761 lines
146 KiB
XML
761 lines
146 KiB
XML
<document id="92AD31CBF6496E146749B8945357EC0A" ID-CLB-Dataset="30883" ID-DOI="10.11646/zootaxa.4377.1.1" ID-GBIF-Dataset="fbc66cd8-6a7b-4cf3-ac3f-850b17b3abe4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1163170" ID-ZooBank="FE92756E-2212-4ADE-9FE0-31C30511583E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517396002872" checkinUser="plazi" docAuthor="Okanishi, Masanori & Fujita, Yoshihisa" docDate="2018" docId="0393AD4FFFC93413FF1AFC3FFDC9FD7B" docLanguage="en" docName="zootaxa.4377.1.1.pdf" docOrigin="Zootaxa 4377 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophiozonella cavernalis Okanishi & Fujita 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FFAAD537FFC33402FF8DFF91FF9FFF8B" masterDocTitle="First finding of anchialine and submarine cave dwelling brittle stars from the Pacific Ocean, with descriptions of new species of Ophiolepis and Ophiozonella (Echinodermata: Ophiuroidea: Amphilepidida)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="11" updateTime="1698517853098" updateUser="ExternalLinkService">
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<mods:mods id="F47EBE920CC35967FDD9F3D276850F26" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A63F958BB37C598022413DFA29A42221">
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<mods:title id="336EC59AB1F848B9563C137D2BEC8CF1">First finding of anchialine and submarine cave dwelling brittle stars from the Pacific Ocean, with descriptions of new species of Ophiolepis and Ophiozonella (Echinodermata: Ophiuroidea: Amphilepidida)</mods:title>
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</mods:titleInfo>
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<mods:name id="16976D7C59D86F2CC8FB91F4833A426D" type="personal">
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<mods:roleTerm id="E1E9F8AA9893F625C6D933F7F759C834">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="381B769E0292314274FEF99F171DCE80">Okanishi, Masanori</mods:namePart>
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</mods:name>
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<mods:name id="4C2F0536784EA1B54BB11C0905A0A612" type="personal">
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<mods:role id="B4C114EA4B2DB85F0ED654C73E6CFBBF">
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<mods:roleTerm id="E958889FC9B5CD2DFE4EF6693AFA4E8F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E3E3E60E3A646A97AEF04F352D7CC51B">Fujita, Yoshihisa</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E87F17EDC036742B8B23D95CAFE33D6E">text</mods:typeOfResource>
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<mods:titleInfo id="C50200C2E5C8106A842D461ADCFBBAC3">
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<mods:title id="25DCC78BC4A31EC4A9435C637C17D6CB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C2A20447D4CDC612471385E802F6D0D0">
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<mods:date id="F3A0CFAC7BF7865B8A49B5F350238A26">2018</mods:date>
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<mods:number id="7B099624E79F263F4F03751AB4B2D129">2018-01-31</mods:number>
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</mods:detail>
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<mods:detail id="F733B8B9387E19EC135A0762C385E972" type="volume">
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<mods:number id="F09F7B0F755E1C0706A1473C1C4D33DD">4377</mods:number>
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</mods:detail>
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<mods:detail id="B98D6EB6D5DD22A9240F276F01B16AD2" type="issue">
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<mods:number id="5381633AE6F6B6951EDE941AF9A7912D">1</mods:number>
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<mods:start id="40A64BFB703F6B1BA9E61FE555677693">1</mods:start>
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<mods:classification id="5496BE65CAD4521F25EF9EFA9EBBFC04">journal article</mods:classification>
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<mods:identifier id="7CE97294349CD30C8BE2110795FC656A" type="CLB-Dataset">30883</mods:identifier>
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<mods:identifier id="A088C2E6CC5135899B85EC3D13CDBD2C" type="DOI">10.11646/zootaxa.4377.1.1</mods:identifier>
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<mods:identifier id="23C55C2846D724252441BC1C8FB58558" type="GBIF-Dataset">fbc66cd8-6a7b-4cf3-ac3f-850b17b3abe4</mods:identifier>
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<mods:identifier id="F806364E4C2978B22789D3205EFAC8DF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="776A132DF20E7BBDEA8FA57D02B8E570" type="Zenodo-Dep">1163170</mods:identifier>
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</mods:mods>
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<treatment id="0393AD4FFFC93413FF1AFC3FFDC9FD7B" ID-DOI="http://doi.org/10.5281/zenodo.3509141" ID-GBIF-Taxon="140561232" ID-Zenodo-Dep="3509141" LSID="urn:lsid:plazi:treatment:0393AD4FFFC93413FF1AFC3FFDC9FD7B" httpUri="http://treatment.plazi.org/id/0393AD4FFFC93413FF1AFC3FFDC9FD7B" lastPageId="17" lastPageNumber="18" pageId="10" pageNumber="11">
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<subSubSection id="C3204FD2FFC93408FF1AFC3FFE88FB85" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8B851C59FFC93408FF1AFC3FFD9AFC43" blockId="10.[151,715,942,1038]" box="[151,517,942,968]" pageId="10" pageNumber="11">
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<heading id="D0CDAB35FFC93408FF1AFC3FFD9AFC43" bold="true" box="[151,517,942,968]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="B94EC04BFFC93408FF1AFC3FFD9AFC43" bold="true" box="[151,517,942,968]" pageId="10" pageNumber="11">
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<taxonomicName id="4C3A67DAFFC93408FF1AFC3FFE24FC43" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[151,443,942,968]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
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<emphasis id="B94EC04BFFC93408FF1AFC3FFE24FC43" bold="true" box="[151,443,942,968]" italics="true" pageId="10" pageNumber="11">Ophiozonella cavernalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27D7D30FFC93408FE4FFC3FFD9AFC43" box="[450,517,942,968]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B851C59FFC93408FF1AFC43FE88FB85" blockId="10.[151,715,942,1038]" pageId="10" pageNumber="11">
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[Japanese Name: Dohkutsu-mozaiku-kumohitode] (
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<figureCitation id="130100DCFFC93408FF12FC64FE8FFB85" box="[159,272,1013,1038]" captionStart-0="FIGURE7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStartId-0="11.[151,250,1419,1441]" captionStartId-1="12.[151,250,1916,1938]" captionStartId-2="14.[151,250,1913,1935]" captionStartId-3="15.[151,250,1794,1816]" captionStartId-4="16.[151,250,1833,1855]" captionTargetBox-0="[164,1426,530,1407]" captionTargetBox-1="[152,1435,409,1895]" captionTargetBox-2="[247,1358,193,1894]" captionTargetBox-3="[234,1343,193,1767]" captionTargetBox-4="[221,1356,193,1805]" captionTargetId-0="figure@11.[151,1436,517,1407]" captionTargetId-1="figure@12.[151,1436,409,1895]" captionTargetId-2="figure@14.[228,1375,193,1894]" captionTargetId-3="figure@15.[234,1352,193,1768]" captionTargetId-4="figure@16.[221,1356,193,1818]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="16" captionText-0="FIGURE7. Ophiozonella cavernalisn. sp. A, living status insitu, aboral view of RUMF-ZE-02008,RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth." captionText-1="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." captionText-2="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." captionText-3="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." captionText-4="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri-0="https://zenodo.org/record/1163184/files/figure.png" httpUri-1="https://zenodo.org/record/1163186/files/figure.png" httpUri-2="https://zenodo.org/record/1163188/files/figure.png" httpUri-3="https://zenodo.org/record/1163190/files/figure.png" httpUri-4="https://zenodo.org/record/1163192/files/figure.png" pageId="10" pageNumber="11">Figs 7–11</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3204FD2FFC93408FF1AFBACFDEEF91D" pageId="10" pageNumber="11" type="materials_examined">
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||
<paragraph id="8B851C59FFC93408FF1AFBACFEC1FB15" blockId="10.[151,1437,1085,2010]" pageId="10" pageNumber="11">
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||
<materialsCitation id="3B521604FFC93408FF1AFBACFEC1FB15" ID-GBIF-Occurrence="1806155411" collectingDate="2016-08-10" collectionCode="SCUBA" collectorName="By Y. & Y. Fujita." country="Japan" elevation="10" location="Hedo-Dome" pageId="10" pageNumber="11" specimenCount="2" stateProvince="Okinawa" typeStatus="holotype">
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||
<emphasis id="B94EC04BFFC93408FF1AFBACFED6FBDD" bold="true" box="[151,329,1085,1110]" pageId="10" pageNumber="11">
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||
<typeStatus id="5481A2FBFFC93408FF1AFBACFF4DFBDD" box="[151,210,1085,1110]" pageId="10" pageNumber="11">Type</typeStatus>
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material.
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</emphasis>
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||
<typeStatus id="5481A2FBFFC93408FED9FBACFE5EFBDD" box="[340,449,1085,1110]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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: RUMF-ZE-02008, submarine cave (
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||
<location id="8EE54A82FFC93408FCFDFBACFC63FBDD" LSID="urn:lsid:plazi:treatment:0393AD4FFFC93413FF1AFC3FFDC9FD7B:8EE54A82FFC93408FCFDFBACFC63FBDD" box="[880,1020,1085,1110]" country="Japan" name="Hedo-Dome" pageId="10" pageNumber="11" stateProvince="Okinawa">Hedo-Dome</location>
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cave) on northern
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<collectingRegion id="49FED2BBFFC93408FB6DFBACFADAFBDD" box="[1248,1349,1085,1110]" country="Japan" name="Okinawa" pageId="10" pageNumber="11">Okinawa</collectingRegion>
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Island, southwestern
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<collectingCountry id="F32D5CC9FFC93408FEBEFBF3FEECFBF2" box="[307,371,1122,1145]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
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||
(26˚51’53.74”N, 128˚14’43.93”E.
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<figureCitation id="130100DCFFC93408FC89FBF0FCD9FBF1" box="[772,838,1121,1146]" captionStart="FIGURE 1" captionStartId="1.[151,250,1946,1968]" captionTargetBox="[207,1389,627,1885]" captionTargetId="figure@1.[197,1390,626,1921]" captionTargetPageId="1" captionText="FIGURE 1. Sampling localities of Ophiolepis cavitata n. sp. and Ophiozonella cavernalis n. sp. Red symbols indicate type localities." httpUri="https://zenodo.org/record/1163172/files/figure.png" pageId="10" pageNumber="11">Fig. 1</figureCitation>
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||
), c.a.
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<elevation id="0017FB6AFFC93408FC1DFBF3FC54FBF2" box="[912,971,1122,1146]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="10.0">
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||
<quantity id="4CC2B1BCFFC93408FC1DFBF3FC54FBF2" box="[912,971,1122,1146]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="10.0">10 m</quantity>
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</elevation>
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||
depth,
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<collectingDate id="EFC0C371FFC93408FBACFBF3FB44FBF1" box="[1057,1243,1121,1146]" pageId="10" pageNumber="11" value="2016-08-10">
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||
<date id="FF843A99FFC93408FBACFBF3FB44FBF1" box="[1057,1243,1121,1146]" pageId="10" pageNumber="11" value="2016-08-10">10 August 2016</date>
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</collectingDate>
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,
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<collectionCode id="ED2B849CFFC93408FB65FBF3FADAFBF1" box="[1256,1349,1122,1146]" pageId="10" pageNumber="11">SCUBA</collectionCode>
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diving, coll.
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<collectorName id="26CF798FFFC93408FF42FB17FE90FB16" box="[207,271,1158,1182]" pageId="10" pageNumber="11">By Y.</collectorName>
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Fujita.
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</materialsCitation>
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||
</paragraph>
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<paragraph id="8B851C59FFC93408FF4AFB3BFDEEF91D" blockId="10.[151,1437,1085,2010]" pageId="10" pageNumber="11">
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||
<materialsCitation id="3B521604FFC93408FF4AFB3BFAA9FB49" ID-GBIF-Occurrence="1806155419" box="[199,1334,1193,1218]" collectingDate="2016-08-10" collectionCode="SCUBA" collectorName="By Y. & Y. Fujita." country="Japan" elevation="10" location="Hedo-Dome" pageId="10" pageNumber="11" specimenCount="19" stateProvince="Okinawa" typeStatus="paratype">
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||
<typeStatus id="5481A2FBFFC93408FF4AFB3BFEABFB49" box="[199,308,1194,1218]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
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||
(
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||
<specimenCount id="9D3CD7D0FFC93408FECBFB3BFE79FB49" box="[326,486,1193,1218]" pageId="10" pageNumber="11" type="generic">18 specimens</specimenCount>
|
||
).
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||
<collectingRegion id="49FED2BBFFC93408FE77FB38FDC0FB49" box="[506,607,1193,1218]" country="Japan" name="Okinawa" pageId="10" pageNumber="11">Okinawa</collectingRegion>
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||
Island: RUMF-ZE-02011 (
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||
<specimenCount id="9D3CD7D0FFC93408FC11FB3BFB80FB49" box="[924,1055,1193,1218]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), same data as holotype
|
||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B521604FFC93408FACEFB3BFD81FAA5" ID-GBIF-Occurrence="1806155412" collectingDate="2017-05-21" collectionCode="SCUBA" collectorName="By Y. & Fujita & M. Okanishi" country="Japan" elevation="12" location="Hedo-Dome" pageId="10" pageNumber="11" specimenCount="5" stateProvince="Okinawa" typeStatus="paratype">
|
||
RUMF- ZE-02009 (
|
||
<specimenCount id="9D3CD7D0FFC93408FE93FB5FFE3CFB6D" box="[286,419,1229,1254]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), RUMF-ZE-02010 (
|
||
<specimenCount id="9D3CD7D0FFC93408FD14FB5FFC81FB6D" box="[665,798,1229,1254]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), RUMF-ZE-02012 (
|
||
<specimenCount id="9D3CD7D0FFC93408FB99FB5FFB06FB6D" box="[1044,1177,1229,1254]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), RUMF-ZE-02013 (
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||
<specimenCount id="9D3CD7D0FFC93408FA1DFB5FFE9AFA81" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), RUMF-ZE-02014 (
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||
<specimenCount id="9D3CD7D0FFC93408FE78FB63FDE8FA81" box="[501,631,1265,1290]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), same locality as holotype,
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||
<elevation id="0017FB6AFFC93408FC3BFB63FC6FFA82" box="[950,1008,1266,1290]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="m" value="12.0">
|
||
<quantity id="4CC2B1BCFFC93408FC3BFB63FC6FFA82" box="[950,1008,1266,1290]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="m" value="12.0">12 m</quantity>
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||
</elevation>
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||
depth,
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||
<collectingDate id="EFC0C371FFC93408FBC9FB63FB42FA81" box="[1092,1245,1266,1290]" pageId="10" pageNumber="11" value="2017-05-21">
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||
<date id="FF843A99FFC93408FBC9FB63FB42FA81" box="[1092,1245,1266,1290]" pageId="10" pageNumber="11" value="2017-05-21">21 May 2017</date>
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</collectingDate>
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||
,
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||
<collectionCode id="ED2B849CFFC93408FB64FB63FAD9FA81" box="[1257,1350,1266,1290]" pageId="10" pageNumber="11">SCUBA</collectionCode>
|
||
diving, coll.
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||
<collectorName id="26CF798FFFC93408FF42FA87FEC9FAA5" box="[207,342,1301,1326]" pageId="10" pageNumber="11">By Y. Fujita</collectorName>
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and
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||
<collectorName id="26CF798FFFC93408FE03FA87FD81FAA5" box="[398,542,1301,1326]" pageId="10" pageNumber="11">M. Okanishi</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B521604FFC93408FDA4FA87FF4DFAD9" ID-GBIF-Occurrence="1806155418" collectingDate="2017-05-21" collectionCode="NSMT" collectorName="By Y. & Y. Fujita." country="Japan" elevation="12" location="Hedo-Dome" pageId="10" pageNumber="11" specimenCount="2" stateProvince="Okinawa" typeStatus="paratype">
|
||
<collectionCode id="ED2B849CFFC93408FDA4FA87FDE8FAA5" box="[553,631,1302,1326]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="10" pageNumber="11">NSMT</collectionCode>
|
||
E-10711 (
|
||
<specimenCount id="9D3CD7D0FFC93408FD60FA87FCE3FAA5" box="[749,892,1301,1326]" pageId="10" pageNumber="11" type="generic">2 specimens</specimenCount>
|
||
), same locality as holotype,
|
||
<elevation id="0017FB6AFFC93408FB34FA87FB6CFAA6" box="[1209,1267,1302,1326]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="m" value="12.0">
|
||
<quantity id="4CC2B1BCFFC93408FB34FA87FB6CFAA6" box="[1209,1267,1302,1326]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="m" value="12.0">12 m</quantity>
|
||
</elevation>
|
||
depth,
|
||
<collectingDate id="EFC0C371FFC93408FACBFA87FF4DFAD9" pageId="10" pageNumber="11" value="2017-05-21">
|
||
<date id="FF843A99FFC93408FACBFA87FF4DFAD9" pageId="10" pageNumber="11" value="2017-05-21">21 May 2017</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B521604FFC93408FF53FAABFA80FAD9" ID-GBIF-Occurrence="1806155414" box="[222,1311,1337,1362]" collectionCode="SCUBA" collectorName="By Y. & Fujita" country="Iceland" location="Okanishi. Ie Island" pageId="10" pageNumber="11" specimenCount="5" typeStatus="paratype">
|
||
<collectionCode id="ED2B849CFFC93408FF53FAABFEA4FAD9" box="[222,315,1338,1362]" pageId="10" pageNumber="11">SCUBA</collectionCode>
|
||
diving, coll.
|
||
<collectorName id="26CF798FFFC93408FE5BFAABFDFFFAD9" box="[470,608,1337,1362]" pageId="10" pageNumber="11">By Y. Fujita</collectorName>
|
||
and M.
|
||
<location id="8EE54A82FFC93408FD4EFAA8FC00FAD9" LSID="urn:lsid:plazi:treatment:0393AD4FFFC93413FF1AFC3FFDC9FD7B:8EE54A82FFC93408FD4EFAA8FC00FAD9" box="[707,927,1337,1362]" country="Iceland" name="Okanishi. Ie Island" pageId="10" pageNumber="11">
|
||
Okanishi. Ie
|
||
<collectingCountry id="F32D5CC9FFC93408FCDBFAA8FC00FAD9" box="[854,927,1337,1362]" name="Iceland" pageId="10" pageNumber="11">Island</collectingCountry>
|
||
</location>
|
||
: RUMF-ZE-02015 (
|
||
<specimenCount id="9D3CD7D0FFC93408FB0AFAA8FA88FAD9" box="[1159,1303,1337,1362]" pageId="10" pageNumber="11" type="generic">5 specimens</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B521604FFC93408FAA6FAABFC3CFA11" ID-GBIF-Occurrence="1806155416" collectingDate="2017-06-25" collectionCode="NSMT" collectorName="Dai-Dohkutsu" country="Japan" elevation="27" location="northeastern Ie Island" pageId="10" pageNumber="11" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2B849CFFC93408FAA6FAABFAE6FAD9" box="[1323,1401,1338,1362]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="10" pageNumber="11">NSMT</collectionCode>
|
||
E- 10712 (
|
||
<specimenCount id="9D3CD7D0FFC93408FF74FACFFE1BFAFD" box="[249,388,1373,1398]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), submarine cave (
|
||
<collectorName id="26CF798FFFC93408FDFEFACCFC8BFAFD" box="[627,788,1373,1398]" pageId="10" pageNumber="11">Dai-Dohkutsu</collectorName>
|
||
cave) at
|
||
<location id="8EE54A82FFC93408FC15FACCFB3DFAFD" LSID="urn:lsid:plazi:treatment:0393AD4FFFC93413FF1AFC3FFDC9FD7B:8EE54A82FFC93408FC15FACCFB3DFAFD" box="[920,1186,1373,1398]" country="Japan" name="northeastern Ie Island" pageId="10" pageNumber="11">northeastern Ie Island</location>
|
||
, southwestern
|
||
<collectingCountry id="F32D5CC9FFC93408FAD1FACFFA03FAFE" box="[1372,1436,1374,1397]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
|
||
(26˚43’27.55’N, 127˚49’53.50”E), c.a.
|
||
<elevation id="0017FB6AFFC93408FDECFA13FD3FFA12" box="[609,672,1410,1434]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="11" unit="m" value="27.0">
|
||
<quantity id="4CC2B1BCFFC93408FDECFA13FD3FFA12" box="[609,672,1410,1434]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="11" unit="m" value="27.0">27 m</quantity>
|
||
</elevation>
|
||
depth,
|
||
<collectingDate id="EFC0C371FFC93408FD73FA10FC3CFA11" box="[766,931,1409,1434]" pageId="10" pageNumber="11" value="2017-06-25">
|
||
<date id="FF843A99FFC93408FD73FA10FC3CFA11" box="[766,931,1409,1434]" pageId="10" pageNumber="11" value="2017-06-25">25 June 2017</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B521604FFC93408FC3EFA10FEADFA69" ID-GBIF-Occurrence="1806155413" collectingDate="2017-06-25" country="Japan" elevation="20" location="northeastern Ie Island" pageId="10" pageNumber="11" specimenCount="1" typeStatus="paratype">
|
||
RUMF-ZE-02016 (
|
||
<specimenCount id="9D3CD7D0FFC93408FB18FA13FA82FA11" box="[1173,1309,1409,1434]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), unnamed submarine cave at
|
||
<location id="8EE54A82FFC93408FEE0FA34FDFAFA35" LSID="urn:lsid:plazi:treatment:0393AD4FFFC93413FF1AFC3FFDC9FD7B:8EE54A82FFC93408FEE0FA34FDFAFA35" box="[365,613,1445,1470]" country="Japan" name="northeastern Ie Island" pageId="10" pageNumber="11">northeastern Ie Island</location>
|
||
, southwestern
|
||
<collectingCountry id="F32D5CC9FFC93408FC83FA37FCD1FA36" box="[782,846,1446,1469]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
|
||
(26˚43’26.31’N, 127˚49’54.86”E), c.a.
|
||
<elevation id="0017FB6AFFC93408FA99FA37FAD1FA36" box="[1300,1358,1446,1470]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="20.0">
|
||
<quantity id="4CC2B1BCFFC93408FA99FA37FAD1FA36" box="[1300,1358,1446,1470]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="20.0">20 m</quantity>
|
||
</elevation>
|
||
depth,
|
||
<collectingDate id="EFC0C371FFC93408FF1AFA58FEADFA69" box="[151,306,1481,1506]" pageId="10" pageNumber="11" value="2017-06-25">
|
||
<date id="FF843A99FFC93408FF1AFA58FEADFA69" box="[151,306,1481,1506]" pageId="10" pageNumber="11" value="2017-06-25">25 June 2017</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B521604FFC93408FEB3FA5BFC63F9A1" ID-GBIF-Occurrence="1806155420" collectingDate="2017-02-04" collectionCode="SCUBA" collectorName="By Y. & Akuma-no-Yakata & Miyako-Island Group" country="Japan" county="Fujita. Shimoji Island" elevation="20" location="Ryukyu Islands" municipality="southern Shimoji Island" pageId="10" pageNumber="11" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2B849CFFC93408FEB3FA5BFE04FA69" box="[318,411,1482,1506]" pageId="10" pageNumber="11">SCUBA</collectionCode>
|
||
diving, coll.
|
||
<collectorName id="26CF798FFFC93408FDB8FA5BFDE9FA6A" box="[565,630,1482,1506]" pageId="10" pageNumber="11">By Y.</collectorName>
|
||
<collectingCounty id="62E464D5FFC93408FDF3FA58FCE6FA69" box="[638,889,1481,1506]" pageId="10" pageNumber="11">Fujita. Shimoji Island</collectingCounty>
|
||
: RUMF-ZE-2017 (
|
||
<specimenCount id="9D3CD7D0FFC93408FBD8FA5BFB48FA69" box="[1109,1239,1481,1506]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), submarine cave (
|
||
<collectorName id="26CF798FFFC93408FF13FA7CFEF1F98D" box="[158,366,1517,1542]" pageId="10" pageNumber="11">Akuma-no-Yakata</collectorName>
|
||
cave) at the
|
||
<collectingMunicipality id="6BE18623FFC93408FD9DFA7CFCB7F98D" box="[528,808,1517,1542]" pageId="10" pageNumber="11">southern Shimoji Island</collectingMunicipality>
|
||
,
|
||
<collectorName id="26CF798FFFC93408FCB5FA7CFBABF98D" box="[824,1076,1517,1542]" pageId="10" pageNumber="11">Miyako-Island Group</collectorName>
|
||
,
|
||
<location id="8EE54A82FFC93408FBCEFA7CFB67F98D" LSID="urn:lsid:plazi:treatment:0393AD4FFFC93413FF1AFC3FFDC9FD7B:8EE54A82FFC93408FBCEFA7CFB67F98D" box="[1091,1272,1517,1542]" country="Japan" county="Fujita. Shimoji Island" municipality="southern Shimoji Island" name="Ryukyu Islands" pageId="10" pageNumber="11">Ryukyu Islands</location>
|
||
, southwestern
|
||
<collectingCountry id="F32D5CC9FFC93408FF1AF983FF48F9A2" box="[151,215,1554,1577]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
|
||
(24˚49’22.51’N, 125˚08’07.84”E), c.a.
|
||
<elevation id="0017FB6AFFC93408FD29F983FD7FF9A2" box="[676,736,1554,1578]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="20.0">
|
||
<quantity id="4CC2B1BCFFC93408FD29F983FD7FF9A2" box="[676,736,1554,1578]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="20.0">20 m</quantity>
|
||
</elevation>
|
||
depth,
|
||
<collectingDate id="EFC0C371FFC93408FCB7F983FC63F9A1" box="[826,1020,1553,1578]" pageId="10" pageNumber="11" value="2017-02-04">
|
||
<date id="FF843A99FFC93408FCB7F983FC63F9A1" box="[826,1020,1553,1578]" pageId="10" pageNumber="11" value="2017-02-04">4 February 2017</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B521604FFC93408FB86F983FA83F9C5" ID-GBIF-Occurrence="1806155410" collectingDate="2017-02-04" collectionCode="SCUBA" collectorName="By Y. & Fujita" country="Japan" county="Fujita. Shimoji Island" elevation="20" location="Ryukyu Islands" municipality="southern Shimoji Island" pageId="10" pageNumber="11" specimenCount="2" typeStatus="paratype">
|
||
<collectionCode id="ED2B849CFFC93408FB86F983FBF7F9A1" box="[1035,1128,1554,1578]" pageId="10" pageNumber="11">SCUBA</collectionCode>
|
||
diving, coll.
|
||
<collectorName id="26CF798FFFC93408FA8AF983FA07F9A1" box="[1287,1432,1553,1578]" pageId="10" pageNumber="11">By Y. Fujita</collectorName>
|
||
, RUMF-ZE-02018 (
|
||
<specimenCount id="9D3CD7D0FFC93408FEFBF9A7FE65F9C5" box="[374,506,1589,1614]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), same locality and sampling data as RUMF-ZE-02017 (
|
||
<specimenCount id="9D3CD7D0FFC93408FB02F9A7FA8BF9C5" box="[1167,1300,1589,1614]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B521604FFC93408FAA7F9A7FB31F9F9" ID-GBIF-Occurrence="1806155417" collectingDate="2017-02-05" collectionCode="NSMT" collectorName="By Y. & Fujita" country="Japan" county="Fujita. Shimoji Island" elevation="8" location="Ryukyu Islands" municipality="southern Shimoji Island" pageId="10" pageNumber="11" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2B849CFFC93408FAA7F9A7FAE7F9C5" box="[1322,1400,1590,1614]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="10" pageNumber="11">NSMT</collectionCode>
|
||
E- 10713 (
|
||
<specimenCount id="9D3CD7D0FFC93408FF79F9CBFEE4F9F9" box="[244,379,1625,1650]" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), same locality as RUMF-ZE-2017 c.a.
|
||
<elevation id="0017FB6AFFC93408FCD4F9CBFC16F9FA" box="[857,905,1626,1650]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="m" value="8.0">
|
||
<quantity id="4CC2B1BCFFC93408FCD4F9CBFC16F9FA" box="[857,905,1626,1650]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="m" value="8.0">8 m</quantity>
|
||
</elevation>
|
||
depth,
|
||
<collectingDate id="EFC0C371FFC93408FC6BF9C8FB31F9F9" box="[998,1198,1625,1650]" pageId="10" pageNumber="11" value="2017-02-05">
|
||
<date id="FF843A99FFC93408FC6BF9C8FB31F9F9" box="[998,1198,1625,1650]" pageId="10" pageNumber="11" value="2017-02-05">5 February 2017</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B521604FFC93408FB33F9CBFDEEF91D" ID-GBIF-Occurrence="1806155415" collectingDate="2017-02-05" collectionCode="NSMT" collectorName="By Y. & Fujita" country="Japan" county="Fujita. Shimoji Island" elevation="8" location="Ryukyu Islands" municipality="southern Shimoji Island" pageId="10" pageNumber="11" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2B849CFFC93408FB33F9CBFA93F9F9" box="[1214,1292,1626,1650]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="10" pageNumber="11">NSMT</collectionCode>
|
||
E-10714 (
|
||
<specimenCount id="9D3CD7D0FFC93408FA1DF9CBFE9AF91D" pageId="10" pageNumber="11" type="generic">1 specimen</specimenCount>
|
||
), same data as RUMF-ZE-2017.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3204FD2FFC93408FF4AF930FC40F88A" pageId="10" pageNumber="11" type="diagnosis">
|
||
<paragraph id="8B851C59FFC93408FF4AF930FC40F88A" blockId="10.[151,1437,1085,2010]" pageId="10" pageNumber="11">
|
||
<emphasis id="B94EC04BFFC93408FF4AF930FEA5F931" bold="true" box="[199,314,1697,1722]" pageId="10" pageNumber="11">Diagnosis</emphasis>
|
||
. Disc scales polygonal, uniform in size; oral shields almost as long as wide, shallow concave proximal sides that end in a fairly sharp point medially; arms about 9 times longer than disc diameter; 3 cylindrical arm spines, of which the oral most one is slightly larger than the other two.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3204FD2FFC9340FFF4AF89CFBCAFAF3" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="8B851C59FFC93409FF4AF89CFF59FE5B" blockId="10.[151,1437,1085,2010]" lastBlockId="11.[151,1437,151,464]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
<emphasis id="B94EC04BFFC93408FF4AF89CFC41F8AD" bold="true" box="[199,990,1805,1830]" pageId="10" pageNumber="11">
|
||
Description of external morphology (
|
||
<typeStatus id="5481A2FBFFC93408FD08F89CFD6AF8AD" box="[645,757,1805,1830]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
|
||
, RUMF-ZE-02008)
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B94EC04BFFC93408FC60F89FFBB7F8AE" box="[1005,1064,1806,1829]" italics="true" pageId="10" pageNumber="11">Disc.</emphasis>
|
||
Pentagonal,
|
||
<quantity id="4CC2B1BCFFC93408FB31F89CFA8BF8AE" box="[1212,1300,1805,1830]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="10" pageNumber="11" unit="mm" value="5.3">5.3 mm</quantity>
|
||
in diameter (
|
||
<figureCitation id="130100DCFFC93408FF12F8A0FF64F8C2" box="[159,251,1841,1866]" captionStart="FIGURE7" captionStartId="11.[151,250,1419,1441]" captionTargetBox="[164,1426,530,1407]" captionTargetId="figure@11.[151,1436,517,1407]" captionTargetPageId="11" captionText="FIGURE7. Ophiozonella cavernalisn. sp. A, living status insitu, aboral view of RUMF-ZE-02008,RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163184/files/figure.png" pageId="10" pageNumber="11">Figs 7B</figureCitation>
|
||
; 8A). Aboral surface with imbricating polygonal and flat scales, approximately 200–400 µm in length at periphery (
|
||
<figureCitation id="130100DCFFC93408FE9CF8C4FEF8F8E5" box="[273,359,1877,1902]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="10" pageNumber="11">Fig. 8B</figureCitation>
|
||
) and approximately 300–500 µm in length at center (
|
||
<figureCitation id="130100DCFFC93408FC31F8C4FB8BF8E5" box="[956,1044,1877,1902]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="10" pageNumber="11">Fig. 8C</figureCitation>
|
||
). Radial shields broadly triangular, approximately 800 µm in width and 500 µm in length, separated by 2 or 3 scales (
|
||
<figureCitation id="130100DCFFC93408FBACF8E8FBE7F819" box="[1057,1144,1913,1938]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="10" pageNumber="11">Fig. 8B</figureCitation>
|
||
). Adoral shield triangular, wider than long, approximately 550 µm in width, 200 µm in length inner side, 300 µm in length outer side, continues around the lateral angle of the oral shield, in contact with first ventral arm plates (
|
||
<figureCitation id="130100DCFFC93408FB1BF850FA8FF851" box="[1174,1296,1985,2010]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="10" pageNumber="11">Fig. 8D–E</figureCitation>
|
||
). Oral plates small, obscured by thin skin (
|
||
<figureCitation id="130100DCFFC83409FE53FF06FDAAFF3B" box="[478,565,151,176]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="11" pageNumber="12">Fig. 8E</figureCitation>
|
||
). Oral shields pointed proximally with slightly concave lateral proximal edges and a strongly convex distal edge, approximately 500 µm in length, 600 µm in width (
|
||
<figureCitation id="130100DCFFC83409FBF0FF2DFB49FF5F" box="[1149,1238,188,213]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="11" pageNumber="12">Fig. 8E</figureCitation>
|
||
). All oral shields similar in size and shape, and oral shield serving as madreporite unrecognizable in external view (
|
||
<figureCitation id="130100DCFFC83409FA9AFF4EFA0FFF73" box="[1303,1424,223,248]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="11" pageNumber="12">Fig. 8E–F</figureCitation>
|
||
). Interradial oral disc also covered by polygonal and imbricating scales, approximately 200–350 µm in length, approximately
|
||
<quantity id="4CC2B1BCFFC83409FECCFEB9FEE4FECB" box="[321,379,295,320]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="11" pageNumber="12" unit="in" value="30.0">30 in</quantity>
|
||
number (
|
||
<figureCitation id="130100DCFFC83409FE64FEB6FDDFFECB" box="[489,576,295,320]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="11" pageNumber="12">Fig. 8F</figureCitation>
|
||
). A scale on midline of lateral edge larger, approximately 600 µm in length (
|
||
<figureCitation id="130100DCFFC83409FF12FEDDFF6BFEEF" box="[159,244,332,357]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="11" pageNumber="12">Fig. 8F</figureCitation>
|
||
). Genital slits long, narrow, almost extending to the disc edge, approximately
|
||
<quantity id="4CC2B1BCFFC83409FBD1FEDDFB2DFEEF" box="[1116,1202,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="mm" value="0.1">0.1 mm</quantity>
|
||
wide (
|
||
<figureCitation id="130100DCFFC83409FB72FEDDFACCFEEF" box="[1279,1363,332,357]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="11" pageNumber="12">Fig. 8F</figureCitation>
|
||
). Four oral papillae at edge of each oral plate, 3 triangular proximal papillae, distal most wider and oblong (
|
||
<figureCitation id="130100DCFFC83409FB78FEFEFAD4FE03" box="[1269,1355,367,392]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="11" pageNumber="12">Fig. 8E</figureCitation>
|
||
). Teeth triangular, pointed, forming vertical row on dental plate. Second tentacle pore completely inside mouth slit (
|
||
<figureCitation id="130100DCFFC83409FAE2FE05FF25FE5B" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="11" pageNumber="12">Fig. 8E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF454CD1FFC83409FF1AFA1AFF4DFA77" httpUri="https://zenodo.org/record/1163184/files/figure.png" pageId="11" pageNumber="12" targetBox="[164,1426,530,1407]" targetPageId="11">
|
||
<paragraph id="8B851C59FFC83409FF1AFA1AFF4DFA77" blockId="11.[151,1436,1419,1532]" pageId="11" pageNumber="12">
|
||
<emphasis id="B94EC04BFFC83409FF1AFA1AFE8DFA2A" bold="true" box="[151,274,1419,1441]" pageId="11" pageNumber="12">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C3A67DAFFC83409FE95FA1AFD8FFA2A" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[280,528,1419,1441]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="11" pageNumber="12" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFC83409FE95FA1AFD8FFA2A" box="[280,528,1419,1441]" italics="true" pageId="11" pageNumber="12">Ophiozonella cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFC83409FD98FA1AFDD1FA2A" box="[533,590,1419,1441]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="B94EC04BFFC83409FD98FA1AFDD1FA2A" bold="true" box="[533,590,1419,1441]" pageId="11" pageNumber="12">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A, living status
|
||
<emphasis id="B94EC04BFFC83409FD7EFA1DFCA8FA2A" box="[755,823,1420,1441]" italics="true" pageId="11" pageNumber="12">in situ,</emphasis>
|
||
aboral view of RUMF-ZE-02008, RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B851C59FFC83409FF4AF9DFFE58F925" blockId="11.[151,1437,1613,2034]" pageId="11" pageNumber="12">
|
||
<emphasis id="B94EC04BFFC83409FF4AF9DFFE95F9EE" box="[199,266,1614,1637]" italics="true" pageId="11" pageNumber="12">Arms.</emphasis>
|
||
Five in number, 3 with evidence of regeneration,
|
||
<quantity id="4CC2B1BCFFC83409FCCEF9DFFC22F9EE" box="[835,957,1614,1638]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.7" metricValueMin="3.1" pageId="11" pageNumber="12" unit="mm" value="34.0" valueMax="37.0" valueMin="31.0">31–37 mm</quantity>
|
||
in length, and 2 entire arms
|
||
<quantity id="4CC2B1BCFFC83409FA88F9DCFAE0F9EE" box="[1285,1407,1613,1638]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="11" pageNumber="12" unit="mm" value="43.5" valueMax="46.0" valueMin="41.0">41–46 mm</quantity>
|
||
in length,
|
||
<quantity id="4CC2B1BCFFC83409FF63F9E0FED8F902" box="[238,327,1649,1674]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="mm" value="1.6">1.6 mm</quantity>
|
||
wide and
|
||
<quantity id="4CC2B1BCFFC83409FE4EF9E3FD83F902" box="[451,540,1650,1674]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="12" unit="mm" value="1.2">1.2 mm</quantity>
|
||
high, arched on dorsal side and flat on ventral side. Arms tapering gradually toward the arm tip (
|
||
<figureCitation id="130100DCFFC83409FEF9F904FE26F925" box="[372,441,1685,1710]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFC8340EFF4AF928FEC6FEEF" blockId="11.[151,1437,1613,2034]" lastBlockId="12.[151,1437,151,357]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
Each arm segment externally composed of a dorsal arm plate, 2 lateral arm plates and a ventral arm plate (
|
||
<figureCitation id="130100DCFFC83409FAE2F928FE98F97D" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9A, D, G</figureCitation>
|
||
). Dorsal arm plate fan-shaped on proximal portion of arm, rounded on distal edge (
|
||
<figureCitation id="130100DCFFC83409FB5BF94CFAADF97D" box="[1238,1330,1757,1782]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9A</figureCitation>
|
||
), sharply triangular on middle portion of arm, slightly separated by lateral arm plates (
|
||
<figureCitation id="130100DCFFC83409FC60F890FBDBF891" box="[1005,1092,1793,1818]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9B</figureCitation>
|
||
), decreasing in size toward the arm tip and finally disappearing (
|
||
<figureCitation id="130100DCFFC83409FD9DF8B4FDF5F8B5" box="[528,618,1829,1854]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9C</figureCitation>
|
||
). Lateral arm plates oblong in external view, slightly wider than high on proximal portion of arm and twice as wide as high on distal portion of arm (
|
||
<figureCitation id="130100DCFFC83409FC7AF8D8FBEAF8E9" box="[1015,1141,1865,1890]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9G–H</figureCitation>
|
||
). Ventral arm plate square with rounded distal edge, contiguous, almost as long as wide on proximal portion of arm (
|
||
<figureCitation id="130100DCFFC83409FB38F8FCFA8DF80D" box="[1205,1298,1901,1926]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9D</figureCitation>
|
||
), triangular, rounded on distal edge, longer than wide (
|
||
<figureCitation id="130100DCFFC83409FDF8F800FD52F821" box="[629,717,1937,1962]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9E</figureCitation>
|
||
), decreasing in size toward the arm tip and finally disappearing (
|
||
<figureCitation id="130100DCFFC83409FF12F824FF6AF845" box="[159,245,1973,1998]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9F</figureCitation>
|
||
). Tentacle pores along entire arm (
|
||
<figureCitation id="130100DCFFC83409FDF3F824FD67F845" box="[638,760,1973,1998]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9D–F</figureCitation>
|
||
). Two oval, flat tentacle scales at each tentacle pore, almost the same size on proximal portion of the arms (
|
||
<figureCitation id="130100DCFFC83409FD35F848FC8BF879" box="[696,788,2009,2034]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="11" pageNumber="12">Fig. 9D</figureCitation>
|
||
). On middle portion of arm, tentacle scales oval and flat decreasing in number to 1 scale at each tentacle pore (
|
||
<figureCitation id="130100DCFFCF340EFC8DFF06FCC7FF3B" box="[768,856,151,176]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="12" pageNumber="13">Fig. 9E</figureCitation>
|
||
). Tentacle scales decreasing in size toward the arm tip and finally disappearing (
|
||
<figureCitation id="130100DCFFCF340EFE50FF2DFDACFF5F" box="[477,563,188,213]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="12" pageNumber="13">Fig. 9F</figureCitation>
|
||
). Three conical arm spines, almost the same length as the corresponding arm segments on proximal portions of arms (
|
||
<figureCitation id="130100DCFFCF340EFDE8FF4EFD5EFF73" box="[613,705,223,248]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="12" pageNumber="13">Fig. 9G</figureCitation>
|
||
). On middle portion of arms, arm spines decreasing in length to one-fourth to one-fifth the length of corresponding arm segments, oral most spine slightly longer than other 2 spines (
|
||
<figureCitation id="130100DCFFCF340EFF60FEB6FED7FECB" box="[237,328,295,320]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="12" pageNumber="13">Fig. 9H</figureCitation>
|
||
). On distal portion of arm, arm spines become ovoid, pointed, decreasing in number to 1 on distal arm tip (
|
||
<figureCitation id="130100DCFFCF340EFF7BFEDDFED4FEEF" box="[246,331,332,357]" captionStart="FIGURE 9" captionStartId="14.[151,250,1913,1935]" captionTargetBox="[247,1358,193,1894]" captionTargetId="figure@14.[228,1375,193,1894]" captionTargetPageId="14" captionText="FIGURE 9. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008).A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate." httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="12" pageNumber="13">Fig. 9F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF454CD1FFCF340EFF1AF8EDFB45F85B" httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="12" pageNumber="13" targetBox="[152,1435,409,1895]" targetPageId="12">
|
||
<paragraph id="8B851C59FFCF340EFF1AF8EDFB45F85B" blockId="12.[151,1437,1916,2000]" pageId="12" pageNumber="13">
|
||
<emphasis id="B94EC04BFFCF340EFF1AF8EDFE87F819" bold="true" box="[151,280,1916,1938]" pageId="12" pageNumber="13">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C3A67DAFFCF340EFEA9F8EDFDBEF819" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[292,545,1916,1938]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFCF340EFEA9F8EDFDBEF819" box="[292,545,1916,1938]" italics="true" pageId="12" pageNumber="13">Ophiozonella cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFCF340EFDA3F8EDFDF3F819" box="[558,620,1916,1938]" pageId="12" pageNumber="13" rank="species">
|
||
<emphasis id="B94EC04BFFCF340EFDA3F8EDFDF3F819" bold="true" box="[558,620,1916,1938]" pageId="12" pageNumber="13">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B851C59FFCE340FFF4AFF06FDE6FE97" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
<emphasis id="B94EC04BFFCE340FFF4AFF06FB95FF3B" bold="true" box="[199,1034,151,176]" pageId="13" pageNumber="14">
|
||
Description by SEM observation (A
|
||
<typeStatus id="5481A2FBFFCE340FFD28FF06FC8DFF3B" box="[677,786,151,176]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
, RUMF-ZE-02009).
|
||
</emphasis>
|
||
Aboral surface with imbricating polygonal and flat scales, approximately 250–500 µm in length at periphery (
|
||
<figureCitation id="130100DCFFCE340FFC76FF2DFBCDFF5F" box="[1019,1106,188,213]" captionStart="FIGURE7" captionStartId="11.[151,250,1419,1441]" captionTargetBox="[164,1426,530,1407]" captionTargetId="figure@11.[151,1436,517,1407]" captionTargetPageId="11" captionText="FIGURE7. Ophiozonella cavernalisn. sp. A, living status insitu, aboral view of RUMF-ZE-02008,RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163184/files/figure.png" pageId="13" pageNumber="14">Fig. 7C</figureCitation>
|
||
) and approximately 200–450 µm in length at center (
|
||
<figureCitation id="130100DCFFCE340FFE12FF4EFE66FF73" box="[415,505,223,248]" captionStart="FIGURE7" captionStartId="11.[151,250,1419,1441]" captionTargetBox="[164,1426,530,1407]" captionTargetId="figure@11.[151,1436,517,1407]" captionTargetPageId="11" captionText="FIGURE7. Ophiozonella cavernalisn. sp. A, living status insitu, aboral view of RUMF-ZE-02008,RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163184/files/figure.png" pageId="13" pageNumber="14">Fig. 7D</figureCitation>
|
||
). Radial shields broadly triangular, approximately 500 µm in width and 700 µm in length, separated by 2 or 3 scales (
|
||
<figureCitation id="130100DCFFCE340FFD94FE95FDEEFE97" box="[537,625,260,285]" captionStart="FIGURE7" captionStartId="11.[151,250,1419,1441]" captionTargetBox="[164,1426,530,1407]" captionTargetId="figure@11.[151,1436,517,1407]" captionTargetPageId="11" captionText="FIGURE7. Ophiozonella cavernalisn. sp. A, living status insitu, aboral view of RUMF-ZE-02008,RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163184/files/figure.png" pageId="13" pageNumber="14">Fig. 7D</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFCE340FFF4AFEB6FD9EFD47" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
<emphasis id="B94EC04BFFCE340FFF4AFEB6FC79FECB" bold="true" box="[199,998,295,320]" pageId="13" pageNumber="14">
|
||
Description of ossicle morphology (A
|
||
<typeStatus id="5481A2FBFFCE340FFD07FEB6FD68FECB" box="[650,759,295,320]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
, RUMF-ZE-02009).
|
||
</emphasis>
|
||
Dorsal arm plates triangular, nearly as long as wide throughout arm (
|
||
<figureCitation id="130100DCFFCE340FFE60FEDDFDE7FEEF" box="[493,632,332,357]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10A, C</figureCitation>
|
||
). A small knob present on proximal tip of dorsal arm plate at proximal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFEC2FEFEFE25FE03" box="[335,442,367,392]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10A</figureCitation>
|
||
). Lateral arm plates on proximal portion of arm slightly longer than high, aboral edge curved to oral-distal side and oral edge shallow convex on proximal side (
|
||
<figureCitation id="130100DCFFCE340FFC43FE05FBC9FE27" box="[974,1110,404,429]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10D–E</figureCitation>
|
||
); distal portion of arm, plates approximately 4 times longer than high, oral and aboral edges both straight (
|
||
<figureCitation id="130100DCFFCE340FFBBBFE26FB22FE5B" box="[1078,1213,439,464]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10H–I</figureCitation>
|
||
). In external view, proximal edge convex with 2 well defined, prominent, horizontally elongated spurs composed of more densely meshed stereom than remaining proximal edge (
|
||
<figureCitation id="130100DCFFCE340FFD39FE6EFC82FD93" box="[692,797,511,536]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10D</figureCitation>
|
||
). External oral spur slightly larger than aboral spur. Such spurs disappear on distal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFDE8FDB5FD79FDB7" box="[613,742,548,573]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10H–I</figureCitation>
|
||
). Bands of conspicuous, fine, horizontal striation on proximal edge (
|
||
<figureCitation id="130100DCFFCE340FFF56FDD6FEDDFDEB" box="[219,322,583,608]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10D</figureCitation>
|
||
), such bands disappearing on distal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFC0BFDD6FC71FDEB" box="[902,1006,583,608]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10H</figureCitation>
|
||
). Three equal-sized spine articulations on distal edge (
|
||
<figureCitation id="130100DCFFCE340FFECEFDFDFD9DFD0F" box="[323,514,620,645]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10D, F, H, J</figureCitation>
|
||
), composed of parallel, horizontal dorsal and ventral lobes (
|
||
<figureCitation id="130100DCFFCE340FFB2BFDFDFA90FD0F" box="[1190,1295,620,645]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10G</figureCitation>
|
||
). Both lobes almost the same length on proximal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFD52FD1EFCDBFD23" box="[735,836,655,680]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10G</figureCitation>
|
||
) and ventral lobes slightly larger than dorsal lobes on distal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFE01FD25FE6BFD47" box="[396,500,692,717]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFCE340FFF4AFD49FF55FC67" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
On inner side, 2 large, well defined, horizontally elongated elevated structures on proximal edges (
|
||
<figureCitation id="130100DCFFCE340FFA99FD46FA12FD7B" box="[1300,1421,727,752]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10E, I</figureCitation>
|
||
). Elevated structure almost the same width throughout arm (
|
||
<figureCitation id="130100DCFFCE340FFCBCFD6DFC31FC9F" box="[817,942,764,789]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10E, I</figureCitation>
|
||
). Distal edge concave, with 2 well defined, oval, horizontally elongate spurs (
|
||
<figureCitation id="130100DCFFCE340FFD95FC8EFDE0FCB3" box="[536,639,799,824]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10E</figureCitation>
|
||
). Oral spur slightly wider than aboral one, on proximal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFF12FCD5FE9AFCD7" box="[159,261,836,861]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10E</figureCitation>
|
||
). Spurs absent on distal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFD48FCD5FCBEFCD7" box="[709,801,836,861]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">
|
||
Fig.
|
||
<date id="FF843A99FFCE340FFD74FCD5FCBEFCD7" box="[761,801,836,860]" pageId="13" pageNumber="14" value="2010-01">10I</date>
|
||
</figureCitation>
|
||
). Spurs composed of more densely meshed stereom than remaining distal edge. Tentacle notches opening on oral side, ending in center of lateral arm plates (
|
||
<figureCitation id="130100DCFFCE340FFB66FCF6FAFAFC0B" box="[1259,1381,871,896]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10E, I</figureCitation>
|
||
). On proximal portion of arm, a single perforation (
|
||
<figureCitation id="130100DCFFCE340FFD21FC1DFC8EFC2F" box="[684,785,908,933]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10E</figureCitation>
|
||
) absent on distal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFB1BFC1DFA9EFC2F" box="[1174,1281,908,933]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">Fig. 10H</figureCitation>
|
||
). Ventral arm plates longer than wide, sharply triangular, notched laterally (
|
||
<figureCitation id="130100DCFFCE340FFCC1FC3EFC47FC43" box="[844,984,943,968]" captionStart="FIGURE 10" captionStartId="15.[151,250,1794,1816]" captionTargetBox="[234,1343,193,1767]" captionTargetId="figure@15.[234,1352,193,1768]" captionTargetPageId="15" captionText="FIGURE 10. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe." httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="13" pageNumber="14">
|
||
Fig.
|
||
<specimenCount id="9D3CD7D0FFCE340FFC0CFC21FC2AFC43" box="[897,949,944,968]" pageId="13" pageNumber="14" type="generic">10L.</specimenCount>
|
||
M
|
||
</figureCitation>
|
||
), notches deeper on proximal portion of arm.
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFCE340FFF4AFC66FB2DFB4F" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
Vertebrae with zygospondylous articulation, elongate, large wing-like muscle flanges distally and proximally (
|
||
<figureCitation id="130100DCFFCE340FFF12FB8DFEB6FBBF" box="[159,297,1052,1077]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11A–B</figureCitation>
|
||
). Muscle flanges of distal portion of arm narrower and longer than those of proximal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFF12FBAEFEB4FBD3" box="[159,299,1087,1112]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11A–D</figureCitation>
|
||
). Longitudinal straight groove on both aboral and oral side (
|
||
<figureCitation id="130100DCFFCE340FFC52FBAEFBF6FBD3" box="[991,1129,1087,1112]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11C–D</figureCitation>
|
||
). A longitudinal large hole opening on median line on distal portion of the arm (
|
||
<figureCitation id="130100DCFFCE340FFD6CFBF5FCD7FBF7" box="[737,840,1124,1149]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11H</figureCitation>
|
||
). No distinct radial water/nerve canals observed from oral view (
|
||
<figureCitation id="130100DCFFCE340FFE9BFB16FE39FB2B" box="[278,422,1159,1184]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11D, H</figureCitation>
|
||
). Depression for tentacles on lateral distal side of vertebra (
|
||
<figureCitation id="130100DCFFCE340FFBEEFB16FB6CFB2B" box="[1123,1267,1159,1184]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11D, H</figureCitation>
|
||
). On proximal portion of the arm, thin flange covering aboral side of depression partly perforated (
|
||
<figureCitation id="130100DCFFCE340FFBCCFB3DFB3AFB4F" box="[1089,1189,1196,1221]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFCE340FFF4AFB41FB04FABB" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
Arm spines cylindrical, with blunt tip on proximal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFC22FB5EFB96FB63" box="[943,1033,1231,1256]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">
|
||
Fig.
|
||
<date id="FF843A99FFCE340FFC6FFB41FB96FB63" box="[994,1033,1232,1256]" pageId="13" pageNumber="14" value="2011-01">11I</date>
|
||
</figureCitation>
|
||
), narrow, with slightly thorny tip on middle portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFE13FB65FE64FA87" box="[414,507,1268,1293]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11J</figureCitation>
|
||
), hook-shaped, with several, 3 or 4, secondary teeth on inner side on distal portion of arm (
|
||
<figureCitation id="130100DCFFCE340FFF7DFA86FEC8FABB" box="[240,343,1303,1328]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">Fig. 11K</figureCitation>
|
||
). Spines with simple and fine stereom without minute thorns (
|
||
<figureCitation id="130100DCFFCE340FFB82FA86FB10FABB" box="[1039,1167,1303,1328]" captionStart="FIGURE 11" captionStartId="16.[151,250,1833,1855]" captionTargetBox="[221,1356,193,1805]" captionTargetId="figure@16.[221,1356,193,1818]" captionTargetPageId="16" captionText="FIGURE 11. Ophiozonella cavernalis n. sp. a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximalportion of arms, proximal view (A), distal view (B), aboral view (C)and oral view (D), a square enlargedfrom D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximalportion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations:ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations:AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth." httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="13" pageNumber="14">
|
||
Fig.
|
||
<date id="FF843A99FFCE340FFBC9FA89FBF7FABB" box="[1092,1128,1304,1328]" pageId="13" pageNumber="14" value="2011-01">11I</date>
|
||
–K
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFCE340FFF4AFAADFBCAFAF3" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
<emphasis id="B94EC04BFFCE340FFF4AFAADFEC7FADE" bold="true" box="[199,344,1340,1365]" pageId="13" pageNumber="14">Color in life</emphasis>
|
||
. Mottled light and dark brown on aboral disc, radial shields darker, arms variegated light brown with dark bands, a darker brown band on proximal portion of each arm spine (
|
||
<figureCitation id="130100DCFFCE340FFB8FFACEFBD8FAF3" box="[1026,1095,1375,1400]" captionStart="FIGURE7" captionStartId="11.[151,250,1419,1441]" captionTargetBox="[164,1426,530,1407]" captionTargetId="figure@11.[151,1436,517,1407]" captionTargetPageId="11" captionText="FIGURE7. Ophiozonella cavernalisn. sp. A, living status insitu, aboral view of RUMF-ZE-02008,RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163184/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3204FD2FFCE340FFF4AFA15FC91FA4B" pageId="13" pageNumber="14" type="etymology">
|
||
<paragraph id="8B851C59FFCE340FFF4AFA15FC91FA4B" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
<emphasis id="B94EC04BFFCE340FFF4AFA15FED9FA16" bold="true" box="[199,326,1412,1437]" pageId="13" pageNumber="14">Etymology</emphasis>
|
||
. The specific name is derived from the Latin feminine noun,
|
||
<taxonomicName id="4C3A67DAFFCE340FFB8EFA15FBE7FA17" authorityName="Okanishi & Fujita" authorityYear="2018" box="[1027,1144,1412,1436]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="cavernalis">
|
||
<emphasis id="B94EC04BFFCE340FFB8EFA15FBE7FA17" box="[1027,1144,1412,1436]" italics="true" pageId="13" pageNumber="14">cavernalis</emphasis>
|
||
</taxonomicName>
|
||
meaning “inhabits cave”, referring to the cavernicolous habits of this new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3204FD2FFCE340FFF4AFA5DFAA0F8A3" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="8B851C59FFCE340FFF4AFA5DFC78F983" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
<emphasis id="B94EC04BFFCE340FFF4AFA5DFEC6FA6E" bold="true" box="[199,345,1484,1509]" pageId="13" pageNumber="14">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F32D5CC9FFCE340FFEE1FA5DFE5AFA6F" box="[364,453,1484,1508]" name="Japan" pageId="13" pageNumber="14">JAPAN</collectingCountry>
|
||
: submarine caves on the Ryukyu Islands (
|
||
<collectingRegion id="49FED2BBFFCE340FFC36FA5DFBBDFA6E" box="[955,1058,1484,1509]" country="Japan" name="Okinawa" pageId="13" pageNumber="14">Okinawa</collectingRegion>
|
||
and Ie Islands,
|
||
<collectingRegion id="49FED2BBFFCE340FFB6AFA5DFAD3FA6E" box="[1255,1356,1484,1509]" country="Japan" name="Okinawa" pageId="13" pageNumber="14">Okinawa</collectingRegion>
|
||
Island Group, and Shimoji Island, Miyako Island Group),
|
||
<quantity id="4CC2B1BCFFCE340FFD5BFA61FCB4F983" box="[726,811,1520,1544]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.7" metricValueMin="0.8" pageId="13" pageNumber="14" unit="m" value="17.5" valueMax="27.0" valueMin="8.0">8–27 m</quantity>
|
||
depth (
|
||
<figureCitation id="130100DCFFCE340FFC0DFA7EFC44F983" box="[896,987,1519,1544]" captionStart="FIGURE7" captionStartId="11.[151,250,1419,1441]" captionTargetBox="[164,1426,530,1407]" captionTargetId="figure@11.[151,1436,517,1407]" captionTargetPageId="11" captionText="FIGURE7. Ophiozonella cavernalisn. sp. A, living status insitu, aboral view of RUMF-ZE-02008,RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163184/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFCE340FFF4AF985FAA0F8A3" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
<emphasis id="B94EC04BFFCE340FFF4AF985FEBCF9A6" bold="true" box="[199,291,1556,1581]" pageId="13" pageNumber="14">Habitat</emphasis>
|
||
. In the Ryukyu Islands, this new species was found in 4 submarine caves “Hedo-Dome cave” at
|
||
<collectingRegion id="49FED2BBFFCE340FFF1AF9A6FF63F9DB" box="[151,252,1591,1616]" country="Japan" name="Okinawa" pageId="13" pageNumber="14">Okinawa</collectingRegion>
|
||
Island, “Dai-Dokutsu cave” and “unnamed cave” at Ie
|
||
<collectingCountry id="F32D5CC9FFCE340FFCF6F9A6FC20F9DB" box="[891,959,1591,1616]" name="Iceland" pageId="13" pageNumber="14">Island</collectingCountry>
|
||
and “Akuma-no-Yakata cave” at Shimoji Island (
|
||
<figureCitation id="130100DCFFCE340FFF61F9CDFEB0F9FF" box="[236,303,1628,1653]" captionStart="FIGURE 1" captionStartId="1.[151,250,1946,1968]" captionTargetBox="[207,1389,627,1885]" captionTargetId="figure@1.[197,1390,626,1921]" captionTargetPageId="1" captionText="FIGURE 1. Sampling localities of Ophiolepis cavitata n. sp. and Ophiozonella cavernalis n. sp. Red symbols indicate type localities." httpUri="https://zenodo.org/record/1163172/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
|
||
), and anchialine environment (low salinity and water-temperature) is confirmed in all caves. At the “Hedo-Dome cave”, this species was collected together with
|
||
<taxonomicName id="4C3A67DAFFCE340FFCD6F911FBA5F91C" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[859,1082,1664,1688]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="cavitata" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFCE340FFCD6F911FBA5F91C" box="[859,1082,1664,1688]" italics="true" pageId="13" pageNumber="14">Ophiolepis cavitata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFCE340FFBC9F9EEFB19F913" box="[1092,1158,1663,1688]" pageId="13" pageNumber="14" rank="species">
|
||
<emphasis id="B94EC04BFFCE340FFBC9F9EEFB19F913" bold="true" box="[1092,1158,1663,1688]" pageId="13" pageNumber="14">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="130100DCFFCE340FFB15F9EEFB6AF913" box="[1176,1269,1663,1688]" captionStart="FIGURE7" captionStartId="11.[151,250,1419,1441]" captionTargetBox="[164,1426,530,1407]" captionTargetId="figure@11.[151,1436,517,1407]" captionTargetPageId="11" captionText="FIGURE7. Ophiozonella cavernalisn. sp. A, living status insitu, aboral view of RUMF-ZE-02008,RUMF-ZE-02009 or HD- 2017-15; B, anesthetized, aboral view of the holotype (RUMF-ZE-02008); C, D, SEM photos of a paratype (RUMF-ZE- 02009), aboral disc (C) and oral disc (D). Abbreviations: ASH, adoral shield; OP, oral plate; OS, oral shield; P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163184/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
|
||
). At the “Dai- Dohkutsu cave”, this species is densely aggregated on the sandy-muddy bottom of the cave. This species was also found in similar environments (under rock on rubble zone or sandy-muddy bottom) in the “unnamed cave” and “Akuma-no-Yakata cave”. Compared to
|
||
<taxonomicName id="4C3A67DAFFCE340FFDEDF97DFCA3F88F" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[608,828,1772,1796]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="cavitata" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFCE340FFDEDF97DFCA3F88F" box="[608,828,1772,1796]" italics="true" pageId="13" pageNumber="14">Ophiolepis cavitata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFCE340FFCC9F97DFC1CF88E" box="[836,899,1772,1797]" pageId="13" pageNumber="14" rank="species">
|
||
<emphasis id="B94EC04BFFCE340FFCC9F97DFC1CF88E" bold="true" box="[836,899,1772,1797]" pageId="13" pageNumber="14">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the number of
|
||
<taxonomicName id="4C3A67DAFFCE340FFBBAF97DFB58F88F" authorityName="Okanishi & Fujita" authorityYear="2018" box="[1079,1223,1772,1796]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="cavernalis">
|
||
<emphasis id="B94EC04BFFCE340FFBBAF97DFBD4F888" box="[1079,1099,1772,1795]" italics="true" pageId="13" pageNumber="14">O</emphasis>
|
||
.
|
||
<emphasis id="B94EC04BFFCE340FFBDFF97DFB58F88F" box="[1106,1223,1772,1796]" italics="true" pageId="13" pageNumber="14">cavernalis</emphasis>
|
||
</taxonomicName>
|
||
specimens is large and its presence in 4 caves suggest that it is indeed a cave endemic species with self-recruiting population.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3204FD2FFCE3413FF4AF8A5FC0FFF73" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14" type="discussion">
|
||
<paragraph id="8B851C59FFCE340FFF4AF8A5FE8EF832" blockId="13.[151,1437,151,1977]" pageId="13" pageNumber="14">
|
||
<emphasis id="B94EC04BFFCE340FFF4AF8A5FEABF8C6" bold="true" box="[199,308,1844,1869]" pageId="13" pageNumber="14">Remarks</emphasis>
|
||
. Among 31 species of
|
||
<taxonomicName id="4C3A67DAFFCE340FFDC1F8A5FD7CF8C7" authorityName="Okanishi & Fujita" authorityYear="2018" box="[588,739,1844,1868]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B94EC04BFFCE340FFDC1F8A5FD7CF8C7" box="[588,739,1844,1868]" italics="true" pageId="13" pageNumber="14">Ophiozonella</emphasis>
|
||
</taxonomicName>
|
||
s. a., the present specimens are most closely related to
|
||
<taxonomicName id="4C3A67DAFFCE340FFA0CF8A5FE02F8FB" authority="(Lyman 1883)" authorityName="Lyman" authorityYear="1883" baseAuthorityName="Lyman" baseAuthorityYear="1883" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="clypeata">
|
||
<emphasis id="B94EC04BFFCE340FFA0CF8A5FF69F8FB" italics="true" pageId="13" pageNumber="14">O. clypeata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAB61A8FFCE340FFE88F8C9FE0AF8FB" author="Lyman" box="[261,405,1880,1904]" pageId="13" pageNumber="14" refString="Lyman, T. (1883) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Caribbean Sea (1878 - 79), and along the Atlantic coast of the United States during the summer of 1880, by the U. S. coast survey steamer " Blake, " commander J. R. Bartlett, U. S. N., commanding. XX. Report on the Ophiuroidea. Bulletin of the Museum of Comparative Zoology, 10 (6), 227 - 287." type="journal article" year="1883">Lyman 1883</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and partly agree with the original description in morphology (
|
||
<bibRefCitation id="EFAB61A8FFCE340FFBD3F8C9FB6AF8FB" author="Lyman" box="[1118,1269,1880,1904]" pageId="13" pageNumber="14" refString="Lyman, T. (1883) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Caribbean Sea (1878 - 79), and along the Atlantic coast of the United States during the summer of 1880, by the U. S. coast survey steamer " Blake, " commander J. R. Bartlett, U. S. N., commanding. XX. Report on the Ophiuroidea. Bulletin of the Museum of Comparative Zoology, 10 (6), 227 - 287." type="journal article" year="1883">Lyman, 1883</bibRefCitation>
|
||
). However,
|
||
<taxonomicName id="4C3A67DAFFCE340FFA0CF8C9FE93F81F" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFCE340FFA0CF8C9FE93F81F" italics="true" pageId="13" pageNumber="14">O. cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFCE340FFE9EF8EDFECCF81E" box="[275,339,1916,1941]" pageId="13" pageNumber="14" rank="species">
|
||
<emphasis id="B94EC04BFFCE340FFE9EF8EDFECCF81E" bold="true" box="[275,339,1916,1941]" pageId="13" pageNumber="14">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="4C3A67DAFFCE340FFD0BF8EDFC97F81F" baseAuthorityName="Lyman" baseAuthorityYear="1883" box="[646,776,1916,1940]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="clypeata">
|
||
<emphasis id="B94EC04BFFCE340FFD0BF8EDFC97F81F" box="[646,776,1916,1940]" italics="true" pageId="13" pageNumber="14">O. clypeata</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the shape of oral shields and length of arm as follows:
|
||
</paragraph>
|
||
<caption id="DF454CD1FFCD340CFF1AF8E8FC01F860" httpUri="https://zenodo.org/record/1163188/files/figure.png" pageId="14" pageNumber="15" targetBox="[247,1358,193,1894]" targetPageId="14">
|
||
<paragraph id="8B851C59FFCD340CFF1AF8E8FC01F860" blockId="14.[151,1436,1913,2027]" pageId="14" pageNumber="15">
|
||
<emphasis id="B94EC04BFFCD340CFF1AF8E8FE8CF804" bold="true" box="[151,275,1913,1935]" pageId="14" pageNumber="15">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C3A67DAFFCD340CFE94F8E8FD8DF804" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[281,530,1913,1935]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFCD340CFE94F8E8FD8DF804" box="[281,530,1913,1935]" italics="true" pageId="14" pageNumber="15">Ophiozonella cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFCD340CFD95F8E8FDCEF804" box="[536,593,1913,1935]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B94EC04BFFCD340CFD95F8E8FDCEF804" bold="true" box="[536,593,1913,1935]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
holotype (RUMF-ZE-02008). A, aboral proximal portion of arm; B, aboral middle portion of arm; C, aboral distal tip of arm; D, oral proximal portion of arm; E, oral middle portion of arm; F, oral distal tip of arm; G, lateral middle portion of arm; H, lateral distal tip of arm. Arrow heads indicate tentacle scale. Abbreviation: ASP, arm spine; DAP, dorsal arm plate; LAP, lateral arm plate; VAP, ventral arm plate.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF454CD1FFCC340DFF1AF893FC7AF865" httpUri="https://zenodo.org/record/1163190/files/figure.png" pageId="15" pageNumber="16" targetBox="[234,1343,193,1767]" targetPageId="15">
|
||
<paragraph id="8B851C59FFCC340DFF1AF893FC7AF865" blockId="15.[151,1436,1794,2030]" pageId="15" pageNumber="16">
|
||
<emphasis id="B94EC04BFFCC340DFF1AF893FEBFF893" bold="true" box="[151,288,1794,1816]" pageId="15" pageNumber="16">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C3A67DAFFCC340DFEABF893FD81F893" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[294,542,1794,1816]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFCC340DFEABF893FD81F893" box="[294,542,1794,1816]" italics="true" pageId="15" pageNumber="16">Ophiozonella cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFCC340DFDA8F893FDC0F893" box="[549,607,1794,1816]" pageId="15" pageNumber="16" rank="species">
|
||
<emphasis id="B94EC04BFFCC340DFDA8F893FDC0F893" bold="true" box="[549,607,1794,1816]" pageId="15" pageNumber="16">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
a paratype (RUMF-ZE-02009). SEM images of ossicles. A–C, dorsal arm plates, from proximal portion of arm, a square enlarged in B (A), and distal tip, an arrow head indicates a distal knob (B), and from distal portion of arm (C); D–G, lateral arm plates, from proximal portion of arms, outer view (D), inner view (E), distal view, a square enlarged in G (F), an arm spine articulation (G); H–K, lateral arm plates, from distal portion of arms, outer view (H), inner view (I), distal view, a square enlarged in K (J), an arm spine articulation (K); L, M, ventral arm plates, outer view from proximal portion (L) and inner view from distal portion (M) of arms. All lateral arm plates (D–K) are aligned in the same orientation. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal side. Abbreviations: DL, dorsal lobe; E, elevated structure; P, perforation; SA, spine articuration; SP, spur; VL, ventral lobe.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF454CD1FFD33412FF1AF8B8FDB6F86C" httpUri="https://zenodo.org/record/1163192/files/figure.png" pageId="16" pageNumber="17" targetBox="[221,1356,193,1805]" targetPageId="16">
|
||
<paragraph id="8B851C59FFD33412FF1AF8B8FDB6F86C" blockId="16.[151,1436,1833,2023]" pageId="16" pageNumber="17">
|
||
<emphasis id="B94EC04BFFD33412FF1AF8B8FEBFF8B4" bold="true" box="[151,288,1833,1855]" pageId="16" pageNumber="17">FIGURE 11.</emphasis>
|
||
<taxonomicName id="4C3A67DAFFD33412FEA5F8B8FDBCF8B4" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[296,547,1833,1855]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFD33412FEA5F8B8FDBCF8B4" box="[296,547,1833,1855]" italics="true" pageId="16" pageNumber="17">Ophiozonella cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFD33412FDA6F8B8FDF8F8B4" box="[555,615,1833,1855]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B94EC04BFFD33412FDA6F8B8FDF8F8B4" bold="true" box="[555,615,1833,1855]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
a paratype (RUMF-ZE-02009). SEM images of ossicles. A–E, vertebrae, from proximal portion of arms, proximal view (A), distal view (B), aboral view (C) and oral view (D), a square enlarged from D (E); F–H, vertebrae, from distal portion of arms, proximal view (F), distal view (G) and oral view (H); I–K, arm spines, from proximal portion of arms (I), distal portion of arms, inner (J) and outer (K) spines. Orientations: ab, aboral side; dis, distal side; o, oral side; pro, proximal. Abbreviations: AF, aboral muscle flange; D, depression for tentacle; OF, oral muscle flange; RWC, radial water canal; ST, secondary tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B851C59FFD23413FF4AFF09FC0FFF73" blockId="17.[151,1436,151,752]" pageId="17" pageNumber="18">
|
||
<emphasis id="B94EC04BFFD23413FF4AFF09FE1DFF3B" box="[199,386,152,176]" italics="true" pageId="17" pageNumber="18">(1) Oral shields.</emphasis>
|
||
The oral shields of
|
||
<taxonomicName id="4C3A67DAFFD23413FDE5FF09FC9EFF3B" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[616,769,152,176]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFD23413FDE5FF09FC9EFF3B" box="[616,769,152,176]" italics="true" pageId="17" pageNumber="18">O. cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFD23413FC84FF06FCD6FF3B" box="[777,841,151,176]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="B94EC04BFFD23413FC84FF06FCD6FF3B" bold="true" box="[777,841,151,176]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are as long as wide, with slightly concave proximal sides that end in a fairly sharp point medially, whereas those of
|
||
<taxonomicName id="4C3A67DAFFD23413FCE2FF2DFC6DFF5F" baseAuthorityName="Lyman" baseAuthorityYear="1883" box="[879,1010,188,212]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="clypeata">
|
||
<emphasis id="B94EC04BFFD23413FCE2FF2DFC6DFF5F" box="[879,1010,188,212]" italics="true" pageId="17" pageNumber="18">O. clypeata</emphasis>
|
||
</taxonomicName>
|
||
are 1.5 times longer than wide, blunt spear-head sharp, with acute proximal point (
|
||
<figureCitation id="130100DCFFD23413FD1CFF4EFD78FF73" box="[657,743,223,248]" captionStart="FIGURE 8" captionStartId="12.[151,250,1916,1938]" captionTargetBox="[152,1435,409,1895]" captionTargetId="figure@12.[151,1436,409,1895]" captionTargetPageId="12" captionText="FIGURE 8. Ophiozonella cavernalis n. sp. holotype (RUMF-ZE-02008). A, aboral disc; B, aboral peripheral disc and proximal portion of arm; C, aboral central disc; D, oral disc and proximal portion of arms; E, oral disc; F, interradial oral disc. Abbreviations: ASH, adoral shield; GS, genital slit; OP, oral plate; OS, oral shield;P, oral papillae; T, tooth." httpUri="https://zenodo.org/record/1163186/files/figure.png" pageId="17" pageNumber="18">Fig. 8E</figureCitation>
|
||
,
|
||
<bibRefCitation id="EFAB61A8FFD23413FD7FFF71FC1CFF73" author="Lyman" box="[754,899,224,248]" pageId="17" pageNumber="18" refString="Lyman, T. (1883) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Caribbean Sea (1878 - 79), and along the Atlantic coast of the United States during the summer of 1880, by the U. S. coast survey steamer " Blake, " commander J. R. Bartlett, U. S. N., commanding. XX. Report on the Ophiuroidea. Bulletin of the Museum of Comparative Zoology, 10 (6), 227 - 287." type="journal article" year="1883">Lyman 1883</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3204FD2FFD23413FF4AFE95FDC9FD7B" pageId="17" pageNumber="18" type="description">
|
||
<paragraph id="8B851C59FFD23413FF4AFE95FDA5FEEE" blockId="17.[151,1436,151,752]" pageId="17" pageNumber="18">
|
||
<emphasis id="B94EC04BFFD23413FF4AFE95FEAEFE90" box="[199,305,260,284]" italics="true" pageId="17" pageNumber="18">(2) Arms.</emphasis>
|
||
Ratio of the longest entire arm to disc diameter is 9 times in
|
||
<taxonomicName id="4C3A67DAFFD23413FC6CFE95FBE7FE97" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[993,1144,260,284]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFD23413FC6CFE95FBE7FE97" box="[993,1144,260,284]" italics="true" pageId="17" pageNumber="18">O. cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFD23413FBF2FE95FB21FE96" box="[1151,1214,260,285]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="B94EC04BFFD23413FBF2FE95FB21FE96" bold="true" box="[1151,1214,260,285]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and at most
|
||
<quantity id="4CC2B1BCFFD23413FAC3FE95FAE5FE96" box="[1358,1402,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="17" pageNumber="18" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="4C3A67DAFFD23413FA0CFE95FE4BFECB" authority="(H. L. Clark, 1908)" authorityName="H. L. Clark" authorityYear="1908" baseAuthorityName="H. L. Clark" baseAuthorityYear="1908" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="clypeata">
|
||
<emphasis id="B94EC04BFFD23413FA0CFE95FF69FECB" italics="true" pageId="17" pageNumber="18">O. clypeata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAB61A8FFD23413FE88FEB9FE53FECB" author="Clark" box="[261,460,295,320]" pageId="17" pageNumber="18" refString="Clark, H. L. (1908) Some Japanese and East Indian echinoderms. Bulletin of the Museum of Comparative Zoology at Harvard College, 51 (11), 279 - 311." type="journal article" year="1908">H. L. Clark, 1908</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. For examination of other
|
||
<specimenCount id="9D3CD7D0FFD23413FD72FEB9FC0FFECB" box="[767,912,296,320]" pageId="17" pageNumber="18" type="generic" typeStatus="paratypes">18 paratypes</specimenCount>
|
||
, the smallest number of the ratio of the longest entire arm/disc diameter was 8 times.
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFD23413FF4AFEE1FDFCFE5B" blockId="17.[151,1436,151,752]" pageId="17" pageNumber="18">
|
||
In conjunction with the wide separation of their distribution (
|
||
<taxonomicName id="4C3A67DAFFD23413FCFDFEE1FB98FE03" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[880,1031,368,392]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFD23413FCFDFEE1FB98FE03" box="[880,1031,368,392]" italics="true" pageId="17" pageNumber="18">O. cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFD23413FB80FEFEFBD3FE03" box="[1037,1100,367,392]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="B94EC04BFFD23413FB80FEFEFBD3FE03" bold="true" box="[1037,1100,367,392]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is from
|
||
<quantity id="4CC2B1BCFFD23413FB24FEE1FA9FFE0C" box="[1193,1280,368,392]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.7" metricValueMin="0.8" pageId="17" pageNumber="18" unit="m" value="17.5" valueMax="27.0" valueMin="8.0">8–27 m</quantity>
|
||
,
|
||
<collectingCountry id="F32D5CC9FFD23413FA86FEE1FAD4FE0C" box="[1291,1355,368,391]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
|
||
and
|
||
<taxonomicName id="4C3A67DAFFD23413FA0CFEE1FF69FE27" baseAuthorityName="H. L. Clark" baseAuthorityYear="1908" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="clypeata">
|
||
<emphasis id="B94EC04BFFD23413FA0CFEE1FF69FE27" italics="true" pageId="17" pageNumber="18">O. clypeata</emphasis>
|
||
</taxonomicName>
|
||
is from
|
||
<quantity id="4CC2B1BCFFD23413FEDEFE05FE4BFE27" box="[339,468,404,429]" metricMagnitude="2" metricUnit="m" metricValue="3.585" metricValueMax="5.59" metricValueMin="1.58" pageId="17" pageNumber="18" unit="m" value="358.5" valueMax="559.0" valueMin="158.0">158–559 m</quantity>
|
||
, Galapagos and Caribbean Sea), these two species should be distinguished as different species (
|
||
<bibRefCitation id="EFAB61A8FFD23413FF7BFE29FE17FE5B" author="Lyman" box="[246,392,440,464]" pageId="17" pageNumber="18" refString="Lyman, T. (1883) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Caribbean Sea (1878 - 79), and along the Atlantic coast of the United States during the summer of 1880, by the U. S. coast survey steamer " Blake, " commander J. R. Bartlett, U. S. N., commanding. XX. Report on the Ophiuroidea. Bulletin of the Museum of Comparative Zoology, 10 (6), 227 - 287." type="journal article" year="1883">Lyman 1883</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAB61A8FFD23413FE19FE29FDC9FE5B" author="Clark" box="[404,598,439,464]" pageId="17" pageNumber="18" refString="Clark, H. L. (1917) Reports on the scientific results of the expedition to the eastern tropical pacific in charge of Alexander Agassiz, by the U. S. Fish Commission Steamer " Albatross, " from October, 1904, to March, 1905, Lieut. Commander L. M. Garrett, U. S. N., Commanding. XXX. Ophiuroidea. Bulletin of the Museum of Comparative Zoology at Harvard College, 61 (12), 429 - 451." type="journal article" year="1917">H. L. Clark 1917</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFD23413FF4AFE4DFA03FD0F" blockId="17.[151,1436,151,752]" pageId="17" pageNumber="18">
|
||
Three species of
|
||
<taxonomicName id="4C3A67DAFFD23413FE19FE4DFDB4FE7F" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[404,555,476,500]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus" sensu="stricto">
|
||
<emphasis id="B94EC04BFFD23413FE19FE4DFDB4FE7F" box="[404,555,476,500]" italics="true" pageId="17" pageNumber="18">Ophiozonella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFD23413FDBBFE4CFDE7FE7F" box="[566,632,477,500]" pageId="17" pageNumber="18" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C3A67DAFFD23413FD05FE4DFC30FE7F" authority="Koehler, 1922" authorityName="Koehler" authorityYear="1922" box="[648,943,476,501]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="subtilis">
|
||
<emphasis id="B94EC04BFFD23413FD05FE4DFC9FFE7F" box="[648,768,476,500]" italics="true" pageId="17" pageNumber="18">O. subtilis</emphasis>
|
||
<bibRefCitation id="EFAB61A8FFD23413FC86FE4DFC30FE7F" author="Koehler" box="[779,943,476,501]" pageId="17" pageNumber="18" refString="Koehler, R. (1922) Contributions to the biology of the Philippine Archipelago and adjacent regions. Ophiurians of the Philippine seas and adjacent waters. Bulletin of the United States National Museum, 100, 1 - 486." type="journal article" year="1922">Koehler, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3A67DAFFD23413FC4FFE4DFB65FE7F" authority="(Lyman, 1878)" authorityName="Lyman" authorityYear="1878" baseAuthorityName="Lyman" baseAuthorityYear="1878" box="[962,1274,476,500]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="insulata">
|
||
<emphasis id="B94EC04BFFD23413FC4FFE4DFBDCFE7F" box="[962,1091,476,500]" italics="true" pageId="17" pageNumber="18">O. insulata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAB61A8FFD23413FBDBFE4DFB6DFE7F" author="Lyman" box="[1110,1266,476,500]" pageId="17" pageNumber="18" refString="Lyman, T. (1878) Ophiurans and Astrophytons of the Exploring Voyage of H. M. S. " Challenger, " under Prof. Wyville Thomson, F. R. S. Bulletin of the Museum of Comparative Zoology, 5, 65 - 168." type="journal article" year="1878">Lyman, 1878</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A67DAFFD23413FAB7FE4DFED4FD93" authority="(Koehler, 1904)" authorityName="Koehler" authorityYear="1904" baseAuthorityName="Koehler" baseAuthorityYear="1904" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="casta">
|
||
<emphasis id="B94EC04BFFD23413FAB7FE4DFA03FE7F" box="[1338,1436,476,500]" italics="true" pageId="17" pageNumber="18">O. casta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAB61A8FFD23413FF13FE6EFEDCFD93" author="Koehler" box="[158,323,511,536]" pageId="17" pageNumber="18" refString="Koehler, R. (1904) Ophiures de l'expedition du Siboga. Part 1. Ophiures de mer profonde. Siboga Expeditie, 45 a, 1 - 176." type="book chapter" year="1904">Koehler, 1904</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
with more than 3 arm spines have been recorded from Indo-Pacific. These three species can be distinguished from
|
||
<taxonomicName id="4C3A67DAFFD23413FEF8FDB5FD15FDB7" authority="Okanishi & Fujita, 2018" authorityName="Okanishi & Fujita" authorityYear="2018" box="[373,650,548,572]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cavernalis" status="sp. nov.">
|
||
<emphasis id="B94EC04BFFD23413FEF8FDB5FD15FDB7" box="[373,650,548,572]" italics="true" pageId="17" pageNumber="18">Ophiozonella cavernalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D7D30FFD23413FD1EFDB5FD4BFDB6" box="[659,724,548,573]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="B94EC04BFFD23413FD1EFDB5FD4BFDB6" bold="true" box="[659,724,548,573]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
mainly by the following characters:
|
||
<taxonomicName id="4C3A67DAFFD23413FBF1FDB5FAF1FDB7" authorityName="Koehler" authorityYear="1922" box="[1148,1390,548,572]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="subtilis">
|
||
<emphasis id="B94EC04BFFD23413FBF1FDB5FAF1FDB7" box="[1148,1390,548,572]" italics="true" pageId="17" pageNumber="18">Ophiozonella subtilis</emphasis>
|
||
</taxonomicName>
|
||
has smaller and narrower oral shields;
|
||
<taxonomicName id="4C3A67DAFFD23413FD97FDD9FD0AFDEB" box="[538,661,584,608]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="insulate">
|
||
<emphasis id="B94EC04BFFD23413FD97FDD9FD0AFDEB" box="[538,661,584,608]" italics="true" pageId="17" pageNumber="18">O. insulate</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3A67DAFFD23413FD46FDD9FCB7FDEB" baseAuthorityName="Koehler" baseAuthorityYear="1904" box="[715,808,584,608]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="casta">
|
||
<emphasis id="B94EC04BFFD23413FD46FDD9FCB7FDEB" box="[715,808,584,608]" italics="true" pageId="17" pageNumber="18">O. casta</emphasis>
|
||
</taxonomicName>
|
||
have approximately 15 and 10 disc scales on interradial oral surface, respectively, which are fewer than 30–50 scales of
|
||
<taxonomicName id="4C3A67DAFFD23413FCD2FDFDFC69FD0F" authorityName="Okanishi & Fujita" authorityYear="2018" box="[863,1014,620,644]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiozonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cavernalis">
|
||
<emphasis id="B94EC04BFFD23413FCD2FDFDFC69FD0F" box="[863,1014,620,644]" italics="true" pageId="17" pageNumber="18">O. cavernalis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFAB61A8FFD23413FB89FDFDFB03FD0F" author="Lyman" box="[1028,1180,620,644]" pageId="17" pageNumber="18" refString="Lyman, T. (1878) Ophiurans and Astrophytons of the Exploring Voyage of H. M. S. " Challenger, " under Prof. Wyville Thomson, F. R. S. Bulletin of the Museum of Comparative Zoology, 5, 65 - 168." type="journal article" year="1878">Lyman, 1878</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAB61A8FFD23413FB2AFDFDFAD6FD0F" author="Koehler" box="[1191,1353,620,645]" pageId="17" pageNumber="18" refString="Koehler, R. (1904) Ophiures de l'expedition du Siboga. Part 1. Ophiures de mer profonde. Siboga Expeditie, 45 a, 1 - 176." type="book chapter" year="1904">Koehler, 1904</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAB61A8FFD23413FAD9FDFDFA0FFD0F" author="Koehler" box="[1364,1424,620,644]" pageId="17" pageNumber="18" refString="Koehler, R. (1922) Contributions to the biology of the Philippine Archipelago and adjacent regions. Ophiurians of the Philippine seas and adjacent waters. Bulletin of the United States National Museum, 100, 1 - 486." type="journal article" year="1922">1922</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B851C59FFD23413FF4AFD1EFDC9FD7B" blockId="17.[151,1436,151,752]" pageId="17" pageNumber="18">
|
||
The first anchialine-endemic ophiuroid species (
|
||
<taxonomicName id="4C3A67DAFFD23413FD60FD01FC45FD23" authorityName="Pomory, Carpenter & Winter" authorityYear="2011" box="[749,986,656,680]" class="Ophiuroidea" family="Amphilepididae" genus="Amphicutis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="stygobita">
|
||
<emphasis id="B94EC04BFFD23413FD60FD01FC45FD23" box="[749,986,656,680]" italics="true" pageId="17" pageNumber="18">Amphicutis stygobita</emphasis>
|
||
</taxonomicName>
|
||
) was found in Bernier Cave,
|
||
<collectingCountry id="F32D5CC9FFD23413FAA3FD1EFA08FD23" box="[1326,1431,655,680]" name="Bahamas" pageId="17" pageNumber="18">Bahamas</collectingCountry>
|
||
, Atlantic Ocean (
|
||
<bibRefCitation id="EFAB61A8FFD23413FEC2FD25FDB3FD47" author="Pomory" box="[335,556,692,716]" pageId="17" pageNumber="18" refString="Pomory, C. M., Carpenter, J. H. & Winter, J. H. (2011) Amphicutis stygobita, a new genus and new species of brittle star (Echinodermata: Ophiuroidea: Ophiurida: Amphilepididae) found in Bernier Cave, an anchialine cave on San Salvador Island, Bahamas. Zootaxa, 3133, 50 - 68." type="journal article" year="2011">
|
||
Pomory
|
||
<emphasis id="B94EC04BFFD23413FE3DFD24FE76FD47" box="[432,489,692,716]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
). This is the second discovery of an anchialine-endemic ophiuroid species and the first finding from the Pacific Ocean.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |