174 lines
18 KiB
XML
174 lines
18 KiB
XML
<document id="337C36FC34104254BE214C9A358D3472" ID-DOI="10.1206/648.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5357410" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630220101912" checkinUser="carolina" docAuthor="Norell, Mark A., Makovicky, Peter J., Bever, Gabe S., Balanoff, Amy M., Clark, James M., Barsbold, Rinchen & Rowe, Timothy" docDate="2009" docId="03A6E839FFB7FFA1FF68BF7EFBD81C46" docLanguage="en" docName="N3654.pdf" docOrigin="American Museum Novitates 3654" docSource="http://www.bioone.org/doi/abs/10.1206/648.1" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Zanabazar junior" docType="treatment" docVersion="4" lastPageNumber="28" masterDocId="FF9F9041FFACFFBAFFF2BE1DFFE31D24" masterDocTitle="A Review of the Mongolian Cretaceous Dinosaur Saurornithoides (Troodontidae: Theropoda)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="28" updateTime="1699191067590" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="53DA43B27B4BA6AE6EB13209B468BBD1">
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<mods:title id="950DBF81108BDE8070522AE92FDFB958">A Review of the Mongolian Cretaceous Dinosaur Saurornithoides (Troodontidae: Theropoda)</mods:title>
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<mods:namePart id="9D95BFD104785C524BFE612CF0D8AF22">Norell, Mark A.</mods:namePart>
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<mods:namePart id="753985A704982694357F48D8483EBE37">Makovicky, Peter J.</mods:namePart>
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<mods:namePart id="1268C2D461BD8D805BD47419CA8C90C9">Bever, Gabe S.</mods:namePart>
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<mods:roleTerm id="C3EA4CC4CD4BA7CB6E7F4FC1F214C4F1">Author</mods:roleTerm>
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<mods:namePart id="3AF0FD68B9532665B305F1EFB7BDDBC3">Balanoff, Amy M.</mods:namePart>
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<mods:name id="12D1F7F989C8DC5B90D27BC3C4C02333" type="personal">
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<mods:namePart id="F47A0FAEDB64B9E0B23D9B41F7FAAF2C">Clark, James M.</mods:namePart>
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<mods:namePart id="BA02891A7DA37B2826E550F2B53E9F8D">Barsbold, Rinchen</mods:namePart>
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<mods:namePart id="F654703A3B7808479CBCAAED1206484F">Rowe, Timothy</mods:namePart>
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<mods:title id="4380E5AA6309B362DAF2983E7CF7A71C">American Museum Novitates</mods:title>
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<mods:part id="DE801118084FE1966D4F426848E60E19">
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<mods:date id="BC9C3F7CAB88A427094741BD73373B2C">2009</mods:date>
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<mods:number id="EF5C28060F4494F8F643701D2189D046">2009-06-25</mods:number>
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<mods:detail id="57839709B0C2029F7A81845C240971D8" type="volume">
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<mods:number id="2ABF7BFAF1A1107BB8254D83D93A4551">3654</mods:number>
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<mods:start id="44F678D90EE9E724780FF407F5B17C26">1</mods:start>
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<mods:url id="2753B5ED7637AF8DAB034F7F696E57F5">http://www.bioone.org/doi/abs/10.1206/648.1</mods:url>
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<mods:classification id="1B156294189329E31A77C2CBE48DED7C">journal article</mods:classification>
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<mods:identifier id="9ABF91F78FF55A2ADC0BC8800F081578" type="DOI">10.1206/648.1</mods:identifier>
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<mods:identifier id="4D597E21FCDEE2268D9BEA033A9B56FE" type="ISSN">0003-0082</mods:identifier>
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<treatment id="03A6E839FFB7FFA1FF68BF7EFBD81C46" ID-DOI="http://doi.org/10.5281/zenodo.5454728" ID-Zenodo-Dep="5454728" LSID="urn:lsid:plazi:treatment:03A6E839FFB7FFA1FF68BF7EFBD81C46" httpUri="http://treatment.plazi.org/id/03A6E839FFB7FFA1FF68BF7EFBD81C46" lastPageNumber="28" pageId="27" pageNumber="28">
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<subSubSection id="C3150AA4FFB7FFA1FF68BF7EFDC71C5E" box="[154,548,354,379]" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BB0592FFFB7FFA1FF68BF7EFDC71C5E" blockId="27.[154,548,354,379]" box="[154,548,354,379]" pageId="27" pageNumber="28">
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<heading id="D0F8EE43FFB7FFA1FF68BF7EFDC71C5E" box="[154,548,354,379]" centered="true" fontSize="36" level="2" pageId="27" pageNumber="28" reason="8">
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<taxonomicName id="4C0F22ACFFB7FFA1FF68BF7EFDC71C5E" authority="(Barsbold, 1974)" baseAuthorityName="Barsbold" baseAuthorityYear="1974" box="[154,548,354,379]" class="Reptilia" family="Troodontidae" genus="Zanabazar" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="junior">
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<emphasis id="B97B853DFFB7FFA1FF68BF7EFEBF1C5F" box="[154,348,355,379]" italics="true" pageId="27" pageNumber="28">Zanabazar junior</emphasis>
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(
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<bibRefCitation id="EF9E24DEFFB7FFA1FE9FBF7FFDF81C5E" author="Barsbold, R." box="[365,539,354,378]" pageId="27" pageNumber="28" pagination="5 - 22" refId="ref23053" refString="Barsbold, R. 1974. Saurornithoididae, a new family of small theropod dinosaurs from central Asia and North America. Palaeontologia Polonica 30: 5 - 22." type="journal article" year="1974">Barsbold, 1974</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3150AA4FFB7FFA1FF80BF80FF3B1EDA" pageId="27" pageNumber="28" type="materials_examined">
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<paragraph id="8BB0592FFFB7FFA1FF80BF80FDD41EE7" blockId="27.[88,614,413,1754]" pageId="27" pageNumber="28">
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<materialsCitation id="3B675372FFB7FFA1FF80BF80FDD41EE7" collectionCode="MT" pageId="27" pageNumber="28" specimenCode="IGM 100" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B4E78DFFB7FFA1FF80BF80FF001C90" box="[114,227,413,437]" pageId="27" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
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:
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<specimenCode id="DBA9F154FFB7FFA1FEF4BF80FE971C91" box="[262,372,413,437]" collectionCode="IGM" country="Mongolia" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="27" pageNumber="28">IGM 100</specimenCode>
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/1, a skull and partial mandible, six sacral and 11 caudal vertebrae, and the distal part of the right hind limb. The skull is largely complete except for the left and right quadrate, quadratojugal, ventral part of the lacrimal, palatine, pterygoid, and ectopterygoid, and the left squamosal, postorbital, and jugal, posterior part of the right jugal, and the postdentary bones other than the splenials. The posterior part of the left nasal is reconstructed in plaster, as are small parts of the left maxilla anterior to the maxillary fenestra and dorsal to the interfenestral bar. Plaster covers the ventral surface of the bones roofing the antorbital fenestra. The mandible comprises only the dentaries and splenials. The right hind limb includes the distal end of the tibia with the coossified astragalus and calcaneum, a distal tarsal, and the proximal ends of
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<collectionCode id="ED1EC1EAFFB7FFA1FE4DBDB6FE081EE7" box="[447,491,939,963]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="27" pageNumber="28">MT</collectionCode>
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II–IV.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB0592FFFB7FFA1FF80BDD4FF3B1EDA" blockId="27.[88,614,413,1754]" pageId="27" pageNumber="28">
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<collectionCode id="ED1EC1EAFFB7FFA1FF80BDD4FE3D1EFB" box="[114,478,969,993]" pageId="27" pageNumber="28">TYPE LOCALITY AND AGE</collectionCode>
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: As for
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<taxonomicName id="4C0F22ACFFB7FFA1FFAABDFBFF321EDA" authorityName="Norell & Makovicky & Bever & Balanoff & Clark & Barsbold & Rowe" authorityYear="2009" box="[88,209,998,1022]" class="Reptilia" family="Troodontidae" genus="Zanabazar" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B97B853DFFB7FFA1FFAABDFBFF321EDA" box="[88,209,998,1022]" italics="true" pageId="27" pageNumber="28">Zanabazar</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3150AA4FFB7FFA1FF80BA19FE1318BC" pageId="27" pageNumber="28" type="diagnosis">
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<paragraph id="8BB0592FFFB7FFA1FF80BA19FE391938" blockId="27.[88,614,413,1754]" box="[114,474,1028,1052]" pageId="27" pageNumber="28">
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DIAGNOSIS: As for
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<taxonomicName id="4C0F22ACFFB7FFA1FEA9BA19FE371938" authorityName="Norell & Makovicky & Bever & Balanoff & Clark & Barsbold & Rowe" authorityYear="2009" box="[347,468,1028,1052]" class="Reptilia" family="Troodontidae" genus="Zanabazar" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B97B853DFFB7FFA1FEA9BA19FE371938" box="[347,468,1028,1052]" italics="true" pageId="27" pageNumber="28">Zanabazar</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BB0592FFFB7FFA1FF80BA3CFE1318BC" blockId="27.[88,614,413,1754]" pageId="27" pageNumber="28">
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REFERRED SPECIMENS: No new material of this species has been described, but two other troodontids have been described from the Nemegt Formation of Mongolia.
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<taxonomicName id="4C0F22ACFFB7FFA1FE05BA64FE7A198A" authority="Osmolska, 1987" authorityName="Osmolska" authorityYear="1987" class="Reptilia" family="Troodontidae" genus="Borogovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="gracilicrus">
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<emphasis id="B97B853DFFB7FFA1FE05BA64FF2B198B" italics="true" pageId="27" pageNumber="28">Borogovia gracilicrus</emphasis>
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<bibRefCitation id="EF9E24DEFFB7FFA1FF29BA8BFE7A198A" author="Osmolska, H." box="[219,409,1174,1198]" pageId="27" pageNumber="28" pagination="133 - 150" refId="ref25109" refString="Osmolska, H. 1987. Borogovia gracilicrus gen. et sp. n., a new troodontid dinosaur from the Late Cretaceous of Mongolia. Acta Palaeontologica Polonica 31: 133 - 150." type="journal article" year="1987">Osmólska, 1987</bibRefCitation>
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</taxonomicName>
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, is known only from a partial hind limb from Altan Uul IV, and
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<taxonomicName id="4C0F22ACFFB7FFA1FF66BACCFEE91822" authority="Kurzanov and Osmolska, 1991" authorityName="Kurzanov and Osmolska" authorityYear="1991" class="Reptilia" family="Troodontidae" genus="Tochisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="nemegtensis">
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<emphasis id="B97B853DFFB7FFA1FF66BACCFE4B19CD" box="[148,424,1233,1257]" italics="true" pageId="27" pageNumber="28">Tochisaurus nemegtensis</emphasis>
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<bibRefCitation id="EF9E24DEFFB7FFA1FE48BACCFEE91822" author="Kurzanov, S. M. & H. Osmolska" pageId="27" pageNumber="28" pagination="69 - 76" refId="ref24468" refString="Kurzanov, S. M., and H. Osmolska. 1991. Tochisaurus nemegtensis gen. et sp. n., a new troodontid (Dinosauria, Theropoda) from Mongolia. Acta Palaeontologica Polonica 36: 69 - 76." type="journal article" year="1991">Kurzanov and Osmólska, 1991</bibRefCitation>
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</taxonomicName>
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, is known only from a metatarsus from the Nemegt area. Because the postcranial elements of
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<taxonomicName id="4C0F22ACFFB7FFA1FEFABB34FE201865" baseAuthorityName="Barsbold" baseAuthorityYear="1974" box="[264,451,1321,1345]" class="Reptilia" family="Troodontidae" genus="Zanabazar" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="junior">
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<emphasis id="B97B853DFFB7FFA1FEFABB34FE201865" box="[264,451,1321,1345]" italics="true" pageId="27" pageNumber="28">Zanabazar junior</emphasis>
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</taxonomicName>
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do not include those parts diagnostic of the other two species, it is possible that one of the other two Nemegt Formation taxa is a junior synonym.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3150AA4FFB7FFA1FF80BB83FBD81C46" pageId="27" pageNumber="28" type="discussion">
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<paragraph id="8BB0592FFFB7FFA1FF80BB83FBD81C46" blockId="27.[88,614,413,1754]" lastBlockId="27.[662,1188,211,354]" pageId="27" pageNumber="28">
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COMMENTS: We chose to construct the name
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<taxonomicName id="4C0F22ACFFB7FFA1FF55BBA6FEFC18F7" authorityName="Norell & Makovicky & Bever & Balanoff & Clark & Barsbold & Rowe" authorityYear="2009" box="[167,287,1467,1491]" class="Reptilia" family="Troodontidae" genus="Zanabazar" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B97B853DFFB7FFA1FF55BBA6FEFC18F7" box="[167,287,1467,1491]" italics="true" pageId="27" pageNumber="28">Zanabazar</emphasis>
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</taxonomicName>
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for
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<taxonomicName id="4C0F22ACFFB7FFA1FE94BBA6FD8518F7" box="[358,614,1467,1491]" class="Reptilia" family="Troodontidae" genus="Saurornithoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="junior">
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<emphasis id="B97B853DFFB7FFA1FE94BBA6FD8518F7" box="[358,614,1467,1491]" italics="true" pageId="27" pageNumber="28">Saurornithoides junior</emphasis>
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</taxonomicName>
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based on the morphological differences between the
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<typeStatus id="54B4E78DFFB7FFA1FF38BBE8FECE1B29" box="[202,301,1525,1549]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
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and that of
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<taxonomicName id="4C0F22ACFFB7FFA1FE4BBBEBFF071B0F" authorityName="Osborn" authorityYear="1924" class="Reptilia" family="Troodontidae" genus="Saurornithoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="mongoliensis">
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<emphasis id="B97B853DFFB7FFA1FE4BBBEBFF071B0F" italics="true" pageId="27" pageNumber="28">Saurornithoides mongoliensis</emphasis>
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</taxonomicName>
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and the questionable monophyly of
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<taxonomicName id="4C0F22ACFFB7FFA1FF8DB82DFECF1B6C" authorityName="Osborn" authorityYear="1924" box="[127,300,1584,1608]" class="Reptilia" family="Troodontidae" genus="Saurornithoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
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<emphasis id="B97B853DFFB7FFA1FF8DB82DFECF1B6C" box="[127,300,1584,1608]" italics="true" pageId="27" pageNumber="28">Saurornithoides</emphasis>
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</taxonomicName>
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as defined by the sistertaxon relationship of
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<taxonomicName id="4C0F22ACFFB7FFA1FEA9B850FDE91B41" authorityName="Osborn" authorityYear="1924" box="[347,522,1613,1637]" class="Reptilia" family="Troodontidae" genus="Saurornithoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="mongoliensis">
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<emphasis id="B97B853DFFB7FFA1FEA9B850FDE91B41" box="[347,522,1613,1637]" italics="true" pageId="27" pageNumber="28">S. mongoliensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0F22ACFFB7FFA1FDBCB850FF7B1BA7" class="Reptilia" family="Troodontidae" genus="Saurornithoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="junior">
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<emphasis id="B97B853DFFB7FFA1FDBCB850FF7B1BA7" italics="true" pageId="27" pageNumber="28">S. junior</emphasis>
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</taxonomicName>
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. This decision also was influenced by the common practice in phylogenetic analyses of assuming monophyly by compositely coding taxa (especially ‘‘genera’’). The differences between
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<taxonomicName id="4C0F22ACFFB7FFA1FD08BECEFC591DCF" baseAuthorityName="Barsbold" baseAuthorityYear="1974" box="[762,954,211,235]" class="Reptilia" family="Troodontidae" genus="Zanabazar" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="junior">
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<emphasis id="B97B853DFFB7FFA1FD08BECEFC591DCF" box="[762,954,211,235]" italics="true" pageId="27" pageNumber="28">Zanabazar junior</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0F22ACFFB7FFA1FC04BECEFCC11C2D" authorityName="Osborn" authorityYear="1924" class="Reptilia" family="Troodontidae" genus="Saurornithoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="mongoliensis">
|
||
<emphasis id="B97B853DFFB7FFA1FC04BECEFCC11C2D" italics="true" pageId="27" pageNumber="28">Saurornithoides mongoliensis</emphasis>
|
||
</taxonomicName>
|
||
certainly merit coding these taxa separately. The temporal difference separating these specimens was not, and should not, be used to support taxonomic decisions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |