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<document id="9D1CFBF411609390A0D9D3000A644A3C" ID-CLB-Dataset="53713" ID-DOI="10.5852/ejt.2023.872.2131" ID-GBIF-Dataset="31ad2cf7-cce8-4202-9a46-d81b0073d561" ID-ISSN="2118-9773" ID-Zenodo-Dep="8018303" ID-ZooBank="05098E38-AB11-486E-8F28-8567DE6BC19C" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1686244863328" checkinUser="felipe" docAuthor="Ebejer, Martin J." docDate="2023" docId="505487F2B008FF9AFD80FEC9084C3E4A" docLanguage="en" docName="ejt-872_1-161.pdf" docOrigin="European Journal of Taxonomy 872" docSource="http://zoobank.org/05098e38-ab11-486e-8f28-8567de6bc19c" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aphaniosoma harteni Ebejer 1996" docType="treatment" docVersion="8" lastPageNumber="77" masterDocId="AC6DFF8AB044FFD6FFC7FFC10B313A78" masterDocTitle="The genus Aphaniosoma Becker, 1903 (Diptera: Chyromyidae) in the Eastern Mediterranean and Middle East, with descriptions of new species" masterLastPageNumber="161" masterPageNumber="1" pageNumber="77" updateTime="1699521912180" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="100ADEC4C53F3D88C4EF97F3995093EB">The genus Aphaniosoma Becker, 1903 (Diptera: Chyromyidae) in the Eastern Mediterranean and Middle East, with descriptions of new species</mods:title>
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<treatment id="505487F2B008FF9AFD80FEC9084C3E4A" ID-DOI="http://doi.org/10.5281/zenodo.8018407" ID-GBIF-Taxon="209675214" ID-Zenodo-Dep="8018407" LSID="urn:lsid:plazi:treatment:505487F2B008FF9AFD80FEC9084C3E4A" httpUri="http://treatment.plazi.org/id/505487F2B008FF9AFD80FEC9084C3E4A" lastPageNumber="77" pageId="76" pageNumber="77">
<subSubSection id="90E7656FB008FF9AFD80FEC908DC3B5B" box="[583,1005,264,291]" pageId="76" pageNumber="77" type="nomenclature">
<paragraph id="D84236E4B008FF9AFD80FEC908DC3B5B" blockId="76.[583,1005,264,326]" box="[583,1005,264,291]" pageId="76" pageNumber="77">
<heading id="830A8188B008FF9AFD80FEC908DC3B5B" box="[583,1005,264,291]" centered="true" fontSize="11" level="2" pageId="76" pageNumber="77" reason="2">
<taxonomicName id="1FFD4D67B008FF9AFD80FEC908DC3B5B" ID-CoL="67LZY" authority="Ebejer, 1996" authorityName="Ebejer" authorityYear="1996" box="[583,1005,264,291]" class="Insecta" family="Chyromyidae" genus="Aphaniosoma" kingdom="Animalia" order="Diptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="harteni">
<emphasis id="EA89EAF6B008FF9AFD80FEC908603B5B" bold="true" box="[583,849,264,291]" italics="true" pageId="76" pageNumber="77">Aphaniosoma harteni</emphasis>
<bibRefCitation id="BC6C4B15B008FF9AFC9FFEC808DC3B5B" author="Ebejer M. J." box="[856,1005,265,291]" pageId="76" pageNumber="77" pagination="280 - 299" refId="ref57223" refString="Ebejer M. J. 1996. Chyromyidae (Diptera: Schizophora) from the Arabian Peninsula with descriptions of twelve new species. Fauna of Saudi Arabia 15: 280 - 299." type="journal article" year="1996">Ebejer, 1996</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="D84236E4B008FF9AFD1FFEED086D3B3E" blockId="76.[583,1005,264,326]" box="[728,860,300,326]" pageId="76" pageNumber="77">
<figureCitation id="40C62A61B008FF9AFD1FFEED086D3B3E" box="[728,860,300,326]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="77.[189,232,1963,1989]" captionStartId-1="78.[189,232,1963,1989]" captionTargetBox-0="[221,1372,265,1925]" captionTargetBox-1="[257,1328,1363,1925]" captionTargetId-0="figure-11@77.[216,1372,265,1925]" captionTargetId-1="figure-256@78.[257,1328,1363,1925]" captionTargetPageId-0="77" captionTargetPageId-1="78" captionText-0="Fig. 39. Aphaniosoma harteni Ebejer, 1996. A. ♂, holotype (NMWC), habitus, lateral view. B. ♂, (MJE), from Israel, habitus, lateral view." captionText-1="Fig. 40. Aphaniosoma harteni Ebejer, 1996, ♂, holotype (NMWC), hypopygium in situ. A. Lateral view. B. Ventro-lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.8018417" figureDoi-1="http://doi.org/10.5281/zenodo.8018421" httpUri-0="https://zenodo.org/record/8018417/files/figure.png" httpUri-1="https://zenodo.org/record/8018421/files/figure.png" pageId="76" pageNumber="77">Figs 39–40</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="90E7656FB008FF9AFF7AFEAC0ACE38BE" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="D84236E4B008FF9AFF7AFEAC0A9C3BFF" blockId="76.[189,429,365,391]" box="[189,429,365,391]" pageId="76" pageNumber="77">
<heading id="830A8188B008FF9AFF7AFEAC0A9C3BFF" bold="true" box="[189,429,365,391]" fontSize="11" level="3" pageId="76" pageNumber="77" reason="3">
<emphasis id="EA89EAF6B008FF9AFF7AFEAC0A9C3BFF" bold="true" box="[189,429,365,391]" pageId="76" pageNumber="77">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="D84236E4B008FF9AFF2DFE5D0A6C3BCE" blockId="76.[189,921,412,474]" box="[234,349,412,438]" pageId="76" pageNumber="77">
<emphasis id="EA89EAF6B008FF9AFF2DFE5D0A6C3BCE" bold="true" box="[234,349,412,438]" pageId="76" pageNumber="77">
<typeStatus id="07468846B008FF9AFF2DFE5D0A6C3BCE" box="[234,349,412,438]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="D84236E4B008FF9AFF7AFE0108A83BA1" blockId="76.[189,921,412,474]" box="[189,921,447,474]" pageId="76" pageNumber="77">
<materialsCitation id="68953CB9B008FF9AFF7AFE0108A83BA1" ID-GBIF-Occurrence="4127865710" box="[189,921,447,474]" collectingDate="1991-08" collectionCode="NMWC" collectorName="A. van Harten" country="Yemen" location="Sana'a" pageId="76" pageNumber="77" specimenCount="1" specimenCount-male="1" stateProvince="Sana'a" typeStatus="holotype">
<collectingCountry id="A0EA7674B008FF9AFF7AFE010A173BA2" box="[189,294,448,474]" name="Yemen" pageId="76" pageNumber="77">YEMEN</collectingCountry>
•
<specimenCount id="CEFBFD6DB008FF9AFEF8FE010A693BA2" box="[319,344,448,474]" pageId="76" pageNumber="77" type="male">♂</specimenCount>
;
<collectingRegion id="1A39F806B008FF9AFEA2FE7E0A883BA1" box="[357,441,447,473]" country="Yemen" name="San'a'" pageId="76" pageNumber="77">Sana’a</collectingRegion>
;
<date id="AC431024B008FF9AFE03FE0109743BA1" box="[452,581,447,474]" pageId="76" pageNumber="77" value="1991-08">
<collectingDate id="BC07E9CCB008FF9AFE03FE0109743BA1" box="[452,581,447,474]" pageId="76" pageNumber="77" value="1991-08">Aug. 1991</collectingDate>
</date>
;
<collectorName id="75085332B008FF9AFD88FE0109C63BA2" box="[591,759,448,474]" pageId="76" pageNumber="77">A. van Harten</collectorName>
leg;
<collectionCode id="BEECAE21B008FF9AFCF5FE7E08A43BA1" box="[818,917,447,473]" collectionName="United Kingdom, Cardiff, National Museum of Wales" country="United Kingdom" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="76" pageNumber="77" type="Museum">NMWC</collectionCode>
.
</materialsCitation>
</paragraph>
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<emphasis id="EA89EAF6B008FF9AFF2DFDC10A993862" bold="true" box="[234,424,512,538]" pageId="76" pageNumber="77">Other material</emphasis>
</paragraph>
<paragraph id="D84236E4B008FF9AFF7AFDE50920381A" blockId="76.[189,1398,512,610]" pageId="76" pageNumber="77">
<materialsCitation id="68953CB9B008FF9AFF7AFDE5093C381A" ID-GBIF-Occurrence="4127865408" collectingDate="2000-03-24" collectionCode="MJE" collectorName="M. J. Ebejer" country="Israel" latitude="30.766666" location="Ezuz" longLatPrecision="1220" longitude="34.483334" municipality="Ezuz" pageId="76" pageNumber="77" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Western Negev">
<collectingCountry id="A0EA7674B008FF9AFF7AFDE50A123846" box="[189,291,548,574]" name="Israel" pageId="76" pageNumber="77">ISRAEL</collectingCountry>
•
<specimenCount id="CEFBFD6DB008FF9AFEF8FDE50A5E3847" box="[319,367,548,575]" pageId="76" pageNumber="77" type="male">1 ♂</specimenCount>
,
<specimenCount id="CEFBFD6DB008FF9AFEB9FDE50AF23847" box="[382,451,548,575]" pageId="76" pageNumber="77" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1A39F806B008FF9AFE15FDE409B83847" box="[466,649,549,575]" pageId="76" pageNumber="77">Western Negev</collectingRegion>
,
<location id="DD22603FB008FF9AFD51FDE409E53847" LSID="urn:lsid:plazi:treatment:505487F2B008FF9AFD80FEC9084C3E4A:DD22603FB008FF9AFD51FDE409E53847" box="[662,724,549,575]" country="Israel" latitude="30.766666" longLatPrecision="1220" longitude="34.483334" municipality="Ezuz" name="Ezuz" pageId="76" pageNumber="77" stateProvince="Western Negev">Ezuz</location>
;
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,
<geoCoordinate id="BDC95023B008FF9AFC99FDE308F83847" box="[862,969,546,575]" degrees="34" direction="east" minutes="29" orientation="longitude" pageId="76" pageNumber="77" precision="925" value="34.483334">34°29ʹ E</geoCoordinate>
;
<date id="AC431024B008FF9AFC10FDE50F4A3846" box="[983,1147,548,575]" pageId="76" pageNumber="77" value="2000-03-24">
<collectingDate id="BC07E9CCB008FF9AFC10FDE50F4A3846" box="[983,1147,548,575]" pageId="76" pageNumber="77" value="2000-03-24">24 Mar. 2000</collectingDate>
</date>
;
<collectorName id="75085332B008FF9AFB4EFDE40E253846" box="[1161,1300,548,575]" pageId="76" pageNumber="77">M.J. Ebejer</collectorName>
leg.; on grazed grass and dung;
<collectionCode id="BEECAE21B008FF9AFE15FD89093C381A" box="[466,525,584,610]" pageId="76" pageNumber="77">MJE</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="D84236E4B008FF9AFF7AFD480ACE38BE" blockId="76.[189,1398,647,710]" pageId="76" pageNumber="77">
<materialsCitation id="68953CB9B008FF9AFF7AFD480ACA38BE" ID-GBIF-Occurrence="4127865840" collectingDate="2014-05-07" collectingDateMax="2014-05-28" collectingDateMin="2014-05-07" collectingMethod="Malaise trap" collectionCode="NMWC" collectorName="H. A. Dawah" country="Saudi Arabia" latitude="17.516666" location="Hay Alshurfa" longLatPrecision="1278" longitude="44.25" pageId="76" pageNumber="77" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Najran">
<collectingCountry id="A0EA7674B008FF9AFF7AFD480ABD38DB" box="[189,396,649,675]" name="Saudi Arabia" pageId="76" pageNumber="77">SAUDI ARABIA</collectingCountry>
•
<specimenCount id="CEFBFD6DB008FF9AFE62FD480AE538DB" box="[421,468,649,675]" pageId="76" pageNumber="77" type="male">1 ♂</specimenCount>
,
<specimenCount id="CEFBFD6DB008FF9AFE26FD48093F38DB" box="[481,526,649,675]" pageId="76" pageNumber="77" type="female">1 ♀</specimenCount>
;
<collectingRegion id="1A39F806B008FF9AFDDAFD48094138DB" box="[541,624,649,675]" country="Saudi Arabia" name="Najran" pageId="76" pageNumber="77">Najran</collectingRegion>
,
<location id="DD22603FB008FF9AFDBBFD48082F38DB" LSID="urn:lsid:plazi:treatment:505487F2B008FF9AFD80FEC9084C3E4A:DD22603FB008FF9AFDBBFD48082F38DB" box="[636,798,649,675]" country="Saudi Arabia" latitude="17.516666" longLatPrecision="1278" longitude="44.25" name="Hay Alshurfa" pageId="76" pageNumber="77" stateProvince="Najran">Hay Alshurfa</location>
;
<geoCoordinate id="BDC95023B008FF9AFCEEFD4608A738DB" box="[809,918,647,675]" degrees="17" direction="north" minutes="31" orientation="latitude" pageId="76" pageNumber="77" precision="925" value="17.516666">17°31ʹ N</geoCoordinate>
,
<geoCoordinate id="BDC95023B008FF9AFC65FD460F3A38DB" box="[930,1035,647,675]" degrees="44" direction="east" minutes="15" orientation="longitude" pageId="76" pageNumber="77" precision="925" value="44.25">44°15ʹ E</geoCoordinate>
;
<date id="AC431024B008FF9AFBDFFD480FE738DB" box="[1048,1238,649,675]" pageId="76" pageNumber="77" value="2014-05-07" valueMax="2014-05-28" valueMin="2014-05-07">
<collectingDate id="BC07E9CCB008FF9AFBDFFD480FE738DB" box="[1048,1238,649,675]" pageId="76" pageNumber="77" value="2014-05-07" valueMax="2014-05-28" valueMin="2014-05-07">7–28 May 2014</collectingDate>
</date>
;
<collectorName id="75085332B008FF9AFB24FD480E4738DB" box="[1251,1398,649,675]" pageId="76" pageNumber="77">H.A. Dawah</collectorName>
leg.;
<collectingMethod id="01BC4EF3B008FF9AFF3FFD6D0ABC38BE" box="[248,397,684,710]" pageId="76" pageNumber="77">Malaise trap</collectingMethod>
;
<collectionCode id="BEECAE21B008FF9AFE5EFD6D0ACA38BE" box="[409,507,684,710]" country="United Kingdom" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="76" pageNumber="77" type="Museum">NMWC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90E7656FB008FF9AFF7AFD2C08A939BB" pageId="76" pageNumber="77" type="discussion">
<paragraph id="D84236E4B008FF9AFF7AFD2C0A01397F" blockId="76.[189,304,749,775]" box="[189,304,749,775]" pageId="76" pageNumber="77">
<heading id="830A8188B008FF9AFF7AFD2C0A01397F" bold="true" box="[189,304,749,775]" fontSize="11" level="3" pageId="76" pageNumber="77" reason="3">
<emphasis id="EA89EAF6B008FF9AFF7AFD2C0A01397F" bold="true" box="[189,304,749,775]" pageId="76" pageNumber="77">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="D84236E4B008FF9AFF7AFCDD08A939BB" blockId="76.[189,1399,796,963]" pageId="76" pageNumber="77">
When this species was described the subtle projection on the hind tronchanter was missed. The type series was re-examined and further illustrations provided (
<bibRefCitation id="BC6C4B15B008FF9AFC9BFCFE0F193921" author="Ebejer M. J." box="[860,1064,831,857]" pageId="76" pageNumber="77" pagination="221 - 236" refId="ref57498" refString="Ebejer M. J. 2016. The Moroccan species of Chyromyidae (Diptera) with descriptions of five new species of Aphaniosoma Becker. Zootaxa 4208 (3): 221 - 236. https: // doi. org / 10.11646 / zootaxa. 4208.3.2" type="journal article" year="2016">Ebejer 2016: 231</bibRefCitation>
, figs 14b, 15a–b). The record from
<collectingCountry id="A0EA7674B008FF9AFF3AFCA30A653904" box="[253,340,866,892]" name="Tunisia" pageId="76" pageNumber="77">Tunisia</collectingCountry>
(
<bibRefCitation id="BC6C4B15B008FF9AFEA4FCA30AC73904" author="Ebejer M. J." box="[355,502,866,892]" pageId="76" pageNumber="77" pagination="191 - 230" refId="ref57257" refString="Ebejer M. J. 1998. A Review of the Palaearctic species of Aphaniosoma Becker (Diptera, Chyromyidae), with descriptions of new species and a key for the identification of adults. Mitteilungen Museum fur Naturkunde Berliner Deutsche entomologische Zeitschrift 45 (2): 191 - 230. https: // doi. org / 10.1002 / mmnd. 4810450208" type="journal article" year="1998">Ebejer 1998</bibRefCitation>
) is now doubtful considering that many similar species have been described since, and a number of additional similar species known to the author are still awaiting description. Photographs of the
<typeStatus id="07468846B008FF9AFE65FC69093839BA" box="[418,521,936,962]" pageId="76" pageNumber="77" type="holotype">holotype</typeStatus>
are provided here (
<figureCitation id="40C62A61B008FF9AFD2AFC69086C39BB" box="[749,861,936,963]" captionStart="Fig" captionStartId="77.[189,232,1963,1989]" captionTargetBox="[221,1372,265,1925]" captionTargetId="figure-11@77.[216,1372,265,1925]" captionTargetPageId="77" captionText="Fig. 39. Aphaniosoma harteni Ebejer, 1996. A. ♂, holotype (NMWC), habitus, lateral view. B. ♂, (MJE), from Israel, habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8018417" httpUri="https://zenodo.org/record/8018417/files/figure.png" pageId="76" pageNumber="77">Figs 39A</figureCitation>
,
<figureCitation id="40C62A61B008FF9AFCAEFC6808BA39BB" box="[873,907,937,963]" captionStart="Fig" captionStartId="78.[189,232,1963,1989]" captionTargetBox="[257,1328,1363,1925]" captionTargetId="figure-256@78.[257,1328,1363,1925]" captionTargetPageId="78" captionText="Fig. 40. Aphaniosoma harteni Ebejer, 1996, ♂, holotype (NMWC), hypopygium in situ. A. Lateral view. B. Ventro-lateral view." figureDoi="http://doi.org/10.5281/zenodo.8018421" httpUri="https://zenodo.org/record/8018421/files/figure.png" pageId="76" pageNumber="77">40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90E7656FB008FF9AFF7AFC2B084C3E4A" box="[189,893,1002,1075]" pageId="76" pageNumber="77" type="distribution">
<paragraph id="D84236E4B008FF9AFF7AFC2B0A693E7C" blockId="76.[189,344,1002,1028]" box="[189,344,1002,1028]" pageId="76" pageNumber="77">
<heading id="830A8188B008FF9AFF7AFC2B0A693E7C" bold="true" box="[189,344,1002,1028]" fontSize="11" level="3" pageId="76" pageNumber="77" reason="3">
<emphasis id="EA89EAF6B008FF9AFF7AFC2B0A693E7C" bold="true" box="[189,344,1002,1028]" pageId="76" pageNumber="77">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D84236E4B008FF9AFF7AFBD8084C3E4A" blockId="76.[189,893,1048,1075]" box="[189,893,1048,1075]" pageId="76" pageNumber="77">
<collectingCountry id="A0EA7674B008FF9AFF7AFBD80A3E3E4B" box="[189,271,1049,1075]" name="Yemen" pageId="76" pageNumber="77">Yemen</collectingCountry>
(
<bibRefCitation id="BC6C4B15B008FF9AFED9FBD90A823E4B" author="Ebejer M. J." box="[286,435,1048,1075]" pageId="76" pageNumber="77" pagination="280 - 299" refId="ref57223" refString="Ebejer M. J. 1996. Chyromyidae (Diptera: Schizophora) from the Arabian Peninsula with descriptions of twelve new species. Fauna of Saudi Arabia 15: 280 - 299." type="journal article" year="1996">Ebejer 1996</bibRefCitation>
). New records for
<collectingCountry id="A0EA7674B008FF9AFD4AFBD909E03E4A" box="[653,721,1048,1074]" name="Israel" pageId="76" pageNumber="77">Israel</collectingCountry>
,
<collectingCountry id="A0EA7674B008FF9AFD1AFBD908483E4A" box="[733,889,1048,1074]" name="Saudi Arabia" pageId="76" pageNumber="77">Saudi Arabia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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