266 lines
35 KiB
XML
266 lines
35 KiB
XML
<document id="ED3D30145036D51A638FCFC9369AC70C" ID-CLB-Dataset="8176" ID-DOI="10.5852/ejt.2021.734.1233" ID-GBIF-Dataset="fd8f5cb9-13e9-4c90-ac15-502fa7ee8d38" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542107" ID-ZooBank="663A0C2D-E3CF-4B0C-8BA1-566B174829A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613423098920" checkinUser="plazi" docAuthor="Montoya-Molina, Santiago & Vaz-De, Fernando Z." docDate="2021" docId="03850669FF8EFFEEFE73274D3F8ACF45" docLanguage="en" docName="ejt-734_1-64.pdf" docOrigin="European Journal of Taxonomy 734" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius (Luederwaldtinia) henripittieri Montoya-Molina & Vaz-De 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="33" masterDocId="FFBC7E11FF90FFCEFFAD26453B59CB56" masterDocTitle="Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="31" updateTime="1698929631379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
|
||
<mods:mods id="5F5182AA45025D0ADC6372AC891F897A" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="0C1F5C3BAFE8E46BA1E9E66EC3222A90">
|
||
<mods:title id="A46D510821F1398820589B8663342048">Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6831A93835C4B33DB8D8CF831EFACF43" type="personal">
|
||
<mods:role id="E5F102A75521976F2A78140863096041">
|
||
<mods:roleTerm id="7F34F8AC14FABF8EBDDB4A438B69B755">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="23D67E696D9E0B1F75DC214CA41127BE">Montoya-Molina, Santiago</mods:namePart>
|
||
<mods:nameIdentifier id="855798F2E01044864CFD16CA7A117783" type="LSID">BC00697E-4F30-497A-A9C3-0C8E7B11B625</mods:nameIdentifier>
|
||
<mods:affiliation id="5A74CCBDE2ECCC3C1265C38A636F3A54">Federal University of Mato Grosso, Institute of Biosciences, Programa de Pós Graduaç „ o em Ecología e Conservaç „ o da Biodiversidade, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil. Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, Praha-Suchdol, 16500, Czech Republic. Federal University of Mato Grosso, Institute of Biosciences, Department of Biology and Zoology, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil.</mods:affiliation>
|
||
<mods:nameIdentifier id="4A83C0312266D71F2CB40FDF28E74756" type="email">horomo@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="BF6C4B4438464C9B6400C5F2EA0C11D6" type="personal">
|
||
<mods:role id="2C436A7381FE59CFD7BADA65A97EF909">
|
||
<mods:roleTerm id="4DC65199A4289EED7054ADE56C42375B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="895C295C1C15747A6FDB7C9B7C57F5EF">Vaz-De, Fernando Z.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D643D83EF93726323FED2A8D6C202D4D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="8C18968A799029A93EB69E56ED4C6868" type="host">
|
||
<mods:titleInfo id="08E3F2A8CD7346F417D7E5E36B393377">
|
||
<mods:title id="60495994B83A61FE297173099E0FFC4C">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="2CA02D791CC5133C90AAD815F90E6E19">
|
||
<mods:date id="29F8F2B6C5DD86F4BC668EC6EBBCDEE2">2021</mods:date>
|
||
<mods:detail id="7A5C21D2E3DC2A52D0127E486B0938FC" type="pubDate">
|
||
<mods:number id="51D4BB15CA2885E0B3BB27111C7DE75B">2021-02-11</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="5002CED57062454269C495CDCB0EAF95" type="volume">
|
||
<mods:number id="FC4ACBB11C6AD43225FA80A7824410D1">734</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="B0E28A4063802920ACE83CAEC9A0B31C" unit="page">
|
||
<mods:start id="C054BBF5622A4E00B58F8B19FF9BAEC3">1</mods:start>
|
||
<mods:end id="C8319935A3BAF81562FA98A4C094F7B7">64</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F30468EDB38307D46E46EB589B840520">journal article</mods:classification>
|
||
<mods:identifier id="62D3C21DBB1F1A29451D656E1362E19C" type="CLB-Dataset">8176</mods:identifier>
|
||
<mods:identifier id="1CB43622E2636CB833B8F62011702CCE" type="DOI">10.5852/ejt.2021.734.1233</mods:identifier>
|
||
<mods:identifier id="954368BC5ECB36DCACE40B947A003B40" type="GBIF-Dataset">fd8f5cb9-13e9-4c90-ac15-502fa7ee8d38</mods:identifier>
|
||
<mods:identifier id="6CF9791741944190E5AB5B458A476DB9" type="ISSN">2118-9773</mods:identifier>
|
||
<mods:identifier id="02C005581CBDF877458B2080035C0D87" type="Zenodo-Dep">4542107</mods:identifier>
|
||
<mods:identifier id="423F006A88A266AF9F6782E6EB877763" type="ZooBank">663A0C2D-E3CF-4B0C-8BA1-566B174829A2</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03850669FF8EFFEEFE73274D3F8ACF45" ID-DOI="http://doi.org/10.5281/zenodo.4545710" ID-GBIF-Taxon="177420814" ID-Zenodo-Dep="4545710" LSID="urn:lsid:plazi:treatment:03850669FF8EFFEEFE73274D3F8ACF45" httpUri="http://treatment.plazi.org/id/03850669FF8EFFEEFE73274D3F8ACF45" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
|
||
<subSubSection id="C336E4F4FF8EFFD0FE73274D3F0CCA75" box="[478,1109,264,291]" pageId="30" pageNumber="31" type="nomenclature">
|
||
<paragraph id="8B93B77FFF8EFFD0FE73274D3F0CCA75" blockId="30.[355,1232,264,360]" box="[478,1109,264,291]" pageId="30" pageNumber="31">
|
||
<heading id="D0DB0013FF8EFFD0FE73274D3F0CCA75" box="[478,1109,264,291]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
|
||
<taxonomicName id="4C2CCCFCFF8EFFD0FE73274D38AACA75" ID-CoL="7VQMS" authority="Montoya-Molina & Vaz-De, 2021" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" box="[478,1011,264,291]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="henripittieri" status="sp. nov." subGenus="Luederwaldtinia">
|
||
<emphasis id="B9586B6DFF8EFFD0FE73274D3920CA75" bold="true" box="[478,633,264,291]" italics="true" pageId="30" pageNumber="31">Dichotomius</emphasis>
|
||
(
|
||
<emphasis id="B9586B6DFF8EFFD0FD27274D3808CA75" bold="true" box="[650,849,264,291]" italics="true" pageId="30" pageNumber="31">Luederwaldtinia</emphasis>
|
||
)
|
||
<emphasis id="B9586B6DFF8EFFD0FCCF274D38AACA75" bold="true" box="[866,1011,264,291]" italics="true" pageId="30" pageNumber="31">henripittieri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26BD616FF8EFFD0FC56274C3F0CCA75" box="[1019,1109,265,291]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336E4F4FF8EFFD0FECE276E382ECA31" pageId="30" pageNumber="31" type="description">
|
||
<paragraph id="8B93B77FFF8EFFD0FECE276E3F89CA10" blockId="30.[355,1232,264,360]" box="[355,1232,299,326]" pageId="30" pageNumber="31">
|
||
<uri id="FFBDBB7DFF8EFFD0FECE276E3F89CA10" box="[355,1232,299,326]" pageId="30" pageNumber="31">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FF8A8DAAFF8EFFD0FD2B276E3F89CA10" box="[646,1232,299,326]" pageId="30" pageNumber="31">CB39A85C-47A6-4EB8-B538-6D4CEED8BC63</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FD112708382ECA31" blockId="30.[355,1232,264,360]" box="[700,887,333,360]" pageId="30" pageNumber="31">
|
||
<figureCitation id="1317ABFAFF8EFFD0FD112708384BCA31" box="[700,786,333,359]" captionStart="Fig" captionStartId="31.[188,231,1890,1916]" captionTargetBox="[216,1380,274,1852]" captionTargetId="figure-77@31.[208,1380,265,1859]" captionTargetPageId="31" captionText="Fig. 10. Dichotomius henripittieri sp. nov. A–B. Dorsal habitus. A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head, cephalic carina medial tubercle rounded at the apex. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth." figureDoi="http://doi.org/10.5281/zenodo.4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="30" pageNumber="31">Fig. 10</figureCitation>
|
||
, Map 11
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336E4F4FF8EFFD0FF1027CB39B9C9D4" pageId="30" pageNumber="31" type="diagnosis">
|
||
<paragraph id="8B93B77FFF8EFFD0FF1027CB3A6FCAFE" blockId="30.[189,310,398,424]" box="[189,310,398,424]" pageId="30" pageNumber="31">
|
||
<heading id="D0DB0013FF8EFFD0FF1027CB3A6FCAFE" bold="true" box="[189,310,398,424]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B9586B6DFF8EFFD0FF1027CB3A6FCAFE" bold="true" box="[189,310,398,424]" pageId="30" pageNumber="31">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF1027F939B9C9D4" blockId="30.[189,1400,444,642]" pageId="30" pageNumber="31">
|
||
<taxonomicName id="4C2CCCFCFF8EFFD0FF1027F93AB1CA80" authority="Montoya-Molina & Vaz-De, 2021" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" box="[189,488,444,470]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="henripittieri" status="sp. nov.">
|
||
<emphasis id="B9586B6DFF8EFFD0FF1027F93AB1CA80" box="[189,488,444,470]" italics="true" pageId="30" pageNumber="31">Dichotomius henripittieri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26BD616FF8EFFD0FE4227F93913CA80" box="[495,586,444,470]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from other species of the group by the following combination of characters: clypeal teeth short and rounded. Clypeo-genal junction not emarginate. Cephalic carina, wide as high, median tubercle rounded at apex. Dense, ocellate punctures covering interocular surface. Pronotum twice as wide as long. Anterior medial disk with weak declivity, not truncate. Impressed fine punctures restricted to antero-lateral edges. Row of uninterrupted ocellate punctures along pronotal margin. Interstriae microsculpture shagreened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336E4F4FF8EFFD0FF1024EC3E2EC9A6" box="[189,1399,681,753]" pageId="30" pageNumber="31" type="etymology">
|
||
<paragraph id="8B93B77FFF8EFFD0FF1024EC3A1DC995" blockId="30.[189,324,681,707]" box="[189,324,681,707]" pageId="30" pageNumber="31">
|
||
<heading id="D0DB0013FF8EFFD0FF1024EC3A1DC995" bold="true" box="[189,324,681,707]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B9586B6DFF8EFFD0FF1024EC3A1DC995" bold="true" box="[189,324,681,707]" pageId="30" pageNumber="31">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF1024923E2EC9A6" blockId="30.[189,1399,726,753]" box="[189,1399,726,753]" pageId="30" pageNumber="31">
|
||
Name based on the locality where this new species was collected, Henri Pittier National Park,
|
||
<collectingCountry id="F33BF7EFFF8EFFD0FB5A24933E2DC9A6" box="[1271,1396,726,752]" name="Venezuela" pageId="30" pageNumber="31">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336E4F4FF8EFFD0FF1025523FF5CF7F" pageId="30" pageNumber="31" type="materials_examined">
|
||
<paragraph id="8B93B77FFF8EFFD0FF1025523AF3C867" blockId="30.[189,426,791,817]" box="[189,426,791,817]" pageId="30" pageNumber="31">
|
||
<heading id="D0DB0013FF8EFFD0FF1025523AF3C867" bold="true" box="[189,426,791,817]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B9586B6DFF8EFFD0FF1025523AF3C867" bold="true" box="[189,426,791,817]" pageId="30" pageNumber="31">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF4725003A4DC890" blockId="30.[189,1399,837,966]" pageId="30" pageNumber="31">
|
||
<materialsCitation id="3B44BD22FF8EFFD0FF4725003A4DC890" ID-GBIF-Occurrence="3037418527" collectingDate="2010-08-18" collectingDateMax="2010-08-20" collectingDateMin="2010-08-18" collectingMethod="human faeces" collectionCode="CEMT" collectorName="R. Acconcia & J. Gamez" country="Venezuela" elevation="1145" latitude="10.3475" location="Mario Briceno Iragorry" longLatPrecision="21" longitude="-67.68806" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Aragua" typeStatus="holotype">
|
||
<emphasis id="B9586B6DFF8EFFD0FF4725003A05C809" bold="true" box="[234,348,837,863]" pageId="30" pageNumber="31">
|
||
<typeStatus id="549709DDFF8EFFD0FF4725003A05C809" box="[234,348,837,863]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33BF7EFFF8EFFD0FF1025223A32C8D7" box="[189,363,871,897]" name="Venezuela" pageId="30" pageNumber="31">VENEZUELA</collectingCountry>
|
||
•
|
||
<emphasis id="B9586B6DFF8EFFD0FE2C25223ACEC8D7" bold="true" box="[385,407,871,897]" pageId="30" pageNumber="31">
|
||
<specimenCount id="9D2A7CF6FF8EFFD0FE2C25223ACEC8D7" box="[385,407,871,897]" pageId="30" pageNumber="31" type="male">♂</specimenCount>
|
||
</emphasis>
|
||
;
|
||
<collectingRegion id="49E8799DFF8EFFD0FE0925223AA4C8D7" box="[420,509,871,897]" country="Venezuela" name="Aragua" pageId="30" pageNumber="31">Aragua</collectingRegion>
|
||
,
|
||
<location id="8EF3E1A4FF8EFFD0FDA525223843C8D4" LSID="urn:lsid:plazi:treatment:03850669FF8EFFEEFE73274D3F8ACF45:8EF3E1A4FF8EFFD0FDA525223843C8D4" box="[520,794,871,898]" country="Venezuela" latitude="10.3475" longLatPrecision="21" longitude="-67.68806" name="Mario Briceno Iragorry" pageId="30" pageNumber="31" stateProvince="Aragua">Mario Briceño Iragorry</location>
|
||
,
|
||
<location id="8EF3E1A4FF8EFFD0FC88252D38B2C8D7" LSID="urn:lsid:plazi:treatment:03850669FF8EFFEEFE73274D3F8ACF45:8EF3E1A4FF8EFFD0FC88252D38B2C8D7" box="[805,1003,871,898]" country="Venezuela" latitude="10.3475" longLatPrecision="21" longitude="-67.68806" name="PN. Henri Pittier" pageId="30" pageNumber="31" stateProvince="Aragua">PN. Henri Pittier</location>
|
||
,
|
||
<location id="8EF3E1A4FF8EFFD0FC58252D3F89C8D7" LSID="urn:lsid:plazi:treatment:03850669FF8EFFEEFE73274D3F8ACF45:8EF3E1A4FF8EFFD0FC58252D3F89C8D7" box="[1013,1232,871,898]" country="Venezuela" latitude="10.3475" longLatPrecision="21" longitude="-67.68806" name="Est. Bio. Periquito" pageId="30" pageNumber="31" stateProvince="Aragua">Est. Bio. Periquito</location>
|
||
;
|
||
<elevation id="0001504CFF8EFFD0FB7625223E29C8D7" box="[1243,1392,871,898]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="30" pageNumber="31" unit="m" value="1145.0">
|
||
<quantity id="4CD41A9AFF8EFFD0FB7625223E6BC8D4" box="[1243,1330,871,898]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="30" pageNumber="31" unit="m" value="1145.0">1145 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<geoCoordinate id="EE18D1B8FF8EFFD0FF1025CD3A09C8F2" box="[189,336,904,932]" degrees="10" direction="north" minutes="20" orientation="latitude" pageId="30" pageNumber="31" precision="15" seconds="51" value="10.3475">10º20′51″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE18D1B8FF8EFFD0FEF125CD3AACC8F2" box="[348,501,904,932]" degrees="67" direction="west" minutes="41" orientation="longitude" pageId="30" pageNumber="31" precision="15" seconds="17" value="-67.68806">67º41′17″ W</geoCoordinate>
|
||
;
|
||
<collectingMethod id="526DCF68FF8EFFD0FDAC25CC39FCC8F5" box="[513,677,905,931]" pageId="30" pageNumber="31">human faeces</collectingMethod>
|
||
;
|
||
<date id="FF9291BFFF8EFFD0FD1C25CF38D9C8F2" box="[689,896,906,932]" pageId="30" pageNumber="31" value="2010-08-18" valueMax="2010-08-20" valueMin="2010-08-18">
|
||
<collectingDate id="EFD66857FF8EFFD0FD1C25CF38D9C8F2" box="[689,896,906,932]" pageId="30" pageNumber="31" value="2010-08-18" valueMax="2010-08-20" valueMin="2010-08-18">18–20 Aug. 2010</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D9D2A9FF8EFFD0FC2125CF3F45C8F5" box="[908,1052,905,932]" pageId="30" pageNumber="31">R. Acconcia</collectorName>
|
||
and
|
||
<collectorName id="26D9D2A9FF8EFFD0FBF825CF3F9BC8F2" box="[1109,1218,906,932]" pageId="30" pageNumber="31">J. Gámez</collectorName>
|
||
leg. [white, p];
|
||
<collectionCode id="ED3D2FBAFF8EFFD0FF1025E93A49C890" box="[189,272,940,966]" pageId="30" pageNumber="31">CEMT</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF4725A83FF5CF7F" blockId="30.[189,1196,1005,1066]" pageId="30" pageNumber="31">
|
||
<materialsCitation id="3B44BD22FF8EFFD0FF4725A83FFECF7F" ID-GBIF-Occurrence="3037418425" collectionCode="CEMT" country="Venezuela" location="VENEZUELA" pageId="30" pageNumber="31" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="paratype">
|
||
<emphasis id="B9586B6DFF8EFFD0FF4725A83A3ECF51" bold="true" box="[234,359,1005,1031]" pageId="30" pageNumber="31">
|
||
<typeStatus id="549709DDFF8EFFD0FF4725A83A3ECF51" box="[234,359,1005,1031]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33BF7EFFF8EFFD0FF10224A3A32CF7F" box="[189,363,1039,1065]" name="Venezuela" pageId="30" pageNumber="31">VENEZUELA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D2A7CF6FF8EFFD0FE2E224A3A9FCF7F" box="[387,454,1037,1065]" pageId="30" pageNumber="31" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D2A7CF6FF8EFFD0FE7E224A394FCF7C" box="[467,534,1039,1066]" pageId="30" pageNumber="31" type="female">3 ♀♀</specimenCount>
|
||
; same collection data as for holotpye [white, p];
|
||
<collectionCode id="ED3D2FBAFF8EFFD0FBF9224A3FFECF7F" box="[1108,1191,1039,1065]" collectionName="CEMT" pageId="30" pageNumber="31">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336E4F4FF8EFFEEFF10220A3FDDC8D5" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
|
||
<paragraph id="8B93B77FFF8EFFD0FF10220A3A17CF3F" blockId="30.[189,334,1103,1129]" box="[189,334,1103,1129]" pageId="30" pageNumber="31">
|
||
<heading id="D0DB0013FF8EFFD0FF10220A3A17CF3F" bold="true" box="[189,334,1103,1129]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
|
||
<emphasis id="B9586B6DFF8EFFD0FF10220A3A17CF3F" bold="true" box="[189,334,1103,1129]" pageId="30" pageNumber="31">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF47223B3A73CFCE" blockId="30.[189,605,1150,1210]" box="[234,298,1150,1176]" pageId="30" pageNumber="31">
|
||
<emphasis id="B9586B6DFF8EFFD0FF47223B3A73CFCE" bold="true" box="[234,298,1150,1176]" pageId="30" pageNumber="31">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF1022E53904CFEC" blockId="30.[189,605,1150,1210]" box="[189,605,1184,1210]" pageId="30" pageNumber="31">
|
||
MEASUREMENTS. Length:
|
||
<quantity id="4CD41A9AFF8EFFD0FE7322E53901CFEC" box="[478,600,1184,1210]" metricMagnitude="-2" metricUnit="m" metricValue="1.936" pageId="30" pageNumber="31" unit="mm" value="19.36">19.36 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF1022A43AF2CEBA" blockId="30.[188,1400,1248,1516]" pageId="30" pageNumber="31">
|
||
HEAD. Wider than long. Anterior edge surface weakly rugose, extending posteriorly not reaching base of cephalic carina. Clypeo-genal surface covered by ocellate punctures spaced by one time diameter of a puncture. Interocular surface with ocellate punctures next to eyes (
|
||
<figureCitation id="1317ABFAFF8EFFD0FC5423603FD7CE69" box="[1017,1166,1317,1343]" captionStart="Fig" captionStartId="31.[188,231,1890,1916]" captionTargetBox="[216,1380,274,1852]" captionTargetId="figure-77@31.[208,1380,265,1859]" captionTargetPageId="31" captionText="Fig. 10. Dichotomius henripittieri sp. nov. A–B. Dorsal habitus. A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head, cephalic carina medial tubercle rounded at the apex. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth." figureDoi="http://doi.org/10.5281/zenodo.4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="30" pageNumber="31">Fig. 10A, C</figureCitation>
|
||
). Clypeus rounded, weak central emargination separating two rounded clypeal teeth. Ventral clypeal process longitudinally enlarged, cylindrical (see
|
||
<figureCitation id="1317ABFAFF8EFFD0FE51232F3906CED2" box="[508,607,1386,1412]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="30" pageNumber="31">Fig. 2D</figureCitation>
|
||
). Cephalic carina as wide as high, posterior surface shagreened microsculpture with ocellate shallow punctation. Median tubercle three times as high as lateral tubercles and curved posteriorly, apex rounded never bifurcate. Lateral tubercles obtuse at apex, vertically flattened (
|
||
<figureCitation id="1317ABFAFF8EFFD0FE9F23943AC7CEBA" box="[306,414,1489,1516]" captionStart="Fig" captionStartId="31.[188,231,1890,1916]" captionTargetBox="[216,1380,274,1852]" captionTargetId="figure-77@31.[208,1380,265,1859]" captionTargetPageId="31" captionText="Fig. 10. Dichotomius henripittieri sp. nov. A–B. Dorsal habitus. A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head, cephalic carina medial tubercle rounded at the apex. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth." figureDoi="http://doi.org/10.5281/zenodo.4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="30" pageNumber="31">Fig. 10C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF1020563F91CD8E" blockId="30.[189,1400,1554,1753]" pageId="30" pageNumber="31">
|
||
PRONOTUM. Twice as wide as long, and as wide as elytra. Anterior medial disc with weak declivity, not truncate. Median disc shiny and smooth, lacking deep punctures and having at most fine punctures. Antero-lateral surface with dense well-impressed simple punctures, scattered ocellate punctures also visible. Row of ocellate punctures along margins, punctures denser at anterior and posterior margins edges. Hypomeron medial disc punctation separated by three times width of a puncture, never smooth or impunctate. External margin bearing single row of organized long setae (see
|
||
<figureCitation id="1317ABFAFF8EFFD0FBCE20FB3FE2CD8E" box="[1123,1211,1726,1752]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="30" pageNumber="31">Fig. 2F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF8EFFD0FF1020BA3F5FCCBE" blockId="30.[188,1399,1791,2025]" pageId="30" pageNumber="31">
|
||
ABDOMEN. Prosternum and mesosternum with ocellate setigerous punctures. Metepisternum surface shagreened, displaying ocellate setigerous punctures, pilosity sparce. Metasternum setigerous punctures restricted mainly to lateral and anterior areas of mestasternal lobe, puncture diameter variable, medial disc smooth. Margins of mesocoxa surrounded by multiple ocellate punctures, reaching lateral portions of metasternum (
|
||
<figureCitation id="1317ABFAFF8EFFD0FE2F21CD3AB7CCF5" box="[386,494,1928,1955]" captionStart="Fig" captionStartId="31.[188,231,1890,1916]" captionTargetBox="[216,1380,274,1852]" captionTargetId="figure-77@31.[208,1380,265,1859]" captionTargetPageId="31" captionText="Fig. 10. Dichotomius henripittieri sp. nov. A–B. Dorsal habitus. A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head, cephalic carina medial tubercle rounded at the apex. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth." figureDoi="http://doi.org/10.5281/zenodo.4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="30" pageNumber="31">Fig. 10D</figureCitation>
|
||
). Lateral surface of metasternum, with ocellate setigerous punctures separated by three times width of a puncture. Ocellate punctures along sternites anterior margins. Pygidium wider than long, basal margin rugose, bearing ocellate punctures in basal half.
|
||
</paragraph>
|
||
<caption id="DF53E7F7FF8FFFD1FF1121273F79CCB0" ID-DOI="http://doi.org/10.5281/zenodo.4542127" ID-Zenodo-Dep="4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="31" pageNumber="32" startId="31.[188,231,1890,1916]" targetBox="[216,1380,274,1852]" targetPageId="31">
|
||
<paragraph id="8B93B77FFF8FFFD1FF1121273F79CCB0" blockId="31.[187,1398,1888,2022]" pageId="31" pageNumber="32">
|
||
<emphasis id="B9586B6DFF8FFFD1FF1121273A47CC2A" bold="true" box="[188,286,1890,1916]" pageId="31" pageNumber="32">Fig. 10.</emphasis>
|
||
<taxonomicName id="4C2CCCFCFF8FFFD1FE8121273904CC2A" authority="Montoya-Molina & Vaz-De, 2021" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" box="[300,605,1890,1916]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="henripittieri" status="sp. nov.">
|
||
<emphasis id="B9586B6DFF8FFFD1FE8121273904CC2A" box="[300,605,1890,1916]" italics="true" pageId="31" pageNumber="32">Dichotomius henripittieri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26BD616FF8FFFD1FDC721263992CC2B" box="[618,715,1891,1917]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B9586B6DFF8FFFD1FD7B21273854CC2A" bold="true" box="[726,781,1890,1916]" pageId="31" pageNumber="32">A–B</emphasis>
|
||
. Dorsal habitus.
|
||
<emphasis id="B9586B6DFF8FFFD1FC4A212738A5CC2A" bold="true" box="[999,1020,1890,1916]" pageId="31" pageNumber="32">A</emphasis>
|
||
. Holotype, ♂ (CEMT).
|
||
<emphasis id="B9586B6DFF8FFFD1FA9D21273E1DCC2A" bold="true" box="[1328,1348,1890,1916]" pageId="31" pageNumber="32">B</emphasis>
|
||
. ♀.
|
||
<emphasis id="B9586B6DFF8FFFD1FF1121C03B88CCC9" bold="true" box="[188,209,1925,1951]" pageId="31" pageNumber="32">C</emphasis>
|
||
. Lateral view of the aedeagus.
|
||
<emphasis id="B9586B6DFF8FFFD1FD9721C0390ECCC9" bold="true" box="[570,599,1925,1951]" pageId="31" pageNumber="32">D.</emphasis>
|
||
Frontal view of the head, cephalic carina medial tubercle rounded at the apex.
|
||
<emphasis id="B9586B6DFF8FFFD1FE8621EC3A66CC95" bold="true" box="[299,319,1961,1987]" pageId="31" pageNumber="32">E</emphasis>
|
||
. Dorsal view of the aedeagus.
|
||
<emphasis id="B9586B6DFF8FFFD1FD3221ED398FCC94" bold="true" box="[671,726,1960,1986]" pageId="31" pageNumber="32">F–G</emphasis>
|
||
. Lamella copulatrix.
|
||
<emphasis id="B9586B6DFF8FFFD1FC7D21EC38BECC95" bold="true" box="[976,999,1961,1987]" pageId="31" pageNumber="32">H</emphasis>
|
||
. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B93B77FFFB0FFEEFF10274C3E2ECA13" blockId="32.[189,1399,265,326]" pageId="32" pageNumber="33">
|
||
ELYTRA. Interstriae microsculpture shagreened, with smooth fine scattered punctures. Striae wellimpressed with ocellate punctures separated by less the diameter of a puncture on elytral disc (
|
||
<figureCitation id="1317ABFAFFB0FFEEFB50276E3E30CA13" box="[1277,1385,299,325]" captionStart="Fig" captionStartId="31.[188,231,1890,1916]" captionTargetBox="[216,1380,274,1852]" captionTargetId="figure-77@31.[208,1380,265,1859]" captionTargetPageId="31" captionText="Fig. 10. Dichotomius henripittieri sp. nov. A–B. Dorsal habitus. A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head, cephalic carina medial tubercle rounded at the apex. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth." figureDoi="http://doi.org/10.5281/zenodo.4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="32" pageNumber="33">Fig. 10A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFB0FFEEFF1027293A11CA9B" blockId="32.[189,1398,364,461]" pageId="32" pageNumber="33">LEGS. Metacoxa posterior edge bearing single row of disorganized ocellate setigerous punctures, separated by twice diameter of a puncture. Profemora displaying aggregation of setigerous punctures on distal ¾.</paragraph>
|
||
<paragraph id="8B93B77FFFB0FFEEFF1027B4381CC95D" blockId="32.[189,837,497,523]" box="[189,837,497,523]" pageId="32" pageNumber="33">
|
||
PARAMERES AND LAMELLA COPULATRIX. As in
|
||
<figureCitation id="1317ABFAFFB0FFEEFD1827B4385FC95D" box="[693,774,497,523]" captionStart="Fig" captionStartId="31.[188,231,1890,1916]" captionTargetBox="[216,1380,274,1852]" captionTargetId="figure-77@31.[208,1380,265,1859]" captionTargetPageId="31" captionText="Fig. 10. Dichotomius henripittieri sp. nov. A–B. Dorsal habitus. A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head, cephalic carina medial tubercle rounded at the apex. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth." figureDoi="http://doi.org/10.5281/zenodo.4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="32" pageNumber="33">Fig. 10</figureCitation>
|
||
E–G.
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFB0FFEEFF1024743AB7C91D" blockId="32.[189,494,561,587]" box="[189,494,561,587]" pageId="32" pageNumber="33">
|
||
<heading id="D0DB0013FFB0FFEEFF1024743AB7C91D" bold="true" box="[189,494,561,587]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
|
||
<emphasis id="B9586B6DFFB0FFEEFF1024743AB7C91D" bold="true" box="[189,494,561,587]" pageId="32" pageNumber="33">Morphological variation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFB0FFEEFF47241A3A8AC92F" blockId="32.[189,722,607,668]" box="[234,467,607,633]" pageId="32" pageNumber="33">
|
||
<emphasis id="B9586B6DFFB0FFEEFF47241A3A08C92F" bold="true" box="[234,337,607,633]" pageId="32" pageNumber="33">Females</emphasis>
|
||
(
|
||
<figureCitation id="1317ABFAFFB0FFEEFECC241A3A93C92F" box="[353,458,607,633]" captionStart="Fig" captionStartId="31.[188,231,1890,1916]" captionTargetBox="[216,1380,274,1852]" captionTargetId="figure-77@31.[208,1380,265,1859]" captionTargetPageId="31" captionText="Fig. 10. Dichotomius henripittieri sp. nov. A–B. Dorsal habitus. A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head, cephalic carina medial tubercle rounded at the apex. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth." figureDoi="http://doi.org/10.5281/zenodo.4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="32" pageNumber="33">Fig. 10B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFB0FFEEFF1024C4398BC9CD" blockId="32.[189,722,607,668]" box="[189,722,641,668]" pageId="32" pageNumber="33">As males except for the following characters:</paragraph>
|
||
<paragraph id="8B93B77FFFB0FFEEFF1024873AC9C876" blockId="32.[188,1399,706,800]" pageId="32" pageNumber="33">
|
||
HEAD. Completely transversely rugose, denser on anterior portion. Clypeus with strong central emargination separating two acute clypeal teeth. Cephalic carina bituberculate, tubercles narrow, almost fading (
|
||
<figureCitation id="1317ABFAFFB0FFEEFEBA25433ADAC876" box="[279,387,774,800]" captionStart="Fig" captionStartId="31.[188,231,1890,1916]" captionTargetBox="[216,1380,274,1852]" captionTargetId="figure-77@31.[208,1380,265,1859]" captionTargetPageId="31" captionText="Fig. 10. Dichotomius henripittieri sp. nov. A–B. Dorsal habitus. A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head, cephalic carina medial tubercle rounded at the apex. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum, mesocoxa surrounded by multiple ocellate punctures, medial disc smooth." figureDoi="http://doi.org/10.5281/zenodo.4542127" httpUri="https://zenodo.org/record/4542127/files/figure.png" pageId="32" pageNumber="33">Fig. 10B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFB0FFEEFF1025023FDDC8D5" blockId="32.[189,1398,839,900]" pageId="32" pageNumber="33">PRONOTUM. Surface covered by fine punctures, denser on antero-lateral areas. Punctuation on margins ocellate and dense, discernible. Hypomeron setae longer and denser than in males.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336E4F4FFB0FFEEFF1025EF3F8ACF45" pageId="32" pageNumber="33" type="discussion">
|
||
<paragraph id="8B93B77FFFB0FFEEFF1025EF3A76C892" blockId="32.[189,303,938,964]" box="[189,303,938,964]" pageId="32" pageNumber="33">
|
||
<heading id="D0DB0013FFB0FFEEFF1025EF3A76C892" bold="true" box="[189,303,938,964]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
|
||
<emphasis id="B9586B6DFFB0FFEEFF1025EF3A76C892" bold="true" box="[189,303,938,964]" pageId="32" pageNumber="33">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFB0FFEEFF1025923F8ACF45" blockId="32.[189,1398,983,1044]" pageId="32" pageNumber="33">
|
||
This species was collected in the Henri Pittier National Park located on the ridge of the Cordillera de la Costa of
|
||
<collectingCountry id="F33BF7EFFFB0FFEEFE8B25BC3AFACF45" box="[294,419,1017,1043]" name="Venezuela" pageId="32" pageNumber="33">Venezuela</collectingCountry>
|
||
. The vegetation of the area is characterized by extensive cloud forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |