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<document id="363BC8C196A9942CED1749CFB1487845" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE3B3FF26FF75F8E3BC7CFE99" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madagaster franzi Jach" docType="treatment" docVersion="9" lastPageNumber="184" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="181" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:title id="210DB4B5F4A27E1F2F0ED63630142025">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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<mods:namePart id="ED5D085EADF7BBB3AF6FD37B16871C80">Perkins, Philip D.</mods:namePart>
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<mods:date id="92E41CF0CA3B89223D7BB3D8D5EBC7A1">2017</mods:date>
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<treatment id="038587BBE3B3FF26FF75F8E3BC7CFE99" ID-DOI="http://doi.org/10.5281/zenodo.5323730" ID-GBIF-Taxon="137233386" ID-Zenodo-Dep="5323730" LSID="urn:lsid:plazi:treatment:038587BBE3B3FF26FF75F8E3BC7CFE99" httpUri="http://treatment.plazi.org/id/038587BBE3B3FF26FF75F8E3BC7CFE99" lastPageId="183" lastPageNumber="184" pageId="180" pageNumber="181">
<subSubSection id="C3366526E3B3FF25FF75F8E3BB98F807" pageId="180" pageNumber="181" type="nomenclature">
<paragraph id="8B9336ADE3B3FF25FF75F8E3B8F6F8E5" blockId="180.[151,676,1862,1922]" box="[151,458,1862,1888]" pageId="180" pageNumber="181">
<heading id="D0DB81C1E3B3FF25FF75F8E3B8F6F8E5" bold="true" box="[151,458,1862,1888]" fontSize="11" level="1" pageId="180" pageNumber="181" reason="1">
<taxonomicName id="4C2C4D2EE3B3FF25FF75F8E3B8F6F8E5" authority="(Jach)" authorityName="Jach" box="[151,458,1862,1888]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B958EABFE3B3FF25FF75F8E3B8F6F8E5" bold="true" box="[151,458,1862,1888]" pageId="180" pageNumber="181">
<emphasis id="B958EABFE3B3FF25FF75F8E3B848F8E5" bold="true" box="[151,372,1862,1888]" italics="true" pageId="180" pageNumber="181">Madagaster franzi</emphasis>
(Jäch)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9336ADE3B3FF25FF75F8CCBB98F807" blockId="180.[151,676,1862,1922]" box="[151,676,1897,1922]" pageId="180" pageNumber="181">
<figureCitation id="13172A28E3B3FF25FF75F8CCB9C3F807" box="[151,255,1897,1922]" captionStart="FIGURE 175" captionStartId="183.[151,250,1551,1573]" captionText="FIGURE 175. Madagaster franzi, holotype, habitus dorsal, lateral, and ventral." pageId="180" pageNumber="181">Figs. 175</figureCitation>
(habitus), 177 (aedeagus), 301 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B3FF24FF75F814B822F94E" lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181" type="reference_group">
<paragraph id="8B9336ADE3B3FF24FF75F814B822F94E" blockId="180.[151,495,1969,1991]" lastBlockId="181.[151,1437,1678,2027]" lastPageId="181" lastPageNumber="182" pageId="180" pageNumber="181">
<materialsCitation id="3B443CF0E3B3FF24FF75F814B822F94E" ID-GBIF-Occurrence="1697157782" collectingDate="11. IV" collectionCode="NMW" collectorName="Plateau du Tampoketsz &amp; H. Franz" country="Madagascar" lastPageId="181" lastPageNumber="182" location="Madagascar" pageId="180" pageNumber="181" specimenCount="2" typeStatus="holotype">
<taxonomicName id="4C2C4D2EE3B3FF25FF75F814B8D3F842" authority="Jach (1994: 92)" authorityName="Jach" authorityPageNumber="92" authorityYear="1994" box="[151,495,1969,1991]" class="Insecta" family="Hydraenidae" genus="Gondraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="franzi">
<emphasis id="B958EABFE3B3FF25FF75F814B877F842" box="[151,331,1969,1991]" italics="true" pageId="180" pageNumber="181">Gondraena franzi</emphasis>
Jäch (1994: 92)
</taxonomicName>
<emphasis id="B958EABFE3B2FF24FF75F92BB8F4F922" bold="true" box="[151,456,1678,1703]" pageId="181" pageNumber="182">
<typeStatus id="5497880FE3B2FF24FF75F92BB9EEF922" box="[151,210,1678,1703]" pageId="181" pageNumber="182">Type</typeStatus>
Material.
<typeStatus id="5497880FE3B2FF24FEB9F92BB8F4F922" box="[347,456,1678,1703]" pageId="181" pageNumber="182" type="holotype">Holotype</typeStatus>
</emphasis>
(male): &quot;
<collectingCountry id="F33B763DE3B2FF24FDD8F92BBBFAF922" box="[570,710,1678,1703]" name="Madagascar" pageId="181" pageNumber="182">Madagascar</collectingCountry>
,
<collectorName id="26D9537BE3B2FF24FD3AF92BBADBF922" box="[728,999,1678,1703]" pageId="181" pageNumber="182">Plateau du Tampoketsz</collectorName>
lg.
<collectorName id="26D9537BE3B2FF24FBFCF92ABDBAF923" box="[1054,1158,1679,1702]" pageId="181" pageNumber="182">H. Franz</collectorName>
[
<collectingDate id="EFD6E985E3B2FF24FB78F92ABDE2F922" box="[1178,1246,1679,1703]" pageId="181" pageNumber="182">
<date id="FF92106DE3B2FF24FB78F92ABDE2F922" box="[1178,1246,1679,1703]" pageId="181" pageNumber="182">11.IV</date>
</collectingDate>
] 1969&quot; (
<collectionCode id="ED3DAE68E3B2FF24FAABF92ABCA8F922" box="[1353,1428,1679,1703]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="181" pageNumber="182">NMW</collectionCode>
) (examined).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF536625E3B2FF24FF75F9F5BBC5F9E2" box="[151,761,1616,1639]" pageId="181" pageNumber="182">
<paragraph id="8B9336ADE3B2FF24FF75F9F5BBC5F9E2" blockId="181.[151,761,1616,1639]" box="[151,761,1616,1639]" pageId="181" pageNumber="182">
<emphasis id="B958EABFE3B2FF24FF75F9F5B851F9E3" bold="true" box="[151,365,1616,1639]" pageId="181" pageNumber="182">FIGURES 172173.</emphasis>
<taxonomicName id="4C2C4D2EE3B2FF24FE91F9F4BB2FF9E2" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[371,531,1617,1639]" class="Insecta" family="Hydraenidae" genus="Menomadraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="170" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B958EABFE3B2FF24FE91F9F4BB2FF9E2" box="[371,531,1617,1639]" italics="true" pageId="181" pageNumber="182">Menomadraena</emphasis>
</taxonomicName>
, aedeagi of holotypes.
</paragraph>
</caption>
<subSubSection id="C3366526E3B2FF24FF25F973BA9FF96B" box="[199,931,1750,1775]" pageId="181" pageNumber="182" type="diagnosis">
<paragraph id="8B9336ADE3B2FF24FF25F973BA9FF96B" blockId="181.[151,1437,1678,2027]" box="[199,931,1750,1775]" pageId="181" pageNumber="182">
<emphasis id="B958EABFE3B2FF24FF25F973B8EEF96A" bold="true" box="[199,466,1750,1775]" pageId="181" pageNumber="182">Differential Diagnosis.</emphasis>
Refer to the diagnosis of
<taxonomicName id="4C2C4D2EE3B2FF24FD11F972BAA0F96A" authorityName="Perkins" authorityYear="2017" box="[755,924,1750,1775]" class="Insecta" family="Hydraenidae" genus="Madagaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="simplissima">
<emphasis id="B958EABFE3B2FF24FD11F972BAA0F96A" box="[755,924,1750,1775]" italics="true" pageId="181" pageNumber="182">M. simplissima</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E3B2FF26FF25F95FBC7CFE99" lastPageId="183" lastPageNumber="184" pageId="181" pageNumber="182" type="description">
<paragraph id="8B9336ADE3B2FF24FF25F95FB85AF8DE" blockId="181.[151,1437,1678,2027]" pageId="181" pageNumber="182">
<emphasis id="B958EABFE3B2FF24FF25F95FB86DF896" bold="true" box="[199,337,1786,1811]" pageId="181" pageNumber="182">Description</emphasis>
. Size:
<typeStatus id="5497880FE3B2FF24FE41F95FBB39F896" box="[419,517,1786,1811]" pageId="181" pageNumber="182" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.70/0.73; head width 0.46; pronotum 0.40/0.61, PA 0.49, PB 0.51; elytra 1.05/0.73. Dorsum and venter dark brown to piceous, legs brown, ocelli light brown.
</paragraph>
<paragraph id="8B9336ADE3B2FF27FF25F8C3BBA6FF50" blockId="181.[151,1437,1678,2027]" lastBlockId="182.[151,1436,151,213]" lastPageId="182" lastPageNumber="183" pageId="181" pageNumber="182">Entire dorsum dull, with very small, closely spaced granules, each with a short recumbent seta. Labrum with V-shaped apicomedian notch, very sparsely punctate and microreticulate. Anterior margin of clypeus not overhanging labrum. Labroclypeal suture moderately deep. Short, deep sulcus in front of each ocellus. Frons with deep concavity between ocelli. Maxillary palpi ca. as along as antenna, almost as long as protibia, last palpomere widest at midlength, distinctly tapering to pointed apex. Antenna with penultimate antennomere larger than ultimate. Posterior margin of eye not carinate.</paragraph>
<caption id="DF536625E3B1FF27FF75F987BA5EF9BD" box="[151,866,1570,1592]" pageId="182" pageNumber="183">
<paragraph id="8B9336ADE3B1FF27FF75F987BA5EF9BD" blockId="182.[151,866,1570,1592]" box="[151,866,1570,1592]" pageId="182" pageNumber="183">
<emphasis id="B958EABFE3B1FF27FF75F987B810F9B2" bold="true" box="[151,300,1570,1592]" pageId="182" pageNumber="183">FIGURE 174.</emphasis>
<taxonomicName id="4C2C4D2EE3B1FF27FED0F987B8D7F9BD" authorityName="Jäch" authorityYear="1994" box="[306,491,1570,1592]" class="Insecta" family="Hydraenidae" genus="Gondraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B958EABFE3B1FF27FED0F987B8D7F9BD" box="[306,491,1570,1592]" italics="true" pageId="182" pageNumber="183">Gondraena indica</emphasis>
</taxonomicName>
, holotype, habitus dorsal and lateral.
</paragraph>
</caption>
<paragraph id="8B9336ADE3B1FF27FF25F9C5BB73F945" blockId="182.[151,1437,1631,2016]" pageId="182" pageNumber="183">Pronotum with two narrowly separated midlongitudinal fovea, anterior ca. twice as long as posterior; each side with two admedian and two sublateral foveae; transverse fovea along base on each side; lateral margin rounded. Postocular emarginations very shallow.</paragraph>
<paragraph id="8B9336ADE3B1FF27FF25F96EBB99F8AE" blockId="182.[151,1437,1631,2016]" pageId="182" pageNumber="183">Elytral suture raised. Each elytron with four very low, subcostate unilinear rows of granules, rows terminating before reaching elytral margin. Areas between rows of granules with four very irregular rows of granules, each granule with short seta. Elytral apices truncate.</paragraph>
<paragraph id="8B9336ADE3B1FF27FF25F89DB8BDF8F1" blockId="182.[151,1437,1631,2016]" pageId="182" pageNumber="183">Mentum small, punctate and microreticulate, dull; postmentum contiguously micropunctulate and microreticulate, dull.</paragraph>
<paragraph id="8B9336ADE3B1FF27FF25F8DABAC4F865" blockId="182.[151,1437,1631,2016]" pageId="182" pageNumber="183">Mesoventrite without tubercle on intercoxal process. Metaventrite densely micropunctulate and with very short dense vestiture, very finely microreticulate or micropunctulate; midlongitudinal depression shallow. First four Abdominal ventrites and anterior margin of fifth with hydrofuge pubescence.</paragraph>
<paragraph id="8B9336ADE3B0FF26FF25FF32BB97FF35" blockId="183.[151,1436,151,285]" box="[199,683,151,176]" pageId="183" pageNumber="184">All tibiae straight, with sparse short spines.</paragraph>
<paragraph id="8B9336ADE3B0FF26FF25FF19BC7CFE99" blockId="183.[151,1436,151,285]" pageId="183" pageNumber="184">
Remarks. The &quot;putative synapomorphy&quot; of the shape of the antennal club, specifically the terminal article being shorter than the preterminal, noted by
<bibRefCitation id="EFBD4B5CE3B0FF26FD73FF7ABA4DFF7D" author="Perkins" box="[657,881,223,248]" pageId="183" pageNumber="184" refString="Perkins, P. D. (1997) Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">Perkins (1997: 179)</bibRefCitation>
must now be put aside. After more species were studied it was found that the relative sizes of the antennomeres varies in both
<emphasis id="B958EABFE3B0FF26FC1EFEA1BDBFFE98" box="[1020,1155,260,285]" italics="true" pageId="183" pageNumber="184">Madagaster</emphasis>
and
<taxonomicName id="4C2C4D2EE3B0FF26FB58FEA1BC05FE98" box="[1210,1337,260,285]" class="Insecta" family="Hydraenidae" genus="Gondraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="B958EABFE3B0FF26FB58FEA1BC05FE98" box="[1210,1337,260,285]" italics="true" pageId="183" pageNumber="184">Gondraena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>