treatments-xml/data/83/80/AF/8380AF1104FE087B077EEE33E82D3F42.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.845.33566" ID-GBIF-Dataset="93d412c8-defe-4417-a5a4-f0b978d55e3f" ID-PMC="PMC6535459" ID-Pensoft-Pub="1313-2970-845-99" ID-PubMed="31156329" ID-ZBK="2C4523C4409749048797D03F55DD47D9" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-845-99" ModsDocOrigin="ZooKeys 845" ModsDocTitle="Two new species of the genus Symphylella (Symphyla, Scolopendrellidae) from Tibet, China" checkinTime="1557997603692" checkinUser="pensoft" docAuthor="Jin, Ya-Li, Bu, Yun &amp; Jiang, Yue" docDate="2019" docId="8380AF1104FE087B077EEE33E82D3F42" docLanguage="en" docName="ZooKeys 845: 99-117" docOrigin="ZooKeys 845" docSource="http://dx.doi.org/10.3897/zookeys.845.33566" docTitle="Symphylella macropora Jin &amp; Bu, sp. n." docType="treatment" docUuid="7AAA91E2-37AF-45CE-B14F-9433EBD3FF75" docUuidSource="ZooBank" docVersion="6" lastPageNumber="105" masterDocId="FFBECB349B6FFFC11442FF9DFFDCFFBE" masterDocTitle="Two new species of the genus Symphylella (Symphyla, Scolopendrellidae) from Tibet, China" masterLastPageNumber="117" masterPageNumber="99" pageNumber="101" updateTime="1668167255525" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of the genus Symphylella (Symphyla, Scolopendrellidae) from Tibet, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jin, Ya-Li</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bu, Yun</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Yue</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>845</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>99</mods:start>
<mods:end>117</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.845.33566</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.845.33566</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-845-99</mods:identifier>
<mods:identifier type="ZBK">2C4523C4409749048797D03F55DD47D9</mods:identifier>
<mods:identifier type="ZooBank">2C4523C4409749048797D03F55DD47D9</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="157167782" LSID="urn:lsid:zoobank.org:act:7AAA91E2-37AF-45CE-B14F-9433EBD3FF75" httpUri="http://treatment.plazi.org/id/8380AF1104FE087B077EEE33E82D3F42" lastPageId="7" lastPageNumber="105" pageId="2" pageNumber="101">
<subSubSection pageId="2" pageNumber="101" type="nomenclature">
<paragraph pageId="2" pageNumber="101">
<taxonomicName LSID="http://zoobank.org/7AAA91E2-37AF-45CE-B14F-9433EBD3FF75" authority="Jin &amp; Bu" class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella macropora" order="Symphyla" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="macropora">
<pageBreakToken pageId="2" pageNumber="101" start="start">Symphylella</pageBreakToken>
macropora Jin &amp; Bu
</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="101">sp. n.</taxonomicNameLabel>
Figures 1, 2, Tables 1, 2, 3
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="101" type="diagnosis">
<paragraph pageId="2" pageNumber="101">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="101">
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella macropora" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="macropora">Symphylella macropora</taxonomicName>
sp. n. is characterized by large, elongated oval openings of the 
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
organs, with their inner margins of these openings covered by minute irregular teeth, as well as by having rudimentary spined sensory organs on the dorsal side of most antennal segments, and cerci with numerous long and slightly curved setae.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="101" type="materials_examined">
<paragraph pageId="2" pageNumber="101">Material examined.</paragraph>
<paragraph pageId="2" pageNumber="101">
Holotype, female (slide no. XZ-SY2015030) (SNHM), China, Tibet, Motuo county, Dexing town, extracted from soil samples from broadleaf forest, alt. 1100 m, 
<geoCoordinate direction="north" orientation="latitude" precision="925" value="29.666666">29°40'N</geoCoordinate>
, 
<geoCoordinate direction="east" orientation="longitude" precision="925" value="95.433334">95°26'E</geoCoordinate>
, 3-XI-2015, coll. Y. Bu &amp; G. Yang. Paratypes, 6 females (slides nos. XZ-SY2015023-XZ-SY2015026, XZ-SY2015029, XZ-SY2015032) (SNHM), 3 males (slides nos. XZ-SY2015027, XZ-SY2015028, XZ-SY2015031) (SNHM), same date as holotype; 1 female (slide no. XZ-SY2015034) (SNHM), China, Tibet, Motuo county, Beibeng town, extracted from soil samples from broadleaf forest, alt. 1500 m, 
<geoCoordinate direction="north" orientation="latitude" precision="925" value="29.5">29°30'N</geoCoordinate>
, 
<geoCoordinate direction="east" orientation="longitude" precision="925" value="95.63333">95°38'E</geoCoordinate>
, 5-XI-2015, coll. Y. Bu &amp; G. Yang. Other materials (SNHM): 1 juvenile with 9 pairs of legs (slide no. XZ-SY2015033) (SNHM), China, Tibet, Motuo county, Beibeng town, extracted from soil samples from broadleaf forest, alt.1500 m, 
<geoCoordinate direction="north" orientation="latitude" precision="925" value="29.5">29°30'N</geoCoordinate>
, 
<geoCoordinate direction="east" orientation="longitude" precision="925" value="95.63333">95°38'E</geoCoordinate>
, 5-XI-2015, coll. Y. Bu &amp; G. Yang; 2 juveniles with 9-10 pairs of legs (slides nos. XZ-SY2015035-XZ-SY2015036) (SNHM), same date as holotype; 10 juveniles with 8-10 pairs of legs (slides nos. XZ-SY2015037-XZ-SY2015046) (SNHM), China, Tibet, Motuo county, Dexing town, Naerdong village, extracted from soil samples from broadleaf forest, alt. 1800 m, 
<geoCoordinate direction="north" orientation="latitude" precision="925" value="29.5">29°30'N</geoCoordinate>
, 
<geoCoordinate direction="east" orientation="longitude" precision="925" value="95.38333">95°23'E</geoCoordinate>
, 4-XI-2015, coll. Y. Bu.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="105" pageId="2" pageNumber="101" type="description">
<paragraph pageId="2" pageNumber="101">Description.</paragraph>
<paragraph pageId="2" pageNumber="101">Adult body 1.90 mm long in average (1.55-2.71 mm, n=11), holotype 1.89 mm (Fig. 1A).</paragraph>
<caption pageId="2" pageNumber="101">
<paragraph pageId="2" pageNumber="101">
Figure 1. 
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella macropora" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="macropora">Symphylella macropora</taxonomicName>
sp. n. A habitus B left antenna, 15 
<normalizedToken originalValue="th–">th-</normalizedToken>
18th segment, dorsal view (bo-bladder-shaped organ, co-cavity-shaped organ) C right 
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
organ D head, dorsal view E first pair of legs (arrows indicate the legs) F stylus and coxal sacs of leg 3 (arrows indicate styli) G 4th tergite, left side (al-anterolateral seta, as-apical seta, cs-central seta, ibs-inner basal seta, is-inserted seta, lms-lateromarginal seta) H 1 
<normalizedToken originalValue="st–">st-</normalizedToken>
3rd tergites I 4 
<normalizedToken originalValue="th–">th-</normalizedToken>
5th tergites J 13 
<normalizedToken originalValue="th–">th-</normalizedToken>
15th tergites. Scale bars: 100 
<normalizedToken originalValue="μm">μm</normalizedToken>
(A); 20 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<normalizedToken originalValue="B–J">B-J</normalizedToken>
).
</paragraph>
</caption>
<paragraph pageId="2" pageNumber="101">
Head length 210-270 
<normalizedToken originalValue="μm">μm</normalizedToken>
, width 225-300 
<normalizedToken originalValue="μm">μm</normalizedToken>
, with widest part somewhat behind the middle on a level with the points of articulation of mandibles. Central rod distinct in both anterior (50-70 
<normalizedToken originalValue="μm">μm</normalizedToken>
) and posterior (58-75 
<normalizedToken originalValue="μm">μm</normalizedToken>
) portions, with a middle knot. Anterior branches well developed, median branches vestigial. Head dorsally moderately covered with setae of different length, longest setae (25-35 
<normalizedToken originalValue="μm">μm</normalizedToken>
) located most anteriorly, at least 3.0 times as long as central ones (8-12 
<normalizedToken originalValue="μm">μm</normalizedToken>
). Cuticle at anterolateral part of head with rather coarse granulation. 
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
organ surrounded by fine granulation, other areas with fine dense granulation (Fig. 1D).
</paragraph>
<paragraph pageId="2" pageNumber="101">
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
organ oval, length 19-32 
<normalizedToken originalValue="μm">μm</normalizedToken>
, width 10-22 
<normalizedToken originalValue="μm">μm</normalizedToken>
, at least half of greatest diameter of 3rd antennal segment (35-50 
<normalizedToken originalValue="μm">μm</normalizedToken>
), openings large and elongated oval (length 13-26 
<normalizedToken originalValue="μm">μm</normalizedToken>
, width 5-10 
<normalizedToken originalValue="μm">μm</normalizedToken>
), with their inner margins covered by minute irregular teeth (Figs 1C, 1D).
</paragraph>
<paragraph pageId="2" pageNumber="101">Mouthparts Mandible with two fused lamellae and 11 teeth in total (Fig. 2A). First maxilla with two lobes, inner lobe with four hook-shaped teeth, palp bud-like with 1 distal point close to outer lobe (Fig. 2B). Anterior part of second maxilla with many small protuberances which carry one seta each, distal setae thickened; posterior part with sparse setae. Cuticle of maxilla and labium covered with pubescence.</paragraph>
<caption pageId="2" pageNumber="101">
<paragraph pageId="2" pageNumber="101">
Figure 2. 
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella macropora" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="macropora">Symphylella macropora</taxonomicName>
sp. n. A mandible B first maxilla 
<normalizedToken originalValue="C–D">C-D</normalizedToken>
15 
<normalizedToken originalValue="th–">th-</normalizedToken>
18th segments of left antenna C dorsal view (bo-bladder-shaped organ, co-cavity-shaped organ, rso-rudimentary spined sensory organ, so-spined sensory organ) D ventral view E 4 
<normalizedToken originalValue="th–">th-</normalizedToken>
5th segments of right antenna, dorsal view F 12th leg G stylus on base of 4th leg H right cercus, dorsal view. Scale bars: 20 
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<normalizedToken originalValue="A–F">A-F</normalizedToken>
, H); 5 
<normalizedToken originalValue="μm">μm</normalizedToken>
(G).
</paragraph>
</caption>
<paragraph lastPageId="4" lastPageNumber="103" pageId="2" pageNumber="101">
Antennae with 16-23 segments (18 in holotype), length 570-680 
<normalizedToken originalValue="μm">μm</normalizedToken>
(620 
<normalizedToken originalValue="μm">μm</normalizedToken>
in holotype), about 0.3 of body length. 1st segment cylindrical, much shorter than follow 
<pageBreakToken pageId="3" pageNumber="102" start="start">ing</pageBreakToken>
ones, greatest diameter wider than long (37-45 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 13-33 
<normalizedToken originalValue="μm">μm</normalizedToken>
), with 2-3 microsetae and 6-9 long mesosetae in one whorl, longest seta (20-24 
<normalizedToken originalValue="μm">μm</normalizedToken>
) inserted at inner side and distinctly longer than outer ones (15-18 
<normalizedToken originalValue="μm">μm</normalizedToken>
). 2nd segment wider (35-44 
<normalizedToken originalValue="μm">μm</normalizedToken>
) than long (28-35 
<normalizedToken originalValue="μm">μm</normalizedToken>
), with eight or nine setae evenly inserted around antennal wall, 
<pageBreakToken pageId="4" pageNumber="103" start="start">interior</pageBreakToken>
setae (23-26 
<normalizedToken originalValue="μm">μm</normalizedToken>
) slightly longer than exterior ones (16-19 
<normalizedToken originalValue="μm">μm</normalizedToken>
). Chaetotaxy of 3rd segment like preceding ones. Setae on basal segments slender and on distal segments shorter. Basal segments of antennae with only primary whorl of setae, in middle and subapical segments with several minute setae in secondary whorl. Three kinds of sensory organs on most antenna segments: rso on dorsal side of most segments (Figs 2C, 2E); co on dorsal side of 3 
<normalizedToken originalValue="rd–">rd-</normalizedToken>
21th segments; bo on subapical 5-9 segments (Figs 1B, 2C, 2D). Apical segment subspherical, somewhat wider than long (width 28-32 
<normalizedToken originalValue="μm">μm</normalizedToken>
, length 24-26 
<normalizedToken originalValue="μm">μm</normalizedToken>
), with 13-15 setae on distal half and 2 baculiform organs on apex, 4-7 so consisting of 3-4 curved spines around a central pillar in depressions in distal surface (Figs 2C, 2D). All segments covered with short pubescence. Chaetotaxy and sensory organs of antennae are given in table 1.
</paragraph>
<caption pageId="4" pageNumber="103">
<paragraph pageId="4" pageNumber="103">Table 1. Numbers of setae and sensory organs of antennae (holotype).</paragraph>
</caption>
<paragraph pageId="4" pageNumber="103">
<table pageId="4" pageNumber="103">
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="2">Segments</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="2">Nos. of primary whorl setae</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="2">Nos. of secondary whorl setae</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="2">Rudimentary spined sensory organs (rso)</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">Cavity-shaped organs (co)</td>
<td colspan="2" pageId="4" pageNumber="103" rowspan="1">Bladder-shaped organs (bo)</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">Dorsal</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">Dorsal</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">Ventral</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">st</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">nd</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">rd</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
</table>
</paragraph>
<paragraph pageId="4" pageNumber="103">
Trunk with 17 tergites. Most tergites overlap, with intertergal zones present between adjacent tergites, except for borders between 14th and 15th, as well as 16th and 17th tergite. Tergites 2-13, and 15 each with one pair of triangular processes. Basal distance between processes of 4 
<normalizedToken originalValue="th–">th-</normalizedToken>
13thand 15th tergites longer than their length from base to tip, and the latter shorter than its basal wide (Table 2). Anterolateral setae of 2nd, 3rd, 4th, 6th, 7th, 9th and10th tergites distinctly longer than other lateromarginal setae, that of 5th, 8th, 11 
<normalizedToken originalValue="th–">th-</normalizedToken>
13th and 15th subequal or slightly longer than other lateromarginal setae. Triangular processes without ovoid swollen ends (Fig. 1G). One to three inserted setae (setae between inner basal seta and apical seta). All tergites pubescent.
</paragraph>
<caption pageId="4" pageNumber="103">
<paragraph pageId="4" pageNumber="103">Table 2. Chaetotaxy of tergites (holotype in brackets).</paragraph>
</caption>
<paragraph pageId="4" pageNumber="103">
<table pageId="4" pageNumber="103">
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">No. of tergites</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">Lateromarginal setae</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">Inserted seta</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">Central setae</td>
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">Other setae</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">st</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">nd</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">rd</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
<tr pageId="4" pageNumber="103">
<td colspan="1" pageId="4" pageNumber="103" rowspan="1">th</td>
</tr>
</table>
</paragraph>
<paragraph lastPageId="5" lastPageNumber="104" pageId="4" pageNumber="103">
Tergites 1st tergite reduced and build up of two short plates separated in the median axis of the body, with 6-9 short setae in a row. 2nd tergite complete, with two posterior 
<pageBreakToken pageId="5" pageNumber="104" start="start">processes</pageBreakToken>
, 5-8 lateromarginal setae, 1-2 inserted setae, 1-4 central setae, with anterolateral setae distinctly longer than other lateromarginal setae, processes approximately 0.9 times as long as broad, basal distance between processes approximately 0.7 times as long as their length. 3rd tergite entire, broader and longer than preceding one with ratios mentioned nearly the same as for the 2nd tergite, 6-10 lateromarginal setae (Fig. 1H). 4th tergite broader than 3rd tergite, with ratios approximately 0.7 and 1.4 respectively, 5-7 lateromarginal setae (Fig. 1I). Shape and chaetotaxy of 5 
<normalizedToken originalValue="th–">th-</normalizedToken>
7th, 8 
<normalizedToken originalValue="th–">th-</normalizedToken>
10th, and 11 
<normalizedToken originalValue="th–">th-</normalizedToken>
13th tergite similar as 2 
<normalizedToken originalValue="nd–">nd-</normalizedToken>
4th tergite. 15th tergite shorter with smaller processes than remaining long tergites (3rd, 6th, 9th and 12th tergites). 14th and 16th tergites without processes, with 11-17 and 6-14 marginal setae respectively (Fig. 1J). 17th tergite with 20-31 setae. Chaetotaxy and measurements of tergites are given in Table 2, 3.
</paragraph>
<caption pageId="5" pageNumber="104">
<paragraph pageId="5" pageNumber="104">
Table 3. Measurements of tergites and processes (mean 
<normalizedToken originalValue="±">+/-</normalizedToken>
se, n = 11) (in 
<normalizedToken originalValue="μm">μm</normalizedToken>
).
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="104">
<table pageId="5" pageNumber="104">
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">No. of tergites</td>
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">Length</td>
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">Width</td>
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">Length of processes</td>
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">Basal width of processes</td>
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">Basal distance between processes</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">st</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">nd</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">rd</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
<tr pageId="5" pageNumber="104">
<td colspan="1" pageId="5" pageNumber="104" rowspan="1">th</td>
</tr>
</table>
</paragraph>
<paragraph lastPageId="6" lastPageNumber="105" pageId="5" pageNumber="104">
Legs 1st pair of legs reduced to two small hairy cupules, each with two long setae (Fig. 1E). Basal areas of legs 2-12 each with 4-6 setae (Fig. 1F). Leg 12 about 0.1 time as long as body, subequal length with head. Trochanter longer than wide (52-76 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 40-56 
<normalizedToken originalValue="μm">μm</normalizedToken>
) with 6-8 subequal setae. Femur as long as wide (32-42 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 30-41 
<normalizedToken originalValue="μm">μm</normalizedToken>
), with 4-6 setae and one of them (17-28 
<normalizedToken originalValue="μm">μm</normalizedToken>
) distinctly longer than others (10-20 
<normalizedToken originalValue="μm">μm</normalizedToken>
); pubescent dorsally, ventrally with cuticular thickenings in pattern of large scales. Tibia nearly 2 times longer than wide (40-60 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 23-30 
<normalizedToken originalValue="μm">μm</normalizedToken>
), with 5-7 dorsal setae and the longest one slightly shorter than greatest diameter of tibia (19-28 
<normalizedToken originalValue="μm">μm</normalizedToken>
).Tarsus subcylindrical, at least 3 times as long as wide (50-70 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 15-20 
<normalizedToken originalValue="μm">μm</normalizedToken>
), with 6-9 setae: 3 straight and protruding, 2 curved and depressed; longest setae (20-27 
<normalizedToken originalValue="μm">μm</normalizedToken>
) about same length of greatest width of podomere, and 2 ventral setae inserted near claw distinctly shorter than dorsal ones. Claws rather curved, anterior one distinctly longer and 
<pageBreakToken pageId="6" pageNumber="105" start="start">broader</pageBreakToken>
than posterior one, the latter more curved than the former (Fig. 2F). All legs covered with dense pubescence except areas with cuticular thickenings.
</paragraph>
<paragraph pageId="6" pageNumber="105">
Coxal sacs present at bases of 3 
<normalizedToken originalValue="rd–">rd-</normalizedToken>
9th pairs of legs, fully developed, each with 4 setae on its surface (Fig. 1F).
</paragraph>
<paragraph pageId="6" pageNumber="105">
Styli present at base of 3 
<normalizedToken originalValue="rd–">rd-</normalizedToken>
12th pairs of legs, subconical (length 6-9 
<normalizedToken originalValue="μm">μm</normalizedToken>
, width 4-6 
<normalizedToken originalValue="μm">μm</normalizedToken>
), basal part with straight hairs; distal one fourth hairless and blunt (3-6 
<normalizedToken originalValue="μm">μm</normalizedToken>
) (Figs 1F, 2G).
</paragraph>
<paragraph pageId="6" pageNumber="105">
<taxonomicName genus="Sense" lsidName="Sense calicles" pageId="6" pageNumber="105" rank="species" species="calicles">Sense calicles</taxonomicName>
with smooth margin to pit, length about the same as outer diameter (18-39 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 20-35 
<normalizedToken originalValue="μm">μm</normalizedToken>
). Sensory seta inserted in cup center, extremely long, length 130-165 
<normalizedToken originalValue="μm">μm</normalizedToken>
, at least 5.5 times longer than other two lateral setae (20-35 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 14-22 
<normalizedToken originalValue="μm">μm</normalizedToken>
respectively) that inserted at the edge of cup.
</paragraph>
<paragraph pageId="6" pageNumber="105">
Cerci subuliform, short, about 0.6 of head length and leg 12, length at least three times as long as its greatest width (126-172 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 40-53 
<normalizedToken originalValue="μm">μm</normalizedToken>
), moderately covered with subequal length and slightly curved setae, with longest (20-40 
<normalizedToken originalValue="μm">μm</normalizedToken>
) at least half of the greatest width of the cerci, terminal area (23-32 
<normalizedToken originalValue="μm">μm</normalizedToken>
) short, circled by 6-8 layers of curved ridges. Terminal setae length 18-28 
<normalizedToken originalValue="μm">μm</normalizedToken>
(Fig. 2H).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="105" type="etymology">
<paragraph pageId="6" pageNumber="105">Etymology.</paragraph>
<paragraph pageId="6" pageNumber="105">
The species name macropora is referring to the large opening of the 
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
organ.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="105" type="distribution">
<paragraph pageId="6" pageNumber="105">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="105">Known only from the type locality.</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="106" pageId="6" pageNumber="105" type="remarks">
<paragraph pageId="6" pageNumber="105">Remarks.</paragraph>
<paragraph lastPageId="7" lastPageNumber="106" pageId="6" pageNumber="105">
<taxonomicName class="Symphyla" family="Scolopendrellidae" genus="Symphylella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Symphylella macropora" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="macropora">Symphylella macropora</taxonomicName>
sp. n. is most similar to 
<taxonomicName lsidName="S. javanensis" pageId="6" pageNumber="105" rank="species" species="javanensis">S. javanensis</taxonomicName>
Scheller, 1988 from Java in the shape of tergites and processes, leg 12 and sensory organs on antennae. However, it can be readily distinguished from the latter by the shape of 
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
organ (oval, openings large and elongated with inner margins covered by minute irregular teeth 
<pageBreakToken pageId="7" pageNumber="106" start="start">vs</pageBreakToken>
subspherical, diameter of opening about half of the organ in 
<taxonomicName lsidName="S. javanensis" pageId="7" pageNumber="106" rank="species" species="javanensis">S. javanensis</taxonomicName>
), central rod (both anterior and posterior portions distinct in 
<taxonomicName lsidName="S. macropora" pageId="7" pageNumber="106" rank="species" species="macropora">S. macropora</taxonomicName>
sp. n. vs anterior half and anterior branches very thin with traces in 
<taxonomicName lsidName="S. javanensis" pageId="7" pageNumber="106" rank="species" species="javanensis">S. javanensis</taxonomicName>
), and the stylus (with blunt apex in 
<taxonomicName lsidName="S. macropora" pageId="7" pageNumber="106" rank="species" species="macropora">S. macropora</taxonomicName>
sp. n. vs with truncate apex in 
<taxonomicName lsidName="S. javanensis" pageId="7" pageNumber="106" rank="species" species="javanensis">S. javanensis</taxonomicName>
). It is also closely related to 
<taxonomicName lsidName="S. asiatica" pageId="7" pageNumber="106" rank="species" species="asiatica">S. asiatica</taxonomicName>
Scheller, 1971 from Indiaand Sri Lanka in the shape and chaetotaxy of tergites 1-3, leg 12 and the sensory organs on antenna, but easily distinguished by characters of 
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
organ (openings very small in 
<taxonomicName lsidName="S. asiatica" pageId="7" pageNumber="106" rank="species" species="asiatica">S. asiatica</taxonomicName>
), and the cerci (most setae subequal length and slightly curved in 
<taxonomicName lsidName="S. macropora" pageId="7" pageNumber="106" rank="species" species="macropora">S. macropora</taxonomicName>
sp. n. vs long and erect setae on dorsal, ventral and outer sides arranged in one or two rows in 
<taxonomicName lsidName="S. asiatica" pageId="7" pageNumber="106" rank="species" species="asiatica">S. asiatica</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>