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<document id="682394FE9AFDDB5D9BD857AC645EB885" ID-DOI="10.5281/zenodo.5732150" ID-ISSN="2064-2474" ID-Zenodo-Dep="5732150" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638041842658" checkinUser="felipe" docAuthor="Makranczy, György" docDate="2014" docId="F21687A65674FFF2FDB8FC0FFB9923B8" docLanguage="en" docName="ActaZoolAcadSciH.60.1.13-38.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 60 (1)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Thinobius micros Fauvel 1871" docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="0E2FFFDE5660FFE7FFF4FFCBFFEE2547" masterDocTitle="Synonymies In The Holarctic Thinobius Major And Linearis Species Groups (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="38" masterPageNumber="13" pageNumber="33" updateTime="1699429339529" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="CF48496BD1A6AA3E3C2F535D72891553">Synonymies In The Holarctic Thinobius Major And Linearis Species Groups (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="BD7D15ECB2BF59207B5FAF5B89A463FA">Makranczy, György</mods:namePart>
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<mods:title id="973A55D77F3CBE72FE53B69F956CDFD0">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date id="80DAE1291DE48074D542D079A5C45F55">2014</mods:date>
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<mods:number id="6556D778F2CCB4D0068CABE05C2BB9C8">60</mods:number>
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<mods:identifier id="DF027ABD2BC571763A87E6DBB1F0011D" type="DOI">10.5281/zenodo.5732150</mods:identifier>
<mods:identifier id="7D359E6E84DCC8B17958CFEFE9863805" type="ISSN">2064-2474</mods:identifier>
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<treatment id="F21687A65674FFF2FDB8FC0FFB9923B8" LSID="urn:lsid:plazi:treatment:F21687A65674FFF2FDB8FC0FFB9923B8" httpUri="http://treatment.plazi.org/id/F21687A65674FFF2FDB8FC0FFB9923B8" lastPageId="21" lastPageNumber="34" pageId="20" pageNumber="33">
<subSubSection id="32A5653B5674FFF3FDB8FC0FFC2C2699" box="[588,962,964,991]" pageId="20" pageNumber="33" type="nomenclature">
<paragraph id="7A0036B05674FFF3FDB8FC0FFC2C2699" blockId="20.[588,962,964,991]" box="[588,962,964,991]" pageId="20" pageNumber="33">
<heading id="214881DC5674FFF3FDB8FC0FFC2C2699" box="[588,962,964,991]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="33" reason="2">
<taxonomicName id="BDBF4D335674FFF3FDB8FC0FFC2C2699" ID-CoL="7BWQ2" authority="Fauvel, 1871" authorityName="Fauvel" authorityYear="1871" box="[588,962,964,991]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="species" species="micros">
<emphasis id="48CBEAA25674FFF3FDB8FC0FFCF72699" box="[588,793,964,990]" italics="true" pageId="20" pageNumber="33">Thinobius micros</emphasis>
<bibRefCitation id="1E2E4B415674FFF3FCD5FC0EFC2C2699" author="FAuvel, A." box="[801,962,964,991]" pageId="20" pageNumber="33" pagination="1869 - 70" refId="ref9454" refString="FAuvel, A. (1871) Faune Gallo-Rhenane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhenanes, et la Valais avec tableaux synoptiques et planches gravees. Bulletin de la Societe Lineenne de Normandie 2 (5) [1869 - 70]: 27 - 192." type="journal article" year="1871">Fauvel, 1871</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="32A5653B5674FFF3FDA7FC2CFC542145" box="[595,954,999,1026]" pageId="20" pageNumber="33" type="description">
<paragraph id="7A0036B05674FFF3FDA7FC2CFC542145" blockId="20.[595,954,999,1026]" box="[595,954,999,1026]" pageId="20" pageNumber="33">
<heading id="214881DC5674FFF3FDA7FC2CFC542145" box="[595,954,999,1026]" centered="true" fontSize="11" level="3" pageId="20" pageNumber="33" reason="3">
(
<figureCitation id="E2842A355674FFF3FDA8FC2CFD452146" box="[604,683,999,1026]" captionStart="Figs 1–6" captionStartId="4.[299,344,1713,1735]" captionTargetBox="[305,1244,405,1695]" captionTargetId="figure-18@4.[305,1245,405,1695]" captionTargetPageId="4" captionText="Figs 1–6. Forebodies of Thinobius species: 1 = T. major Kraatz, 1857, 2 = T. angusticeps Fauvel, 1889, 3 = T. procerus Eppelsheim, 1893, 4 = T. bicolor Joy, 1911, 5 = T. petzi Bernhauer, 1908, 6 = T. micros Fauvel, 1871. Scales: 0.07 mm for Fig. 6, 0.10 mm for Figs 2, 4–5, 0.14 mm for Figs 1, 3." figureDoi="http://doi.org/10.5281/zenodo.5732154" httpUri="https://zenodo.org/record/5732154/files/figure.png" pageId="20" pageNumber="33">Figs 6</figureCitation>
,
<figureCitation id="E2842A355674FFF3FD43FC23FD382145" box="[695,726,1000,1026]" captionStart="Figs 13–26" captionStartId="9.[336,381,1630,1652]" captionTargetBox="[357,1268,405,1603]" captionTargetId="figure-11@9.[356,1268,405,1605]" captionTargetPageId="9" captionText="Figs 13–26. Antennae of Thinobius species: 13 = T. major Kraatz, 1857, 14 = T. angusticeps Fauvel, 1889, 15 = T. procerus Eppelsheim, 1893, 16 = T. bicolor Joy, 1911, 17 = T. petzi Bernhauer, 1908, 18 = T. micros Fauvel, 1871, 19 = T. paramicros sp. n. 20–22 = T. major Kraatz, 1857, male, 20 = sternite VIII, 21 = tergite IX, 22 = sternite IX. 23–24 = T. angusticeps Fauvel, 1889, male, 23 = tergite IX, 24 = sternite IX. 25–26 = T. procerus Eppelsheim, 1893, male, 25 = tergite IX, 26 = sternite IX. Scales: 0.05 mm for Figs 13, 15, 0.06 mm for Figs 14, 16–17, 26, 0.07 mm for Fig." figureDoi="http://doi.org/10.5281/zenodo.5732160" httpUri="https://zenodo.org/record/5732160/files/figure.png" pageId="20" pageNumber="33">18</figureCitation>
,
<figureCitation id="E2842A355674FFF3FD17FC2CFCC12145" box="[739,815,999,1026]" captionStart="Figs 34–40" captionStartId="15.[336,381,1709,1731]" captionTargetBox="[346,1279,406,1683]" captionTargetId="figure-10@15.[345,1280,405,1689]" captionTargetPageId="15" captionText="Figs 34–40. 34 = T. petzi Bernhauer, 1908, male sternite VIII. 35–37 = T. micros Fauvel, 1871, male, 35 = sternite VIII, 36 = tergite IX, 37 = sternite IX. 38–40 = T. paramicros sp. n., male, 38 = sternite VIII, 39 = tergite IX, 40 = sternite IX. Scales: 0.08 mm for Fig. 36, 0.09 mm for Fig." figureDoi="http://doi.org/10.5281/zenodo.5732168" httpUri="https://zenodo.org/record/5732168/files/figure.png" pageId="20" pageNumber="33">35–37</figureCitation>
,
<figureCitation id="E2842A355674FFF3FCCFFC23FC5C2145" box="[827,946,999,1026]" captionStart="Figs 41–53" captionStartId="17.[336,381,1625,1647]" captionText="Figs 41–53. Aedeagi of Thinobius species: 41 = T. major Kraatz, 1857, 42 = T. angusticeps Fauvel, 1889, 43 = T. procerus Eppelsheim, 1893, 44 = T. micros Fauvel, 1871, 45 = T. paramicros sp. n. Femalegenitaltraits: 46–47 = T. angusticeps Fauvel, 1889, neotype, 46 = genitalappendage, 47 = spermatheca. 48–50 = T. micros Fauvel, 1871, 48 = genitalappendage, 49–50 = spermathecae. 51–53 = T. paramicros sp. n., 51 = genitalappendage, 52–53 = spermathecae. Scales: 0.05 mm for Figs 44, 48–50, 52–53, 0.065 mm for Figs 45, 51, 0.08 mm for Fig. 42, 0.10" pageId="20" pageNumber="33">44, 48–50</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="32A5653B5674FFF3FEDFFBD4FB542128" pageId="20" pageNumber="33" type="reference_group">
<paragraph id="7A0036B05674FFF3FEDFFBD4FCF02172" blockId="20.[299,1210,1055,1135]" box="[299,798,1055,1077]" pageId="20" pageNumber="33">
<treatmentCitationGroup id="5AAF119E5674FFF3FEDFFBD4FCF02172" box="[299,798,1055,1077]" pageId="20" pageNumber="33">
<treatmentCitation id="FB1E10A15674FFF3FEDFFBD4FD612172" author="FAuvel, A." box="[299,655,1055,1077]" page="163" pageId="20" pageNumber="33" year="1871">
<taxonomicName id="BDBF4D335674FFF3FEDFFBD4FD612172" ID-CoL="7BWQ2" authority="FAuvel, 1871: 163" authorityName="Fauvel" authorityYear="1871" box="[299,655,1055,1077]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="species" species="micros">
<emphasis id="48CBEAA25674FFF3FEDFFBD4FE3C2172" box="[299,466,1055,1077]" italics="true" pageId="20" pageNumber="33">Thinobius micros</emphasis>
<bibRefCitation id="1E2E4B415674FFF3FE2CFBEBFD612172" author="FAuvel, A." box="[472,655,1055,1077]" pageId="20" pageNumber="33" pagination="1869 - 70" refId="ref9454" refString="FAuvel, A. (1871) Faune Gallo-Rhenane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhenanes, et la Valais avec tableaux synoptiques et planches gravees. Bulletin de la Societe Lineenne de Normandie 2 (5) [1869 - 70]: 27 - 192." type="journal article" year="1871">FAuvel, 1871: 163</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[=
<treatmentCitation id="FB1E10A15674FFF3FD5BFBEBFCFD2172" author="FAuvel, A." box="[687,787,1055,1077]" page="137" pageId="20" pageNumber="33" year="1872">
<bibRefCitation id="1E2E4B415674FFF3FD5BFBEBFCFD2172" author="FAuvel, A." box="[687,787,1055,1077]" pageId="20" pageNumber="33" pagination="1 - 214" refId="ref9516" refString="FAuvel, A. (1872) Faune Gallo-Rhenane ou Species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhenane, le Nassau et le Valais avec tableaux synoptiques et planches gravees, Coleopteres, Tome Troisieme: Staphylinides / Catalogue systematique des Staphylinides de la Faune Gallo-Rhenane avec l'addition synonymique des especes europeennes, siberiennes, caucasiques et mediterraneennes et descriptions nouvelles. Le Blanc- Hardel, Caen. 1 - 214 pp., plates I - II." type="book chapter" year="1872">1872: 137</bibRefCitation>
</treatmentCitation>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="7A0036B05674FFF3FEDFFBF7FD1F2116" blockId="20.[299,1210,1055,1135]" box="[299,753,1084,1106]" pageId="20" pageNumber="33">
<treatmentCitationGroup id="5AAF119E5674FFF3FEDFFBF7FD1F2116" box="[299,753,1084,1106]" pageId="20" pageNumber="33">
<treatmentCitation id="FB1E10A15674FFF3FEDFFBF7FD752116" author="FAuvel, A." box="[299,667,1084,1106]" page="94" pageId="20" pageNumber="33" year="1898">
<taxonomicName id="BDBF4D335674FFF3FEDFFBF7FD752116" ID-CoL="56GSW" authority="FAuvel, 1898: 94" authorityName="Fauvel. Thefollowingnewsynonymiesareproposed" authorityYear="1898" box="[299,667,1084,1106]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="13" phylum="Arthropoda" rank="species" species="algiricus">
<emphasis id="48CBEAA25674FFF3FEDFFBF7FE082116" box="[299,486,1084,1105]" italics="true" pageId="20" pageNumber="33">Thinobius algiricus</emphasis>
<bibRefCitation id="1E2E4B415674FFF3FE19FBF7FD752116" author="FAuvel, A." box="[493,667,1084,1106]" pageId="20" pageNumber="33" pagination="93 - 113" refId="ref9633" refString="FAuvel, A. (1898) Catalogue des Staphylinides de Barbarie et des Iles Acores, Maderes, Salvages et Canaries. Supplement. Description des especes nouvelles. Revue d'Entomologie 17: 93 - 113." type="journal article" year="1898">FAuvel, 1898: 94</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.,
<emphasis id="48CBEAA25674FFF3FD5DFBF7FD1F2116" bold="true" box="[681,753,1084,1105]" pageId="20" pageNumber="33">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="7A0036B05674FFF3FEDFFB92FB542128" blockId="20.[299,1210,1055,1135]" box="[299,1210,1113,1135]" pageId="20" pageNumber="33">
<emphasis id="48CBEAA25674FFF3FEDFFB92FE3A2129" box="[299,468,1113,1134]" italics="true" pageId="20" pageNumber="33">Thinobiusernesti</emphasis>
BeRNhAueR, 1908: 332. (newsynonyminSChülKe &amp; MAKRANCZY 2011)
</paragraph>
</subSubSection>
<subSubSection id="32A5653B5674FFF2FE93FB58FB9923B8" lastPageId="21" lastPageNumber="34" pageId="20" pageNumber="33" type="description">
<paragraph id="7A0036B05674FFF3FE93FB58FC2F20C9" blockId="20.[298,1251,1171,1423]" pageId="20" pageNumber="33">
<materialsCitation id="CAD73CED5674FFF3FE93FB58FC5320C9" collectionCode="R, ISNB, MNHP" collectorName="I. Sc. &amp; A. Fauvel &amp; Peyerimhoff" country="Algeria" determinerName="Makranczy" elevation="1" location="Thinobiusalgiricus" pageId="20" pageNumber="33" specimenCount="6" typeStatus="lectotype">
Examinedtypematerial –
<emphasis id="48CBEAA25674FFF3FD72FB58FCDE21EF" box="[646,816,1171,1192]" italics="true" pageId="20" pageNumber="33">Thinobiusmicros</emphasis>
Fauvel –
<typeStatus id="A50488125674FFF3FC68FB58FBEB21EF" box="[924,1029,1171,1192]" pageId="20" pageNumber="33" type="lectotype">Lectotype</typeStatus>
(m), bypresentdesignation: “Fréjus \ micros; Fauv. \
<collectionCode id="1CAEAE755674FFF3FD44FB7BFD2D2182" box="[688,707,1200,1221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="33">R</collectionCode>
.
<collectorName id="D74A53665674FFF3FD32FB7BFD012182" box="[710,751,1200,1221]" pageId="20" pageNumber="33">I.Sc.</collectorName>
N.B. 17.479; Coll. etdet.
<collectorName id="D74A53665674FFF3FBF6FB7BFB852182" box="[1026,1131,1199,1221]" pageId="20" pageNumber="33">A. Fauvel</collectorName>
\
<typeStatus id="A50488125674FFF3FB78FB7BFB0D2182" box="[1164,1251,1200,1221]" pageId="20" pageNumber="33" type="syntype">Syntype</typeStatus>
\ m \
<typeStatus id="A50488125674FFF3FE80FB06FE0221A5" box="[372,492,1229,1250]" pageId="20" pageNumber="33" type="lectotype">Lectotypus</typeStatus>
;
<taxonomicName id="BDBF4D335674FFF3FE0DFB07FD8821A5" authorityName="Kiesenwetter" authorityYear="1844" box="[505,614,1228,1250]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="genus">Thinobius</taxonomicName>
; microsFauvel; des. Makranczy, 2001 \
<taxonomicName id="BDBF4D335674FFF3FBD5FB07FB6121A5" authorityName="Kiesenwetter" authorityYear="1844" box="[1057,1167,1228,1250]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="genus">Thinobius</taxonomicName>
;
<taxonomicName id="BDBF4D335674FFF3FB6FFB07FE9A21B8" authority="Fauvel" authorityName="Fauvel" authorityYear="1871" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="species" species="micros">micros Fauvel</taxonomicName>
; det.
<determinerName id="8FDCDA745674FFF3FE5AFB22FDCD21B8" box="[430,547,1257,1279]" pageId="20" pageNumber="33">Makranczy</determinerName>
, 2010” (
<collectionCode id="1CAEAE755674FFF3FD8EFB22FD5B21B8" box="[634,693,1257,1279]" pageId="20" pageNumber="33">ISNB</collectionCode>
);
<typeStatus id="A50488125674FFF3FD3CFB22FC8C21B9" box="[712,866,1257,1279]" pageId="20" pageNumber="33" type="paralectotype">Paralectotypes</typeStatus>
(4): samedataaslectotype (
<quantity id="BD479B555674FFF3FB61FB22FB5C21B8" box="[1173,1202,1257,1279]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="33" unit="m" value="1.0">
<elevation id="F192D1835674FFF3FB61FB22FB5C21B8" box="[1173,1202,1257,1279]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="33" unit="m" value="1.0">1m</elevation>
</quantity>
, 3f,
<collectionCode id="1CAEAE755674FFF3FEDFFACDFE88205B" box="[299,358,1286,1308]" pageId="20" pageNumber="33">ISNB</collectionCode>
);
<emphasis id="48CBEAA25674FFF3FE81FACDFDDE205C" box="[373,560,1286,1307]" italics="true" pageId="20" pageNumber="33">
<location id="7F60606B5674FFF3FE81FACDFDDE205C" LSID="urn:lsid:plazi:treatment:F21687A65674FFF2FDB8FC0FFB9923B8:7F60606B5674FFF3FE81FACDFDDE205C" box="[373,560,1286,1307]" country="Algeria" name="Thinobiusalgiricus" pageId="20" pageNumber="33">Thinobiusalgiricus</location>
</emphasis>
FAuvel –
<typeStatus id="A50488125674FFF3FD61FACDFD10205C" box="[661,766,1286,1307]" pageId="20" pageNumber="33" type="lectotype">Lectotype</typeStatus>
(f), bypresentdesignation: “[
<collectingCountry id="02A876205674FFF3FBCFFACDFB63205C" box="[1083,1165,1286,1308]" name="Algeria" pageId="20" pageNumber="33">Algeria</collectingCountry>
: Djebel] Ouarsenis \ confonduavec; longipennis \ algiricus; Fvl. n. sp.; Désiré [underlined] \ Ty- pus \ Collection; Vauloger \
<typeStatus id="A50488125674FFF3FDAEFA8BFD3C2012" box="[602,722,1344,1365]" pageId="20" pageNumber="33" type="lectotype">Lectotypus</typeStatus>
;
<taxonomicName id="BDBF4D335674FFF3FD2FFAF4FCA62012" authorityName="Kiesenwetter" authorityYear="1844" box="[731,840,1343,1365]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="genus">Thinobius</taxonomicName>
; algiricusFauvel; des. Makranczy, 2010 \
<taxonomicName id="BDBF4D335674FFF3FEB6FA97FE412035" authorityName="Kiesenwetter" authorityYear="1844" box="[322,431,1372,1394]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="genus">Thinobius</taxonomicName>
; microsFauvel; det.
<determinerName id="8FDCDA745674FFF3FD67FA96FCE62035" box="[659,776,1372,1394]" pageId="20" pageNumber="33">Makranczy</determinerName>
, 2010” (coll.
<collectorName id="D74A53665674FFF3FC6CFA96FBF32035" box="[920,1053,1372,1394]" pageId="20" pageNumber="33">Peyerimhoff</collectorName>
,
<collectionCode id="1CAEAE755674FFF3FBDDFA96FB972035" box="[1065,1145,1373,1394]" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="20" pageNumber="33">MNHP</collectionCode>
); [Forthe type of
<taxonomicName id="BDBF4D335674FFF3FE8FFAB2FDCD20C8" authorityName="Bernhauer" authorityYear="1908" box="[379,547,1401,1423]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="species" species="ernesti">
<emphasis id="48CBEAA25674FFF3FE8FFAB2FDCD20C8" box="[379,547,1401,1423]" italics="true" pageId="20" pageNumber="33">Thinobius ernesti</emphasis>
</taxonomicName>
, see Makranczy &amp;
<bibRefCitation id="1E2E4B415674FFF3FD19FAB2FC5D20C9" author="SChulKe, M." box="[749,947,1400,1423]" pageId="20" pageNumber="33" pagination="185 - 193" refId="ref9934" refString="MAKRANCZY, GY. &amp; SChulKe, M. (2001) Typenstudien an den mitteleuropaischen Vertretern der Artengruppe des Thinobius linearis Kraatz, 1857 (Coleoptera, Staphylinidae, Oxy- telinae). Entomologische Blatter fur Biologie und Systematik der Kafer 97 (2): 185 - 193." type="journal article" year="2001">Schülke, 2001: 191</bibRefCitation>
.]
</materialsCitation>
.
</paragraph>
<paragraph id="7A0036B05674FFF2FE93FA78FC5227E8" blockId="20.[298,1251,1458,1797]" lastBlockId="21.[336,1289,406,1493]" lastPageId="21" lastPageNumber="34" pageId="20" pageNumber="33">
Redescription – Measurements (in mm, n=6): HW = 0.20 (0.19–0.20); TW = 0.19 (0.185– 0.20); PW = 0.21 (0.205–0.22); SW = 0.22 (0.21–0.22); AW = 0.24 (0.23–0.24); HL = 0.18 (0.17– 0.19); EL = 0.07 (0.06–0.07); TL = 0.05 (0.05–0.06); PL = 0.17 (0.16–0.175); SL = 0.28 (0.26–0.29); SC = 0.24 (0.23–0.25); FB = 0.61 (0.59–0.64); BL = 1.10 (1.03–1.19). Body (
<figureCitation id="E2842A355674FFF3FBE0F9C2FBBE2358" box="[1044,1104,1545,1567]" captionStart="Figs 1–6" captionStartId="4.[299,344,1713,1735]" captionTargetBox="[305,1244,405,1695]" captionTargetId="figure-18@4.[305,1245,405,1695]" captionTargetPageId="4" captionText="Figs 1–6. Forebodies of Thinobius species: 1 = T. major Kraatz, 1857, 2 = T. angusticeps Fauvel, 1889, 3 = T. procerus Eppelsheim, 1893, 4 = T. bicolor Joy, 1911, 5 = T. petzi Bernhauer, 1908, 6 = T. micros Fauvel, 1871. Scales: 0.07 mm for Fig. 6, 0.10 mm for Figs 2, 4–5, 0.14 mm for Figs 1, 3." figureDoi="http://doi.org/10.5281/zenodo.5732154" httpUri="https://zenodo.org/record/5732154/files/figure.png" pageId="20" pageNumber="33">Fig. 6</figureCitation>
) very slightly bicoloured. Head, pronotumandabdomenslightlyreddishdarkbrown, elytrareddish mediumbrownwithadarkerareabehindscutellumextendingtoshoulders. Legsand mouthpartsmediumbrown, apicesoffemoraandbothendsoftibiaeoftenlighter, together withtarsi; basalpartofantennamuchlighter (yellowish) thanapicalarticles, latteroften significantlyblackish. Forebodywithveryfinecoriaceousmicroreticulation, ratherdull. Punctationonforebodyratherdenseandsofinethathardlydiscernible. Discsofabdominaltergiteswithaparticulartypeoftransversalmicrosculptureandlongitudinalmicrosetationarrangedintheminutetransversalridgesthatcausesaslightiridescenteffect. Pu- bescenceonforebodydepressedandveryshort, fineanddense (onelytrathemostdense), onabdomenapicesoftergiteswitharowoflonghairs, andasparsesetationcomposedof similarbutshorterhairs; abdominalsidesandapexwithafewdarkerandstrongerbristles. Strongdarkbristleonoutersideofsupraantennaltubercleatanteriorborderofeye (thosenearouterposterioredgeofvertexweaklydeveloped), similaronesrightbehind anteriorpronotalcornerandonsideat 3/5 length. Setationanteriorlydirectedonheadand pronotum, directionposterioronelytraandabdomen. Antennaewithlonger, darkerhairs onbasalarticles, legsandmouthpartswithveryshort, notconspicuoussetation, exceptfor thestiff, darkermid-tibialhairsondonesnearapicesoffemora. Sidesofelytrawiththree suchstiffsetaeaboutequallydistributedonlengthofside.
</paragraph>
<paragraph id="7A0036B05675FFF2FE79FD7DFB1E263F" blockId="21.[336,1289,406,1493]" pageId="21" pageNumber="34">
Headslightlywiderthanlong, templesusuallystraight, withagentlecurveonthe side, roundedonposteriorpart. Middleofvertexverygentlyimpressed. Anteriorborder ofneckisstronglymarkedwithashiny, arched, deepgroove. Frontoclypealsutureappear- ingasweakgrooveandadarkline (visibleonlyifheadlighter) connectingsupraantennal tubercles, similarlydarkerspotssituatedatbothsidesonmiddleofvertex. Supraantennal tuberclesmoderatelydeveloped, separatedfromvertexbylongitudinalimpression. An- tennaemoderatelyshort (
<figureCitation id="E2842A355675FFF2FD94FCA8FD47263E" box="[608,681,867,889]" captionStart="Figs 13–26" captionStartId="9.[336,381,1630,1652]" captionTargetBox="[357,1268,405,1603]" captionTargetId="figure-11@9.[356,1268,405,1605]" captionTargetPageId="9" captionText="Figs 13–26. Antennae of Thinobius species: 13 = T. major Kraatz, 1857, 14 = T. angusticeps Fauvel, 1889, 15 = T. procerus Eppelsheim, 1893, 16 = T. bicolor Joy, 1911, 17 = T. petzi Bernhauer, 1908, 18 = T. micros Fauvel, 1871, 19 = T. paramicros sp. n. 20–22 = T. major Kraatz, 1857, male, 20 = sternite VIII, 21 = tergite IX, 22 = sternite IX. 23–24 = T. angusticeps Fauvel, 1889, male, 23 = tergite IX, 24 = sternite IX. 25–26 = T. procerus Eppelsheim, 1893, male, 25 = tergite IX, 26 = sternite IX. Scales: 0.05 mm for Figs 13, 15, 0.06 mm for Figs 14, 16–17, 26, 0.07 mm for Fig." figureDoi="http://doi.org/10.5281/zenodo.5732160" httpUri="https://zenodo.org/record/5732160/files/figure.png" pageId="21" pageNumber="34">Fig. 18</figureCitation>
), 6thantennomereappearingaswideasadjacentones.
</paragraph>
<paragraph id="7A0036B05675FFF2FE79FC4BFC6A21DE" blockId="21.[336,1289,406,1493]" pageId="21" pageNumber="34">Pronotumweaklytransverse, 1.25× widerthanlong, justatinybitwiderthanhead, withonlytracesofimpressionsinanteriorhalfofmiddleofdiscandnearanteriorcor- ners. Anteriorcornersmoderatelyweaklyrounded, posteriorcornersverybroadly, sides almostinarchwithposteriormargin. Pronotalmarginalbeadvisibleonposteriormargin, anteriormarginslightlypulledaheadinmiddlebutwithoutvisiblemarginalbead. Scutellumsmall, ifexposed, shinierthanadjoiningareas. Elytralong, oneelytronabouttwice aslongasbroad, quiteflatandparallel-sided (verygentlydilatingposteriorly). Shoulders moderatelydeveloped. Alongsuturewithouteventracesofmarginalbead; suturalcorners broadlyrounded. Legsofmediumlength, tarsallobessmallandthin, inconspicuous, tibiae fusiform, appearmuchwiderinmiddlethanatends.</paragraph>
<paragraph id="7A0036B05675FFF2FE79FB54FC1921A8" blockId="21.[336,1289,406,1493]" pageId="21" pageNumber="34">Abdomenveryweaklyfusiform, sidesmoreorlessstraightandparallel; widestat 1/3 length, justveryslightlylesswidethanelytraatbroadestpoint. Posteriormarginoftergite VIIwithpalisadefringeandposteriorcornersbroadlyrounded.</paragraph>
<paragraph id="7A0036B05675FFF2FE79FB3DFBBD2025" blockId="21.[336,1289,406,1493]" pageId="21" pageNumber="34">
Primaryandsecondarysexualfeatures: Sexesnotappearingdifferentinregulardor- salview. PosterioredgeoftergiteVIII (similarinbothsexes) gentlyconcave, arched. Male
<figureCitation id="E2842A355675FFF2FEA4FAFBFDBE2002" box="[336,592,1327,1349]" captionStart="Figs 34–40" captionStartId="15.[336,381,1709,1731]" captionTargetBox="[346,1279,406,1683]" captionTargetId="figure-10@15.[345,1280,405,1689]" captionTargetPageId="15" captionText="Figs 34–40. 34 = T. petzi Bernhauer, 1908, male sternite VIII. 35–37 = T. micros Fauvel, 1871, male, 35 = sternite VIII, 36 = tergite IX, 37 = sternite IX. 38–40 = T. paramicros sp. n., male, 38 = sternite VIII, 39 = tergite IX, 40 = sternite IX. Scales: 0.08 mm for Fig. 36, 0.09 mm for Fig." figureDoi="http://doi.org/10.5281/zenodo.5732168" httpUri="https://zenodo.org/record/5732168/files/figure.png" pageId="21" pageNumber="34">sterniteVIIIasinFig. 35</figureCitation>
,
<figureCitation id="E2842A355675FFF2FDADFAFBFCD22002" box="[601,828,1327,1349]" captionStart="Figs 34–40" captionStartId="15.[336,381,1709,1731]" captionTargetBox="[346,1279,406,1683]" captionTargetId="figure-10@15.[345,1280,405,1689]" captionTargetPageId="15" captionText="Figs 34–40. 34 = T. petzi Bernhauer, 1908, male sternite VIII. 35–37 = T. micros Fauvel, 1871, male, 35 = sternite VIII, 36 = tergite IX, 37 = sternite IX. 38–40 = T. paramicros sp. n., male, 38 = sternite VIII, 39 = tergite IX, 40 = sternite IX. Scales: 0.08 mm for Fig. 36, 0.09 mm for Fig." figureDoi="http://doi.org/10.5281/zenodo.5732168" httpUri="https://zenodo.org/record/5732168/files/figure.png" pageId="21" pageNumber="34">tergiteIXasinFig. 36</figureCitation>
,
<figureCitation id="E2842A355675FFF2FCB2FAFBFBDB2002" box="[838,1077,1327,1349]" captionStart="Figs 34–40" captionStartId="15.[336,381,1709,1731]" captionTargetBox="[346,1279,406,1683]" captionTargetId="figure-10@15.[345,1280,405,1689]" captionTargetPageId="15" captionText="Figs 34–40. 34 = T. petzi Bernhauer, 1908, male sternite VIII. 35–37 = T. micros Fauvel, 1871, male, 35 = sternite VIII, 36 = tergite IX, 37 = sternite IX. 38–40 = T. paramicros sp. n., male, 38 = sternite VIII, 39 = tergite IX, 40 = sternite IX. Scales: 0.08 mm for Fig. 36, 0.09 mm for Fig." figureDoi="http://doi.org/10.5281/zenodo.5732168" httpUri="https://zenodo.org/record/5732168/files/figure.png" pageId="21" pageNumber="34">sterniteIXasinFig. 37</figureCitation>
.
<figureCitation id="E2842A355675FFF2FBC8FAE4FE852026" captionStart="Figs 41–53" captionStartId="17.[336,381,1625,1647]" captionText="Figs 41–53. Aedeagi of Thinobius species: 41 = T. major Kraatz, 1857, 42 = T. angusticeps Fauvel, 1889, 43 = T. procerus Eppelsheim, 1893, 44 = T. micros Fauvel, 1871, 45 = T. paramicros sp. n. Femalegenitaltraits: 46–47 = T. angusticeps Fauvel, 1889, neotype, 46 = genitalappendage, 47 = spermatheca. 48–50 = T. micros Fauvel, 1871, 48 = genitalappendage, 49–50 = spermathecae. 51–53 = T. paramicros sp. n., 51 = genitalappendage, 52–53 = spermathecae. Scales: 0.05 mm for Figs 44, 48–50, 52–53, 0.065 mm for Figs 45, 51, 0.08 mm for Fig. 42, 0.10" pageId="21" pageNumber="34">AedeagusasinFig. 44</figureCitation>
.
<figureCitation id="E2842A355675FFF2FE80FA86FCFE2026" box="[372,784,1356,1378]" captionStart="Figs 41–53" captionStartId="17.[336,381,1625,1647]" captionText="Figs 41–53. Aedeagi of Thinobius species: 41 = T. major Kraatz, 1857, 42 = T. angusticeps Fauvel, 1889, 43 = T. procerus Eppelsheim, 1893, 44 = T. micros Fauvel, 1871, 45 = T. paramicros sp. n. Femalegenitaltraits: 46–47 = T. angusticeps Fauvel, 1889, neotype, 46 = genitalappendage, 47 = spermatheca. 48–50 = T. micros Fauvel, 1871, 48 = genitalappendage, 49–50 = spermathecae. 51–53 = T. paramicros sp. n., 51 = genitalappendage, 52–53 = spermathecae. Scales: 0.05 mm for Figs 44, 48–50, 52–53, 0.065 mm for Figs 45, 51, 0.08 mm for Fig. 42, 0.10" pageId="21" pageNumber="34">FemalegenitalappendageasinFig. 48</figureCitation>
,
<figureCitation id="E2842A355675FFF2FCEEFA86FBA12025" box="[794,1103,1356,1378]" captionStart="Figs 41–53" captionStartId="17.[336,381,1625,1647]" captionText="Figs 41–53. Aedeagi of Thinobius species: 41 = T. major Kraatz, 1857, 42 = T. angusticeps Fauvel, 1889, 43 = T. procerus Eppelsheim, 1893, 44 = T. micros Fauvel, 1871, 45 = T. paramicros sp. n. Femalegenitaltraits: 46–47 = T. angusticeps Fauvel, 1889, neotype, 46 = genitalappendage, 47 = spermatheca. 48–50 = T. micros Fauvel, 1871, 48 = genitalappendage, 49–50 = spermathecae. 51–53 = T. paramicros sp. n., 51 = genitalappendage, 52–53 = spermathecae. Scales: 0.05 mm for Figs 44, 48–50, 52–53, 0.065 mm for Figs 45, 51, 0.08 mm for Fig. 42, 0.10" pageId="21" pageNumber="34">spermathecaasinFigs 49–50</figureCitation>
.
</paragraph>
<paragraph id="7A0036B05675FFF2FE79FAA2FCDF2092" blockId="21.[336,1289,406,1493]" pageId="21" pageNumber="34">Distribution – Obviouslyaveryrareandlocalspecies, possiblyextinctfrommost ofitsformerrange. ThisisindicatedbyitsoldrecordfromAustria, whilenotcollectedin CentralEuropeinthepastcentury. TheNorthernAfricanoccurrencesuggeststhatitoriginallyhadcircum-mediterraneandistribution.</paragraph>
<paragraph id="7A0036B05675FFF2FE79FA30FB9923B8" blockId="21.[336,1291,1530,1792]" pageId="21" pageNumber="34">
Remarks – Theaedeagusdrawingof “
<taxonomicName id="BDBF4D335675FFF2FCA2FA30FC552353" authorityName="Bernhauer" authorityYear="1908" box="[854,955,1531,1556]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="34" phylum="Arthropoda" rank="species" species="ernesti">
<emphasis id="48CBEAA25675FFF2FCA2FA30FC552353" box="[854,955,1531,1556]" italics="true" pageId="21" pageNumber="34">T. ernesti</emphasis>
</taxonomicName>
” inMAKRANCZYandSChül- Ke (2001) wasmadebasedonitspoorlypreservedholotypewiththeaedeagus ratherdistorted. Theherepresentednewillustrationisbasedonmorespecimens, improved, soshouldsubstitutefortheearlierfigure. Therearestillonly therespectivetypematerials (morethanahundredyearsold) availableforthis species. SincePeYeRimhoff’ s (1931) identification (basedonatypestudy) of “
<taxonomicName id="BDBF4D335675FFF2FB07F968FE5E2399" authorityName="Fauvel. Thefollowingnewsynonymiesareproposed" authorityYear="1898" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="34" phylum="Arthropoda" rank="species" species="algiricus">
<emphasis id="48CBEAA25675FFF2FB07F968FE5E2399" italics="true" pageId="21" pageNumber="34">T. algiricus</emphasis>
</taxonomicName>
” isincorrectandinfactreferstoaverysimilarbutsignificantlylarger, clearlydistinctspecies, thelatterhastobedescribedasnewbelow.
</paragraph>
</subSubSection>
</treatment>
</document>
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