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<document ID-DOI="http://doi.org/10.5281/zenodo.6213080" ID-GBIF-Dataset="e04117e5-9306-489d-a0a3-9dbd97774f74" ID-GBIF-Taxon="119613328" ID-ISSN="1175-5326" ID-Zenodo-Dep="6213080" checkinTime="1461057042134" checkinUser="plazi" docAuthor="Hong, Yong &amp; James, Samuel W." docDate="2009" docId="03858791FFEDFFF3FF67FE5DFD84F389" docLanguage="en" docName="zt02059p045.pdf" docOrigin="Zootaxa 2059" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@46f0bf0d" docStyleName="zootaxa.2007.monograph" docTitle="Pheretima doriae Hong &amp; James, 2009, sp. nov." docType="treatment" docVersion="8" lastPageNumber="39" masterDocId="FFBCFFE9FFEBFFF5FFF0FF8AFFEBF41B" masterDocTitle="New earthworms of the Pheretima urceolata species group (Oligochaeta: Megascolecidae) from southern Luzon, Philippines" masterLastPageNumber="45" masterPageNumber="33" pageId="6" pageNumber="39" updateTime="1645486389336" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New earthworms of the Pheretima urceolata species group (Oligochaeta: Megascolecidae) from southern Luzon, Philippines</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hong, Yong</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>James, Samuel W.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="volume">
<mods:number>2059</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>33</mods:start>
<mods:end>45</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.186796</mods:identifier>
<mods:identifier type="GBIF-Dataset">e04117e5-9306-489d-a0a3-9dbd97774f74</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">186796</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6213080" ID-GBIF-Taxon="119613328" ID-Zenodo-Dep="6213080" LSID="urn:lsid:plazi:treatment:03858791FFEDFFF3FF67FE5DFD84F389" httpUri="http://treatment.plazi.org/id/03858791FFEDFFF3FF67FE5DFD84F389" lastPageNumber="39" pageId="6" pageNumber="39">
<subSubSection pageId="6" pageNumber="39" type="nomenclature">
<paragraph blockId="6.[151,461,471,532]" box="[151,461,471,497]" pageId="6" pageNumber="39">
<heading bold="true" box="[151,461,471,497]" fontSize="11" level="1" pageId="6" pageNumber="39" reason="1">
<emphasis bold="true" box="[151,461,471,497]" pageId="6" pageNumber="39">
<taxonomicName box="[151,360,471,497]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="doriae" status="sp. nov.">
<emphasis bold="true" box="[151,360,471,497]" italics="true" pageId="6" pageNumber="39">Pheretima doriae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[367,461,471,497]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[151,461,471,532]" box="[151,294,506,532]" pageId="6" pageNumber="39">
(
<figureCitation box="[159,228,506,532]" captionStart="FIGURE 2" captionStartId="7.[151,255,1253,1277]" captionTargetBox="[332,1218,187,1219]" captionTargetId="figure@7.[327,1227,177,1240]" captionTargetPageId="7" captionText="FIGURE 2. Pheretima doriae sp. nov. A: ventral view; B: spermatheca. Pheretima batoensis sp. nov. C: ventral view; D: spermatheca. Scale bars = 2.5 mm (ventral views), 1 mm (spermathecae)." httpUri="https://zenodo.org/record/186798/files/figure.png" pageId="6" pageNumber="39">
Fig.
<number box="[214,228,506,532]" pageId="6" pageNumber="39" value="2.0">2</number>
</figureCitation>
AB)
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="39" type="materials_examined">
<paragraph blockId="6.[151,1437,586,1938]" pageId="6" pageNumber="39">
<emphasis bold="true" box="[151,267,586,612]" pageId="6" pageNumber="39">Material.</emphasis>
<typeStatus box="[277,391,586,612]" pageId="6" pageNumber="39">Holotype</typeStatus>
: Clitellate (NMA 004318):
<collectingCountry box="[727,862,586,612]" name="Philippines" pageId="6" pageNumber="39">Philippines</collectingCountry>
, Camarines Sur, Mt. Isarog (
<geoCoordinate box="[1214,1365,586,612]" direction="north" orientation="latitude" pageId="6" pageNumber="39" precision="9" value="13.661667">
<number box="[1214,1244,586,612]" pageId="6" pageNumber="39" value="13.0">13</number>
°
<number box="[1266,1338,586,612]" pageId="6" pageNumber="39" value="39.7">39.70</number>
N
</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" pageId="6" pageNumber="39" precision="9" value="123.367836">
<number box="[1380,1424,586,612]" pageId="6" pageNumber="39" value="123.0">123</number>
°
<number box="[151,224,626,652]" pageId="6" pageNumber="39" value="22.07">22.07</number>
E
</geoCoordinate>
),
<quantity box="[269,361,626,652]" metricMagnitude="3" metricUnit="m" metricValue="1.7449999999999999" pageId="6" pageNumber="39" unit="m" value="1745.0">1745 m</quantity>
, arboreal,
<date box="[484,644,626,652]" pageId="6" pageNumber="39" value="2001-05-16">
<number box="[484,513,626,652]" pageId="6" pageNumber="39" value="16.0">16</number>
May
<number box="[583,644,626,652]" pageId="6" pageNumber="39" value="2001.0">2001</number>
</date>
, S. W. James coll.
<number box="[866,880,626,652]" pageId="6" pageNumber="39" value="2.0">2</number>
<typeStatus box="[888,1005,626,652]" pageId="6" pageNumber="39">paratypes</typeStatus>
:
<number box="[1016,1030,626,652]" pageId="6" pageNumber="39" value="1.0">1</number>
clitellate (NMA 004323),
<number box="[1345,1360,626,652]" pageId="6" pageNumber="39" value="1.0">1</number>
clitellate (
<collectionCode box="[207,320,666,692]" pageId="6" pageNumber="39">KUNHM</collectionCode>
003014): Same data as for
<typeStatus box="[639,745,666,692]" pageId="6" pageNumber="39">holotype</typeStatus>
. Other material:
<number box="[940,955,666,692]" pageId="6" pageNumber="39" value="1.0">1</number>
clitellate,
<number box="[1076,1091,666,692]" pageId="6" pageNumber="39" value="2.0">2</number>
semi-clitellates, same data as for
<typeStatus box="[196,303,706,732]" pageId="6" pageNumber="39">holotype</typeStatus>
;
<number box="[318,333,706,732]" pageId="6" pageNumber="39" value="1.0">1</number>
aclitellate, Camarines Sur, Mt. Isarog (
<geoCoordinate box="[815,967,706,732]" direction="north" orientation="latitude" pageId="6" pageNumber="39" precision="9" value="13.661667">
<number box="[815,845,706,732]" pageId="6" pageNumber="39" value="13.0">13</number>
°
<number box="[868,940,706,732]" pageId="6" pageNumber="39" value="39.7">39.70</number>
N
</geoCoordinate>
,
<geoCoordinate box="[983,1147,706,732]" direction="east" orientation="longitude" pageId="6" pageNumber="39" precision="9" value="123.367836">
<number box="[983,1027,706,732]" pageId="6" pageNumber="39" value="123.0">123</number>
°
<number box="[1049,1122,706,732]" pageId="6" pageNumber="39" value="22.07">22.07</number>
E
</geoCoordinate>
),
<quantity box="[1171,1266,706,732]" metricMagnitude="3" metricUnit="m" metricValue="1.7449999999999999" pageId="6" pageNumber="39" unit="m" value="1745.0">1745 m</quantity>
, soil,
<date pageId="6" pageNumber="39" value="2001-05-16">
<number box="[1342,1371,706,732]" pageId="6" pageNumber="39" value="16.0">16</number>
May
<number box="[151,212,746,772]" pageId="6" pageNumber="39" value="2001.0">2001</number>
</date>
, S. W. James coll.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="39" type="etymology">
<paragraph blockId="6.[151,1437,586,1938]" pageId="6" pageNumber="39">
<emphasis bold="true" box="[197,338,786,812]" pageId="6" pageNumber="39">Etymology.</emphasis>
The species is named for Dorothy (Dori) Franke, a student participant in the fieldwork. On the ascent of Mt. Isarog, she elected to remain at the
<number box="[770,829,826,852]" pageId="6" pageNumber="39" value="1745.0">1745</number>
meters site and search there.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="39" type="diagnosis">
<paragraph blockId="6.[151,1437,586,1938]" pageId="6" pageNumber="39">
<emphasis bold="true" box="[197,325,866,892]" pageId="6" pageNumber="39">Diagnosis.</emphasis>
<taxonomicName box="[335,446,866,891]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="urceolata">
<emphasis box="[335,446,866,891]" italics="true" pageId="6" pageNumber="39">urceolata</emphasis>
</taxonomicName>
-group with spermathecal pores and openings of copulatory bursae
<number box="[1252,1303,866,892]" pageId="6" pageNumber="39" value="0.32">0.32</number>
circumference apart, intestinal origin in XVI, hearts in XIXIII, and septum
<number box="[933,947,906,932]" pageId="6" pageNumber="39" value="9.0">9</number>
/
<number box="[957,985,906,932]" pageId="6" pageNumber="39" value="10.0">10</number>
present.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="39" type="description">
<paragraph blockId="6.[151,1437,586,1938]" pageId="6" pageNumber="39">
<emphasis bold="true" box="[197,349,946,972]" pageId="6" pageNumber="39">Description.</emphasis>
Red-brown dorsal pigment, except on setal rings; preclitellate segments with lighter pigment. Dimensions
<quantity box="[375,504,986,1012]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.5" metricValueMin="3.4" pageId="6" pageNumber="39" unit="mm" value="39.5" valueMax="45.0" valueMin="34.0">3445 mm</quantity>
by
<number box="[551,588,986,1012]" pageId="6" pageNumber="39" value="2.4">2.4</number>
at segment X, 2.0 mm at XXX,
<quantity box="[984,1076,986,1012]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="39" unit="mm" value="2.1">2.1 mm</quantity>
at clitellum, segments
<number box="[1355,1432,986,1012]" pageId="6" pageNumber="39" value="81.5" valueMax="86.0" valueMin="77.0">7786</number>
; body circular in cross-section. Setae numbering
<number box="[730,803,1026,1052]" pageId="6" pageNumber="39" value="37.5" valueMax="40.0" valueMin="35.0">3540</number>
at VII,
<number box="[901,974,1026,1052]" pageId="6" pageNumber="39" value="41.5" valueMax="43.0" valueMin="40.0">4043</number>
at XX;
<number box="[1075,1118,1026,1052]" pageId="6" pageNumber="39" value="6.5" valueMax="8.0" valueMin="5.0">58</number>
between male pores, setal formula AA:AB:YZ:ZZ=
<number box="[457,475,1066,1092]" pageId="6" pageNumber="39" value="4.0">4</number>
:
<number box="[480,498,1066,1092]" pageId="6" pageNumber="39" value="2.0">2</number>
:
<number box="[502,518,1066,1092]" pageId="6" pageNumber="39" value="3.0">3</number>
:
<number box="[522,540,1066,1092]" pageId="6" pageNumber="39" value="4.0">4</number>
at XIII. Clitellum annular XIVXVI; setae invisible externally.
</paragraph>
<paragraph blockId="6.[151,1437,586,1938]" pageId="6" pageNumber="39">
First dorsal pores
<number box="[413,441,1109,1135]" pageId="6" pageNumber="39" value="11.0">11</number>
/
<number box="[452,483,1109,1135]" pageId="6" pageNumber="39" value="12.0">12</number>
. One pair of spermathecae deep in
<number box="[907,920,1109,1135]" pageId="6" pageNumber="39" value="5.0">5</number>
/
<number box="[930,944,1109,1135]" pageId="6" pageNumber="39" value="6.0">6</number>
at
<number box="[983,998,1109,1135]" pageId="6" pageNumber="39" value="7.0">7</number>
th
<number box="[1027,1040,1109,1135]" pageId="6" pageNumber="39" value="8.0">8</number>
th setal lines, sub-laterally placed,
<number box="[151,202,1149,1175]" pageId="6" pageNumber="39" value="0.32">0.32</number>
circumference apart, distance between spermathecal pores
<quantity box="[906,1000,1149,1175]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="39" unit="mm" value="2.3">2.3 mm</quantity>
;
<quantity box="[1013,1155,1149,1175]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" pageNumber="39" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.30.4 mm</quantity>
openings of copulatory bursae paired in XVIII at
<number box="[464,479,1192,1218]" pageId="6" pageNumber="39" value="7.0">7</number>
th setal lines, ventro-laterally placed,
<number box="[919,970,1192,1218]" pageId="6" pageNumber="39" value="0.32">0.32</number>
circumference apart, distance between openings
<quantity box="[264,357,1232,1258]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="39" unit="mm" value="1.8">1.8 mm</quantity>
. Genital markings lacking.
</paragraph>
<paragraph blockId="6.[151,1437,586,1938]" pageId="6" pageNumber="39">
<collectingCountry box="[197,262,1272,1298]" name="Spain" pageId="6" pageNumber="39">Septa</collectingCountry>
<number box="[270,283,1272,1298]" pageId="6" pageNumber="39" value="5.0">5</number>
/
<number box="[293,338,1272,1298]" pageId="6" pageNumber="39" value="6.5" valueMax="7.0" valueMin="6.0">67</number>
/
<number box="[347,360,1272,1298]" pageId="6" pageNumber="39" value="8.0">8</number>
thin,
<number box="[429,442,1272,1298]" pageId="6" pageNumber="39" value="8.0">8</number>
/
<number box="[452,466,1272,1298]" pageId="6" pageNumber="39" value="9.0">9</number>
absent,
<number box="[565,579,1272,1298]" pageId="6" pageNumber="39" value="9.0">9</number>
/
<number box="[589,617,1272,1298]" pageId="6" pageNumber="39" value="10.0">10</number>
membranous,
<date box="[793,940,1272,1298]" pageId="6" pageNumber="39" value="1914-10-11" valueMax="1914-10-13" valueMin="1914-10-11">
<number box="[793,822,1272,1298]" pageId="6" pageNumber="39" value="10.0">10</number>
/
<number box="[831,901,1272,1298]" pageId="6" pageNumber="39" value="12.0" valueMax="13.0" valueMin="11.0">1113</number>
/
<number box="[911,940,1272,1298]" pageId="6" pageNumber="39" value="14.0">14</number>
</date>
thin. Gizzard in VIIIIX, intestine begins in XVI, small paired lymph glands from XXVII along dorsal vessel; intestinal caeca simple originating in XXVII, and extending anteriorly about to XXV, short sac with strong septal indentations, typhlosole very low from XXVII. Hearts XIXIII, esophageal, X absent.
</paragraph>
<paragraph blockId="6.[151,1437,586,1938]" pageId="6" pageNumber="39">Ovaries and funnels in XIII. Spermathecae one pair in VI, with nephridia on spermathecal ducts; spermatheca with small broad spherical ampulla, duct shorter than ampulla; diverticulum very small egg-shaped chamber, with short stalk. Male sexual system holandric, testes and funnels in ventral paired sacs in XXI. Seminal vesicles two pairs in XIXII, with elongate fuzzy dorsal lobes, prostates in XVIIXIX bilobed, with muscular ducts, duct crossing top of copulatory bursae from lateral edge to center, attached to top; copulatory bursae without coelomic glands, glandular pads at anterior medial, posterior medial, and anterior lateral sides of copulatory bursa cavity; penis a small bump on roof.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="39" type="discussion">
<paragraph blockId="6.[151,1437,586,1938]" pageId="6" pageNumber="39">
<emphasis bold="true" box="[197,318,1712,1738]" pageId="6" pageNumber="39">Remarks.</emphasis>
<taxonomicName box="[330,539,1712,1737]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="doriae" status="sp. nov.">
<emphasis box="[330,539,1712,1737]" italics="true" pageId="6" pageNumber="39">Pheretima doriae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[550,648,1712,1738]" pageId="6" pageNumber="39">
<taxonomicNameLabel box="[550,648,1712,1738]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has widely spaced spermathecal pores (
<number box="[1140,1193,1712,1738]" pageId="6" pageNumber="39" value="0.32">0.32</number>
circumference) and male openings (
<number box="[341,398,1752,1778]" pageId="6" pageNumber="39" value="0.32">0.32</number>
). Others in the species group with relatively wide spermathecal pore spacing, hearts in XIXIII and an intestinal origin in XVI are
<taxonomicName box="[663,910,1792,1817]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="viracensis" status="sp. nov.">
<emphasis box="[663,910,1792,1817]" italics="true" pageId="6" pageNumber="39">Pheretima viracensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[917,1010,1792,1818]" pageId="6" pageNumber="39">
<taxonomicNameLabel box="[917,1010,1792,1818]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName box="[1025,1144,1792,1818]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="abiadai">
<emphasis box="[1025,1040,1792,1817]" italics="true" pageId="6" pageNumber="39">P</emphasis>
.
<emphasis box="[1054,1144,1792,1817]" italics="true" pageId="6" pageNumber="39">abiadai</emphasis>
</taxonomicName>
, and
<taxonomicName box="[1208,1354,1792,1818]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="nagaensis">
<emphasis box="[1208,1223,1792,1817]" italics="true" pageId="6" pageNumber="39">P</emphasis>
.
<emphasis box="[1237,1354,1792,1817]" italics="true" pageId="6" pageNumber="39">nagaensis</emphasis>
</taxonomicName>
(Table
<number box="[151,165,1832,1858]" pageId="6" pageNumber="39" value="1.0">1</number>
). Among these,
<taxonomicName box="[364,569,1832,1857]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="doriae" status="sp. nov.">
<emphasis box="[364,569,1832,1857]" italics="true" pageId="6" pageNumber="39">Pheretima doriae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[578,673,1832,1858]" pageId="6" pageNumber="39">
<taxonomicNameLabel box="[578,673,1832,1858]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only one with three copulatory bursal pads located on the sides and to have septum
<number box="[462,476,1872,1898]" pageId="6" pageNumber="39" value="9.0">9</number>
/
<number box="[486,514,1872,1898]" pageId="6" pageNumber="39" value="10.0">10</number>
present.
<taxonomicName box="[625,832,1872,1897]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="39" phylum="Annelida" rank="species" species="doriae" status="sp. nov.">
<emphasis box="[625,832,1872,1897]" italics="true" pageId="6" pageNumber="39">Pheretima doriae</emphasis>
</taxonomicName>
<emphasis bold="true" box="[841,937,1872,1898]" pageId="6" pageNumber="39">
<taxonomicNameLabel box="[841,937,1872,1898]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has wider male opening spacing than the other three, whose spacing is
<number box="[497,618,1912,1938]" pageId="6" pageNumber="39" value="0.225" valueMax="0.25" valueMin="0.2">0.200.25</number>
.
</paragraph>
</subSubSection>
</treatment>
</document>