, L 11; 60, L 01; 61, L 07; 62, L 16; 63, E. gedeensis, holotype; 64 – 67, E. timidus; 64, L 08; 65, L 02; 66, L 17; 67, L 21; 68, E. sabahensis, L 14; 69, E. cechovskyi, L 28; 70 – 73, E. timidus; 70, L 09; 71, L 06; 72, L 20; 73, L 13; 74, E. sabahensis, after Krikken 2008; 75, E. cechovskyi, paratype; 76, E. javanus, after Krikken (2008), holotype." httpUri="https://zenodo.org/record/241186/files/figure.png" pageId="1" pageNumber="554">57–58</figureCitation>
, 77, 90, 103)
</paragraph>
</subSubSection>
<subSubSection id="C32033925A697550D39FAFCBF3DBFDB1" pageId="1" pageNumber="554" type="materials_examined">
<paragraph id="8B8560195A697550D39FAFCBF3DBFDB1" blockId="1.[151,1436,428,1173]" pageId="1" pageNumber="554">
<emphasis id="B94EBC0B5A697550D39FAFCBF29AFD9C" bold="true" box="[151,255,428,453]" pageId="1" pageNumber="554">Material</emphasis>
.
<emphasis id="B94EBC0B5A697550D205AFCBF3C6FD9C" bold="true" box="[269,419,428,453]" pageId="1" pageNumber="554">
<collectingCountry id="F32D20895A697550D205AFCBF3C6FD9C" box="[269,419,428,453]" name="Malaysia" pageId="1" pageNumber="554">MALAYSIA</collectingCountry>
</emphasis>
:
<specimenCount id="9D3CAB905A697550D2BBAFCBF3B8FD9D" box="[435,477,428,452]" pageId="1" pageNumber="554" type="generic">1 ♂</specimenCount>
(L12), 1 ♀ (L10), Kelantan,
<quantity id="4CC2CDFC5A697550D02FAFCBF10AFD9C" box="[807,879,428,453]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="554" unit="km" value="60.0">60 km</quantity>
NE of Tanah Rata, Tanah Kerajaan,
<quantity id="4CC2CDFC5A697550D606AFCBF703FD9D" box="[1294,1382,428,452]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="554" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FF8446D95A697550D67AAFCBF373FDB1" pageId="1" pageNumber="554" value="2007-04-12" valueMax="2007-04-30" valueMin="2007-04-12">12– 30.IV.2007</date>
, P. Cechovsky.
</paragraph>
</subSubSection>
<subSubSection id="C32033925A697550D3CFAF93F6EFFF2C" pageId="1" pageNumber="554" type="discussion">
<paragraph id="8B8560195A697550D3CFAF93F716FEE6" blockId="1.[151,1436,428,1173]" pageId="1" pageNumber="554">
<emphasis id="B94EBC0B5A697550D3CFAF93F351FE54" bold="true" box="[199,308,500,525]" pageId="1" pageNumber="554">Remarks</emphasis>
. The following previously unnoticed characters of this species should be noted, in comparison with the species we have examined. Third antennomere similar to that of
<taxonomicName id="4C3A1B9A5A697550D0A5AC7EF630FE69" box="[941,1109,536,560]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="B94EBC0B5A697550D0A5AC7EF630FE69" box="[941,1109,536,560]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A1B9A5A697550D799AC7EF748FE69" box="[1169,1325,536,560]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="B94EBC0B5A697550D799AC7EF748FE69" box="[1169,1325,536,560]" italics="true" pageId="1" pageNumber="554">E. sabahensis</emphasis>
</taxonomicName>
in shape, more elongate than in
<emphasis id="B94EBC0B5A697550D291AC5AF0CFFE0D" box="[409,682,572,596]" italics="true" pageId="1" pageNumber="554">
<taxonomicName id="4C3A1B9A5A697550D291AC5AF06AFE0D" box="[409,527,572,596]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="timidus">E. timidus</taxonomicName>
,
<taxonomicName id="4C3A1B9A5A697550D114AC5AF0CFFE0D" authority="Huang &amp; Wu, 2011" authorityName="Huang &amp; Wu" authorityYear="2011" box="[540,682,572,596]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="gedeensis">E. gedeensis</taxonomicName>
</emphasis>
<bibRefCitation id="EFAB1DE85A697550D1BBAC5AF1F0FE0D" author="Huang" box="[691,917,572,597]" pageId="1" pageNumber="554" refString="Huang, H., Imura, Y. &amp; Wu, L. (2011) Description of three new taxa of Echinoaesalus Zelenka from Borneo and Java (Coleoptera, Lucanidae). Kogane, 12, 97 - 107." type="journal article" year="2011">Huang &amp; Wu, 2011</bibRefCitation>
, and
<taxonomicName id="4C3A1B9A5A697550D0DCAC5AF60BFE0D" box="[980,1134,572,596]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="cechovskyi" status="sp. nov.">
<emphasis id="B94EBC0B5A697550D0DCAC5AF60BFE0D" box="[980,1134,572,596]" italics="true" pageId="1" pageNumber="554">E. cechovskyi</emphasis>
</taxonomicName>
<emphasis id="B94EBC0B5A697550D77FAC5BF766FE0C" bold="true" box="[1143,1283,572,597]" pageId="1" pageNumber="554">
<taxonomicNameLabel id="A27D01705A697550D77FAC5BF766FE0C" box="[1143,1283,572,597]" pageId="1" pageNumber="554" rank="species">new species</taxonomicNameLabel>
</emphasis>
.. Male ligula similar to that of
<taxonomicName id="4C3A1B9A5A697550D250AC06F3A5FE2E" box="[344,448,608,632]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="arayai">
<emphasis id="B94EBC0B5A697550D250AC06F3A5FE2E" box="[344,448,608,632]" italics="true" pageId="1" pageNumber="554">E. arayai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A1B9A5A697550D2FEAC06F0F9FE21" box="[502,668,608,632]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="B94EBC0B5A697550D2FEAC06F0F9FE21" box="[502,668,608,632]" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
</taxonomicName>
, but markedly larger than in
<emphasis id="B94EBC0B5A697550D0EBAC06F361FEC5" italics="true" pageId="1" pageNumber="554">
<taxonomicName id="4C3A1B9A5A697550D0EBAC06F634FE21" box="[995,1105,608,632]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="chungi">E. chungi</taxonomicName>
,
<taxonomicName id="4C3A1B9A5A697550D754AC06F6B5FE21" box="[1116,1232,608,632]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="timidus">E. timidus</taxonomicName>
,
<taxonomicName id="4C3A1B9A5A697550D7D4AC06F71CFE21" box="[1244,1401,608,632]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="sabahensis">E. sabahensis</taxonomicName>
,
<taxonomicName id="4C3A1B9A5A697550D68CAC06F361FEC5" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="gedeensis">E. gedeensis</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C3A1B9A5A697550D249ACE2F3BFFEC5" box="[321,474,644,668]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="cechovskyi">
<emphasis id="B94EBC0B5A697550D249ACE2F3BFFEC5" box="[321,474,644,668]" italics="true" pageId="1" pageNumber="554">E. cechovskyi</emphasis>
</taxonomicName>
. Labial palpus in both sexes with all palpomeres nearly as long as in
<taxonomicName id="4C3A1B9A5A697550D7EDACE2F728FEC5" box="[1253,1357,645,668]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="arayai">
<emphasis id="B94EBC0B5A697550D7EDACE2F728FEC5" box="[1253,1357,645,668]" italics="true" pageId="1" pageNumber="554">E. arayai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A1B9A5A697550D68CACE2F37BFE99" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="borneoensis">
<emphasis id="B94EBC0B5A697550D68CACE2F37BFE99" italics="true" pageId="1" pageNumber="554">E. borneoensis</emphasis>
</taxonomicName>
, but markedly longer than in
<emphasis id="B94EBC0B5A697550D162ACCEF6F2FE99" box="[618,1175,680,704]" italics="true" pageId="1" pageNumber="554">
<taxonomicName id="4C3A1B9A5A697550D162ACCEF0BDFE99" box="[618,728,680,704]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="chungi">E. chungi</taxonomicName>
,
<taxonomicName id="4C3A1B9A5A697550D1ECACCEF13DFE99" box="[740,856,680,704]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="timidus">E. timidus</taxonomicName>
,
<taxonomicName id="4C3A1B9A5A697550D06BACCEF19AFE99" box="[867,1023,680,704]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="sabahensis">E. sabahensis</taxonomicName>
,
<taxonomicName id="4C3A1B9A5A697550D703ACCEF6F2FE99" box="[1035,1175,680,704]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="gedeensis">E. gedeensis</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C3A1B9A5A697550D7DCACCEF709FE99" box="[1236,1388,680,704]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="cechovskyi">
<emphasis id="B94EBC0B5A697550D7DCACCEF709FE99" box="[1236,1388,680,704]" italics="true" pageId="1" pageNumber="554">E. cechovskyi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8560195A697550D3CFACABF6EFFF2C" blockId="1.[151,1436,428,1173]" pageId="1" pageNumber="554">Sexual dimorphism is found in width of head, size of labrum, galea of maxilla, ligula of labium, distance between palp insertions on prementum, size and shape of mentum, length of intercoxal process of prosternum, detailed shape of outer apex of metatibia, and presence or absence of a central projection on anterior margin of last abdominal ventrite; but is not found in shape of tooth, shape and length of third antennomere, detailed apex of protibia, and posterior margin of last visible abdominal ventrite and venation of hindwing.</paragraph>
</subSubSection>
<subSubSection id="C32033925A697550D3CFAD18F000F814" pageId="1" pageNumber="554" type="description">
<paragraph id="8B8560195A697550D3CFAD18F6B7FFE4" blockId="1.[151,1436,428,1173]" pageId="1" pageNumber="554">
<emphasis id="B94EBC0B5A697550D3CFAD18F317FFC1" bold="true" box="[199,370,895,920]" pageId="1" pageNumber="554">Male genitalia</emphasis>
(
<figureCitation id="13017C9C5A697550D28AAD18F3B3FFC1" box="[386,470,895,920]" captionStart="FIGURES 90 – 102" captionStartId="5.[151,264,1797,1820]" captionTargetBox="[151,1436,519,1775]" captionTargetId="figure@5.[151,1436,519,1775]" captionTargetPageId="5" captionText="FIGURES 90 – 102. Aedeagus in dorsal, left lateral, ventral, and right lateral views under same scale (scale bar 1 mm). 90, Echinoaesalus hidakai, L 12, western Malaysia; 91, E. borneoensis, L 11, Borneo; 92, E. borneoensis, L 01, Borneo; 93 – 98, E. timidus; 93, L 08; 94, L 02; 95, L 17; 96, L 23; 97, L 21; 98, L 22; 99, E. sabahensis, L 14, Borneo; 100, E. timidus, L 18; 101, E. cechovskyi, holotype, western Malaysia; 102, E. gedeensis, holotype, Java. Arrows directed to ends of parameres." httpUri="https://zenodo.org/record/241187/files/figure.png" pageId="1" pageNumber="554">Fig. 90</figureCitation>
). The male genitalia examined by us fit the original description (
<bibRefCitation id="EFAB1DE85A697550D7B8ADE7F712FFC1" author="Araya" box="[1200,1399,896,920]" pageId="1" pageNumber="554" refString="Araya, K., Kon, M. &amp; Johki, Y. (1993) A new species of the genus Aesalus (Coleoptera, Lucanidae) from the Malay Peninsula. Elytra, 21, 93 - 107." type="journal article" year="1993">
Araya
<emphasis id="B94EBC0B5A697550D7F5ADE6F753FFC1" box="[1277,1334,896,920]" italics="true" pageId="1" pageNumber="554">et al.</emphasis>
1993
</bibRefCitation>
) in all details. The median lobe of parameres is entirely pigmented, without colorless splits laterally.
</paragraph>
<paragraph id="8B8560195A697550D3CFADA0F000F814" blockId="1.[151,1436,428,1173]" pageId="1" pageNumber="554">
<emphasis id="B94EBC0B5A697550D3CFADA0F3EEFFB9" bold="true" box="[199,395,967,992]" pageId="1" pageNumber="554">Female genitalia</emphasis>
(
<figureCitation id="13017C9C5A697550D292ADA0F399FFB9" box="[410,508,967,992]" captionStart="FIGURES 103 – 106" captionStartId="6.[151,264,1501,1524]" captionTargetBox="[151,1436,193,1480]" captionTargetId="figure@6.[151,1436,193,1481]" captionTargetPageId="6" captionText="FIGURES 103 – 106. Female genitalia under same scale (scale bar 1 mm). 103, Echinoaesalus hidakai, L 10; 104 – 106, E. borneoensis; 104, L 16; 105, L 07; 106, L 27. Abbreviations: st, stylus; lv, last abdominal ventrite; mo, median oviduct; bd, bursal duct; bc, bursa copulatrix; sd, spermathecal duct; s, spermatheca; sg, spermathecal gland; ag, accessory gland." httpUri="https://zenodo.org/record/241188/files/figure.png" pageId="1" pageNumber="554">Fig. 103</figureCitation>
). Accessory gland rooted just on terminal end of bursa copulatrix. Bursa copulatrix nearly as large as accessory gland when fully inflated, easily separable from bursal duct in width. Bursal duct rather long and wide. Spermatheca and its duct originated near the beginning of bursa copulatrix. Spermathecal duct rather long, nearly as long as bursal duct.
</paragraph>
</subSubSection>
<subSubSection id="C32033925A697550D3CFAA30F150F8CC" pageId="1" pageNumber="554" type="distribution">
<paragraph id="8B8560195A697550D3CFAA30F150F8CC" blockId="1.[151,1436,428,1173]" pageId="1" pageNumber="554">
<emphasis id="B94EBC0B5A697550D3CFAA30F33CF829" bold="true" box="[199,345,1111,1136]" pageId="1" pageNumber="554">Distribution</emphasis>
. Restricted to Malay Peninsula. The previously described subspecies,
<taxonomicName id="4C3A1B9A5A697550D79AAA3EF7FEF829" box="[1170,1435,1112,1136]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="554" phylum="Arthropoda" rank="subSpecies" species="hidakai" subSpecies="borneoensis">
<emphasis id="B94EBC0B5A697550D79AAA3EF6C6F829" box="[1170,1187,1113,1136]" italics="true" pageId="1" pageNumber="554">E</emphasis>
.
<emphasis id="B94EBC0B5A697550D7BDAA3FF7FEF829" box="[1205,1435,1112,1136]" italics="true" pageId="1" pageNumber="554">hidakai borneoensis</emphasis>
</taxonomicName>
from Borneo deserves full specific rank as discussed below.
</paragraph>
</subSubSection>
</treatment>
</document>
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