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<document id="6BACA11955A812C0520F2D02ED9B4CED" ID-DOI="10.11646/zootaxa.5296.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984382" ID-ZooBank="2EE722D2-463B-463F-8EA3-E9A099E85154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685428400234" checkinUser="plazi" docAuthor="Schawaller, Wolfgang &amp; Bellersheim, Aron" docDate="2023" docId="CB5E87F7FA75FFE2FF69FA5080CCC1C9" docLanguage="en" docName="zootaxa.5296.4.3.pdf" docOrigin="Zootaxa 5296 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laena faillei Schawaller &amp; Bellersheim 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="543" masterDocId="3767FF8FFA74FFE1FFFEFFCC8013C040" masterDocTitle="New species and records of Laena Dejean (Coleoptera: Tenebrionidae: Lagriinae) from Vietnam, southeastern China, and Korea" masterLastPageNumber="550" masterPageNumber="540" pageNumber="541" updateTime="1685429945776" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D1436418F399FBD2DC5738B77B981164" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F7D0DD044E84BCC628BD137FD07F184">
<mods:title id="2B22D5678E45F03C8FC2BEA62AD98519">New species and records of Laena Dejean (Coleoptera: Tenebrionidae: Lagriinae) from Vietnam, southeastern China, and Korea</mods:title>
</mods:titleInfo>
<mods:name id="BDE173866AB79367B258745E6E7DF503" type="personal">
<mods:role id="BE29814AFFA8EF2E6F4386277CCB0623">
<mods:roleTerm id="8E8560A5A3A54372875501DA091FA51F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A8C73210C797842CC10718A05D45F83">Schawaller, Wolfgang</mods:namePart>
<mods:nameIdentifier id="70B9347944915E759F12DEBE86B784FC" type="ORCID">0000-0003-1482-7386</mods:nameIdentifier>
<mods:affiliation id="99D9667EFAD173ACFC500B60A13D81B5">Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany &amp; schawaller. ehrenamt @ smns-bw. de; https: // orcid. org / 0000 - 0003 - 1482 - 7386</mods:affiliation>
<mods:nameIdentifier id="F7D0DC263B4888B71C923952957F610E" type="email">schawaller.ehrenamt@smns-bw.de</mods:nameIdentifier>
</mods:name>
<mods:name id="96E399DD4266ABE632FE45E822424E41" type="personal">
<mods:role id="65D8F15BE7A5C2C6A59792A525A9569B">
<mods:roleTerm id="BA83F79D0FC8514565EF31029342A76F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E1A633CFB2210655BC930B22578DFFE0">Bellersheim, Aron</mods:namePart>
<mods:nameIdentifier id="7A2DAE383F85C86DEEDA4FA6F8361520" type="ORCID">0000-0002-5607-328X</mods:nameIdentifier>
<mods:affiliation id="8533967DA4D3268CA41D3B239F9635DA">Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany &amp; bellersheim. aron @ smns-bw. de; https: // orcid. org / 0000 - 0002 - 5607 - 328 X</mods:affiliation>
<mods:nameIdentifier id="255890C8B64744ACD48675D6C94B8FC8" type="email">bellersheim.aron@smns-bw.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="014F59AB8BEB94B05904A094F4DA18B9">text</mods:typeOfResource>
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<mods:titleInfo id="D3ABEAAC6299EA356FF4EA8A1F6C4FEB">
<mods:title id="1E61222A37769C6E29ED35B6A6ED5596">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4C2E826E116F5936E3A9ABED7B64B2D9">
<mods:date id="5988BA84062A4F960FE045BB2F7F115D">2023</mods:date>
<mods:detail id="DEA9BBC6C337644920C13E1EC58FCB3B" type="pubDate">
<mods:number id="AD5E9CC6AF560F15B9E1D186EAEFEC51">2023-05-30</mods:number>
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<mods:number id="298EB1F53D4CC91621E92414F324537D">5296</mods:number>
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<mods:number id="979D304F0B2B4A47C787B820EA513F59">4</mods:number>
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<mods:url id="9A6FB64C393874D26EFC9DD0FB10987C">http://dx.doi.org/10.11646/zootaxa.5296.4.3</mods:url>
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<mods:classification id="6656A7609DEA171076403667638B2CC2">journal article</mods:classification>
<mods:identifier id="104994FF5AD54B9997F36464B611936F" type="DOI">10.11646/zootaxa.5296.4.3</mods:identifier>
<mods:identifier id="54FBC43C1B890E586946C3B15B9159CB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B872750CAF250F9F4F3AA2CF256DC4ED" type="Zenodo-Dep">7984382</mods:identifier>
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<treatment id="CB5E87F7FA75FFE2FF69FA5080CCC1C9" ID-DOI="http://doi.org/10.5281/zenodo.7984221" ID-Zenodo-Dep="7984221" LSID="urn:lsid:plazi:treatment:CB5E87F7FA75FFE2FF69FA5080CCC1C9" httpUri="http://treatment.plazi.org/id/CB5E87F7FA75FFE2FF69FA5080CCC1C9" lastPageId="3" lastPageNumber="543" pageId="1" pageNumber="541">
<subSubSection id="0BED656AFA75FFE0FF69FA508168C5F6" box="[151,379,1435,1462]" pageId="1" pageNumber="541" type="nomenclature">
<paragraph id="434836E1FA75FFE0FF69FA508168C5F6" blockId="1.[151,379,1435,1499]" box="[151,379,1435,1462]" pageId="1" pageNumber="541">
<heading id="1800818DFA75FFE0FF69FA508168C5F6" bold="true" box="[151,379,1435,1462]" fontSize="11" level="1" pageId="1" pageNumber="541" reason="1">
<emphasis id="7183EAF3FA75FFE0FF69FA508168C5F6" bold="true" box="[151,379,1435,1462]" pageId="1" pageNumber="541">
<taxonomicName id="84F74D62FA75FFE0FF69FA508122C5F6" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[151,305,1435,1462]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="541" phylum="Arthropoda" rank="species" species="faillei" status="sp. n.">
<emphasis id="7183EAF3FA75FFE0FF69FA508122C5F6" bold="true" box="[151,305,1435,1462]" italics="true" pageId="1" pageNumber="541">Laena faillei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AB05788FA75FFE0FEC6FA508168C5F6" box="[312,379,1436,1462]" pageId="1" pageNumber="541" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA75FFE0FF69FA0C810AC59B" box="[151,281,1472,1499]" pageId="1" pageNumber="541" type="description">
<paragraph id="434836E1FA75FFE0FF69FA0C810AC59B" blockId="1.[151,379,1435,1499]" box="[151,281,1472,1499]" pageId="1" pageNumber="541">
(
<figureCitation id="DBCC2A64FA75FFE0FF61FA0C80F5C59A" box="[159,230,1472,1498]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1801,1825]" captionTargetBox="[192,1388,207,1748]" captionTargetId="figure-17@2.[151,1436,179,1775]" captionTargetPageId="2" captionText="FIGURES 1–7. Dorsal view of Laena species from Vietnam and southeastern China. 1. L. faillei sp. n., ♀ paratype, Vietnam, SMNS. 2. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 3. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 4. L. hajeki sp. n., ♁ paratype, Guangdong, SMNS. 5. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 6. L. logshengica sp. n., ♁ paratype, Guangxi, NMPC. 7. L. honba sp. n., ♁ holotype, Vietnam, SMNS. Scale line: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984384" httpUri="https://zenodo.org/record/7984384/files/figure.png" pageId="1" pageNumber="541">Figs 1</figureCitation>
,
<figureCitation id="DBCC2A64FA75FFE0FF0AFA0C8102C59B" box="[244,273,1472,1499]" captionStart="FIGURES 11–20" captionStartId="5.[152,270,1254,1278]" captionTargetBox="[152,1435,185,1226]" captionTargetId="figure-20@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURES 11–20. Aedeagi of Laena species from Vietnam and southeastern China. 11. L. tramtonica sp. n., ♁ holotype, Vietnam, NHMW. 12. L. tamdaoensis Masumoto, 1996, non-type, Vietnam, NHMW. 13. L. fanjingshanana Ren &amp; Hua, 2006, ♁ non-type, Guangxi, SMNS. 14. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 15. L. faillei sp. n., ♁ holotype, Vietnam, SMNS. 16. L. logshengica sp. n., ♁ holotype, Guangxi, NMPC. 17. L. honba sp. n., ♁ holotype, Vietnam, SMNS. 18. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 19. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 20. L. hajeki sp. n., ♁ holotype, Guangdong, NMPC. Scale line: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984388" httpUri="https://zenodo.org/record/7984388/files/figure.png" pageId="1" pageNumber="541">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA75FFE0FF69F9C483A4C6CE" pageId="1" pageNumber="541" type="materials_examined">
<paragraph id="434836E1FA75FFE0FF69F9C483A4C6CE" blockId="1.[151,1437,1544,2039]" pageId="1" pageNumber="541">
<emphasis id="7183EAF3FA75FFE0FF69F9C4815BC662" bold="true" box="[151,328,1544,1570]" pageId="1" pageNumber="541">Type material.</emphasis>
<materialsCitation id="F39F3CBCFA75FFE0FEACF9C48202C606" collectingDate="2013-11-14" collectionCode="SMNS" collectorName="Hon Ba NR &amp; B. Fontaine &amp; O. Gargomini" country="Vietnam" county="Yersin" elevation="1500" location="Cam Lam" municipality="Suoi Cat" pageId="1" pageNumber="541" specimenCount="1" typeStatus="holotype">
<emphasis id="7183EAF3FA75FFE0FEACF9C481DBC662" bold="true" box="[338,456,1544,1570]" pageId="1" pageNumber="541">
<typeStatus id="9C4C8843FA75FFE0FEACF9C481D0C662" box="[338,451,1544,1570]" pageId="1" pageNumber="541">Holotype</typeStatus>
:
</emphasis>
♁,
<collectingCountry id="3BE07671FA75FFE0FE06F9C48249C662" box="[504,602,1544,1570]" name="Vietnam" pageId="1" pageNumber="541">Vietnam</collectingCountry>
,
<location id="4628603AFA75FFE0FD96F9C482CEC663" LSID="urn:lsid:plazi:treatment:CB5E87F7FA75FFE2FF69FA5080CCC1C9:4628603AFA75FFE0FD96F9C482CEC663" box="[616,733,1544,1571]" country="Vietnam" county="Yersin" municipality="Suoi Cat" name="Cam Lam" pageId="1" pageNumber="541">Cam Lam</location>
,
<collectingMunicipality id="A32CAC9BFA75FFE0FD12F9C48340C662" box="[748,851,1544,1570]" pageId="1" pageNumber="541">Suoi Cat</collectingMunicipality>
,
<collectorName id="EE025337FA75FFE0FC9FF9C583FCC663" box="[865,1007,1545,1571]" pageId="1" pageNumber="541">Hon Ba NR</collectorName>
,
<collectingCounty id="AA294E6DFA75FFE0FC02F9C48454C662" box="[1020,1095,1544,1570]" pageId="1" pageNumber="541">Yersin</collectingCounty>
,
<quantity id="840F9B04FA75FFE0FBABF9C484BCC663" box="[1109,1199,1544,1571]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="541" unit="m" value="1500.0">
<elevation id="C8DAD1D2FA75FFE0FBABF9C484BCC663" box="[1109,1199,1544,1571]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="541" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="37491021FA75FFE0FB40F9C48552C662" box="[1214,1345,1544,1570]" pageId="1" pageNumber="541" value="2013-11-14">
<collectingDate id="270DE9C9FA75FFE0FB40F9C48552C662" box="[1214,1345,1544,1570]" pageId="1" pageNumber="541" value="2013-11-14">14.XI.2013</collectingDate>
</date>
, leg.
<collectorName id="EE025337FA75FFE0FA7DF9C580EAC606" pageId="1" pageNumber="541">B. Fontaine</collectorName>
&amp;
<collectorName id="EE025337FA75FFE0FEE4F9E081A4C606" box="[282,439,1580,1606]" pageId="1" pageNumber="541">O. Gargomini</collectorName>
,
<collectionCode id="25E6AE24FA75FFE0FE3FF9E08202C606" box="[449,529,1580,1606]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="541">SMNS</collectionCode>
</materialsCitation>
.
<materialsCitation id="F39F3CBCFA75FFE0FDE4F9E08440C606" box="[538,1107,1580,1607]" collectingDate="2013-11-14" collectionCode="SMNS" collectorName="Hon Ba NR &amp; B. Fontaine &amp; O. Gargomini" country="Vietnam" county="Yersin" elevation="1500" location="Same" municipality="Suoi Cat" pageId="1" pageNumber="541" specimenCount="3" typeStatus="paratype">
<emphasis id="7183EAF3FA75FFE0FDE4F9E0828AC606" bold="true" box="[538,665,1580,1606]" pageId="1" pageNumber="541">
<typeStatus id="9C4C8843FA75FFE0FDE4F9E08287C606" box="[538,660,1580,1606]" pageId="1" pageNumber="541">Paratypes</typeStatus>
.
</emphasis>
<location id="4628603AFA75FFE0FD60F9E082CFC606" LSID="urn:lsid:plazi:treatment:CB5E87F7FA75FFE2FF69FA5080CCC1C9:4628603AFA75FFE0FD60F9E082CFC606" box="[670,732,1580,1606]" name="Same" pageId="1" pageNumber="541">Same</location>
data as for holotype,
<specimenCount id="55F1FD68FA75FFE0FC37F9E083EDC607" box="[969,1022,1580,1607]" pageId="1" pageNumber="541" type="generic">3 ex.</specimenCount>
<collectionCode id="25E6AE24FA75FFE0FBFAF9E08440C606" box="[1028,1107,1580,1606]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="541">SMNS</collectionCode>
</materialsCitation>
,
<materialsCitation id="F39F3CBCFA75FFE0FBA3F9E084C9C606" box="[1117,1242,1580,1607]" collectingDate="2013-11-14" collectionCode="IEBH" collectorName="Hon Ba NR &amp; B. Fontaine &amp; O. Gargomini" country="Vietnam" county="Yersin" elevation="1500" isTailEnumeration="true" location="Same" municipality="Suoi Cat" pageId="1" pageNumber="541" specimenCount="1" typeStatus="paratype">
<specimenCount id="55F1FD68FA75FFE0FBA3F9E08481C607" box="[1117,1170,1580,1607]" pageId="1" pageNumber="541" type="generic">1 ex.</specimenCount>
<collectionCode id="25E6AE24FA75FFE0FB66F9E084C9C606" box="[1176,1242,1580,1606]" pageId="1" pageNumber="541">IEBH</collectionCode>
</materialsCitation>
.—
<materialsCitation id="F39F3CBCFA75FFE0FB04F9E08445C62A" collectingDate="2013-11-13" collectionCode="SMNS" collectorName="Hon Ba NR &amp; A. Faille" country="Vietnam" county="Yersin" elevation="1500" location="Cam Lam" municipality="Suoi Cat" pageId="1" pageNumber="541" specimenCount="1" typeStatus="paratype">
<collectingCountry id="3BE07671FA75FFE0FB04F9E0854EC606" box="[1274,1373,1580,1606]" name="Vietnam" pageId="1" pageNumber="541">Vietnam</collectingCountry>
,
<location id="4628603AFA75FFE0FA99F9E080DEC62B" LSID="urn:lsid:plazi:treatment:CB5E87F7FA75FFE2FF69FA5080CCC1C9:4628603AFA75FFE0FA99F9E080DEC62B" country="Vietnam" county="Yersin" municipality="Suoi Cat" name="Cam Lam" pageId="1" pageNumber="541">Cam Lam</location>
,
<collectingMunicipality id="A32CAC9BFA75FFE0FF27F99C812CC62A" box="[217,319,1616,1642]" pageId="1" pageNumber="541">Suoi Cat</collectingMunicipality>
,
<collectorName id="EE025337FA75FFE0FEB5F99D81C7C62B" box="[331,468,1617,1643]" pageId="1" pageNumber="541">Hon Ba NR</collectorName>
,
<collectingCounty id="AA294E6DFA75FFE0FE21F99C8239C62A" box="[479,554,1616,1642]" pageId="1" pageNumber="541">Yersin</collectingCounty>
,
<quantity id="840F9B04FA75FFE0FDCBF99C829EC62B" box="[565,653,1616,1643]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="541" unit="m" value="1500.0">
<elevation id="C8DAD1D2FA75FFE0FDCBF99C829EC62B" box="[565,653,1616,1643]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="541" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="37491021FA75FFE0FD64F99C830FC62A" box="[666,796,1616,1642]" pageId="1" pageNumber="541" value="2013-11-13">
<collectingDate id="270DE9C9FA75FFE0FD64F99C830FC62A" box="[666,796,1616,1642]" pageId="1" pageNumber="541" value="2013-11-13">13.XI.2013</collectingDate>
</date>
, leg.
<collectorName id="EE025337FA75FFE0FCA6F99C83ACC62A" box="[856,959,1616,1642]" pageId="1" pageNumber="541">A. Faille</collectorName>
,
<specimenCount id="55F1FD68FA75FFE0FC34F99C8411C62B" box="[970,1026,1616,1643]" pageId="1" pageNumber="541" type="generic">1 ex.</specimenCount>
<collectionCode id="25E6AE24FA75FFE0FBF4F99C8445C62A" box="[1034,1110,1616,1642]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="541">SMNS</collectionCode>
</materialsCitation>
.—
<materialsCitation id="F39F3CBCFA75FFE0FB89F99C83A0C6CE" collectingDate="2013-11-19" collectionCode="SMNS" collectorName="Hon Boa NR &amp; A. Faille" country="Vietnam" elevation="1030" location="Khan Hoa" municipality="Suoi Cat" pageId="1" pageNumber="541" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="3BE07671FA75FFE0FB89F99C84C8C62A" box="[1143,1243,1616,1642]" name="Vietnam" pageId="1" pageNumber="541">Vietnam</collectingCountry>
,
<location id="4628603AFA75FFE0FB19F99C854FC62B" LSID="urn:lsid:plazi:treatment:CB5E87F7FA75FFE2FF69FA5080CCC1C9:4628603AFA75FFE0FB19F99C854FC62B" box="[1255,1372,1616,1643]" country="Vietnam" municipality="Suoi Cat" name="Khan Hoa" pageId="1" pageNumber="541">Khan Hoa</location>
,
<collectingMunicipality id="A32CAC9BFA75FFE0FA97F99C80D2C6CE" pageId="1" pageNumber="541">Suoi Cat</collectingMunicipality>
,
<collectorName id="EE025337FA75FFE0FF32F9B98172C6CF" box="[204,353,1653,1679]" pageId="1" pageNumber="541">Hon Boa NR</collectorName>
,
<quantity id="840F9B04FA75FFE0FE92F9B881E9C6CF" box="[364,506,1652,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.03" metricValueMax="1.16" metricValueMin="0.9" pageId="1" pageNumber="541" unit="m" value="1030.0" valueMax="1160.0" valueMin="900.0">
<elevation id="C8DAD1D2FA75FFE0FE92F9B881E9C6CF" box="[364,506,1652,1679]" metricMagnitude="3" metricUnit="m" metricValue="1.03" metricValueMax="1.16" metricValueMin="0.9" pageId="1" pageNumber="541" unit="m" value="1030.0" valueMax="1160.0" valueMin="900.0">900–1160 m</elevation>
</quantity>
,
<date id="37491021FA75FFE0FDF8F9B8829BC6CE" box="[518,648,1652,1678]" pageId="1" pageNumber="541" value="2013-11-19">
<collectingDate id="270DE9C9FA75FFE0FDF8F9B8829BC6CE" box="[518,648,1652,1678]" pageId="1" pageNumber="541" value="2013-11-19">19.XI.2013</collectingDate>
</date>
, leg.
<collectorName id="EE025337FA75FFE0FD3CF9B88334C6CE" box="[706,807,1652,1678]" pageId="1" pageNumber="541">A. Faille</collectorName>
,
<specimenCount id="55F1FD68FA75FFE0FCCCF9B8834FC6CF" box="[818,860,1652,1679]" pageId="1" pageNumber="541" type="female">1 ♀</specimenCount>
<collectionCode id="25E6AE24FA75FFE0FC9DF9B883A0C6CE" box="[867,947,1652,1678]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="541">SMNS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA75FFE0FF39F95483F0C7B7" pageId="1" pageNumber="541" type="description">
<paragraph id="434836E1FA75FFE0FF39F95483F0C7B7" blockId="1.[151,1437,1544,2039]" pageId="1" pageNumber="541">
<emphasis id="7183EAF3FA75FFE0FF39F954814AC6F2" bold="true" box="[199,345,1688,1714]" pageId="1" pageNumber="541">Description.</emphasis>
Body length 4.0–
<quantity id="840F9B04FA75FFE0FDDEF9548265C6F3" box="[544,630,1688,1715]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="541" unit="mm" value="4.8">4.8 mm</quantity>
, dorsal surface dark brownish with a feeble greenish lustre, legs distictly lighter. Eyes not prominent. Pronotum trapezoid, widest across anterior angles, posterior margin distinctly narrower than the anterior one; with large separated punctures, distance 0.5–3 puncture diameters, a few punctures laterally with long erect setae; surface convex and shining, disc in middle without impression(s), lateral margins distinctly beaded; posterior corners rounded; prothoracic hypomera with similar punctures as on pronotum but without visible setation. Elytra elongate oval with rows of punctures in feeble striae, punctures of rows similar as punctures on pronotum, some punctures laterally and at elytral declivity with long erect setae; intervals with very few glabrous punctures, interval 9 with 5 distinct setiferous pores, intervals convex, external intervals stronger convex and nearly keel-like; surface shining.Apex of elytra prolonged.All femora without teeth or sharp angles. Tibiae simple, without peculiarities. Apicale of aedeagus long triangular with acute apex (
<figureCitation id="DBCC2A64FA75FFE0FC7CF81083C5C7B7" box="[898,982,2012,2039]" captionStart="FIGURES 11–20" captionStartId="5.[152,270,1254,1278]" captionTargetBox="[152,1435,185,1226]" captionTargetId="figure-20@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURES 11–20. Aedeagi of Laena species from Vietnam and southeastern China. 11. L. tramtonica sp. n., ♁ holotype, Vietnam, NHMW. 12. L. tamdaoensis Masumoto, 1996, non-type, Vietnam, NHMW. 13. L. fanjingshanana Ren &amp; Hua, 2006, ♁ non-type, Guangxi, SMNS. 14. L. lianhua sp. n., ♁ holotype, Jiangxi, SMNS. 15. L. faillei sp. n., ♁ holotype, Vietnam, SMNS. 16. L. logshengica sp. n., ♁ holotype, Guangxi, NMPC. 17. L. honba sp. n., ♁ holotype, Vietnam, SMNS. 18. L. brunkei sp. n., ♁ holotype, Vietnam, NHMW. 19. L. hainanica sp. n., ♁ holotype, Hainan, NMPC. 20. L. hajeki sp. n., ♁ holotype, Guangdong, NMPC. Scale line: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7984388" httpUri="https://zenodo.org/record/7984388/files/figure.png" pageId="1" pageNumber="541">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="17886669FA76FFE3FF66F8C58414C7CD" ID-DOI="http://doi.org/10.5281/zenodo.7984384" ID-Zenodo-Dep="7984384" httpUri="https://zenodo.org/record/7984384/files/figure.png" pageId="2" pageNumber="542" startId="2.[152,270,1801,1825]" targetBox="[192,1388,207,1748]" targetPageId="2" targetType="figure">
<paragraph id="434836E1FA76FFE3FF66F8C58414C7CD" blockId="2.[151,1437,1801,1936]" pageId="2" pageNumber="542">
<emphasis id="7183EAF3FA76FFE3FF66F8C58152C761" bold="true" box="[152,321,1801,1825]" pageId="2" pageNumber="542">FIGURES 1–7.</emphasis>
Dorsal view of
<taxonomicName id="84F74D62FA76FFE3FE1BF8C68267C761" box="[485,628,1801,1825]" pageId="2" pageNumber="542">
<emphasis id="7183EAF3FA76FFE3FE1BF8C68237C761" box="[485,548,1802,1825]" italics="true" pageId="2" pageNumber="542">Laena</emphasis>
species
</taxonomicName>
from Vietnam and southeastern China.
<emphasis id="7183EAF3FA76FFE3FBF9F8C58409C761" bold="true" box="[1031,1050,1801,1825]" pageId="2" pageNumber="542">1.</emphasis>
<taxonomicName id="84F74D62FA76FFE3FBDFF8C68464C761" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[1057,1143,1801,1825]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="542" phylum="Arthropoda" rank="species" species="faillei" status="sp. n.">
<emphasis id="7183EAF3FA76FFE3FBDFF8C68464C761" box="[1057,1143,1801,1825]" italics="true" pageId="2" pageNumber="542">L. faillei</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA76FFE3FB83F8C684A5C762" bold="true" box="[1149,1206,1802,1826]" pageId="2" pageNumber="542">
<taxonomicNameLabel id="6AB05788FA76FFE3FB83F8C684A5C762" box="[1149,1206,1802,1826]" pageId="2" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♀ paratype, Vietnam, SMNS.
<emphasis id="7183EAF3FA76FFE3FF14F8E180EEC705" bold="true" box="[234,253,1837,1861]" pageId="2" pageNumber="542">2.</emphasis>
<taxonomicName id="84F74D62FA76FFE3FEFBF8E2817DC705" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[261,366,1837,1861]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="542" phylum="Arthropoda" rank="species" species="brunkei" status="sp. n.">
<emphasis id="7183EAF3FA76FFE3FEFBF8E2817DC705" box="[261,366,1837,1861]" italics="true" pageId="2" pageNumber="542">L. brunkei</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA76FFE3FE88F8E281A3C706" bold="true" box="[374,432,1838,1862]" pageId="2" pageNumber="542">
<taxonomicNameLabel id="6AB05788FA76FFE3FE88F8E281A3C706" box="[374,432,1838,1862]" pageId="2" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♁ holotype, Vietnam, NHMW.
<emphasis id="7183EAF3FA76FFE3FCFCF8E18306C705" bold="true" box="[770,789,1837,1861]" pageId="2" pageNumber="542">3.</emphasis>
<taxonomicName id="84F74D62FA76FFE3FCE2F8E28395C705" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[796,902,1837,1861]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="542" phylum="Arthropoda" rank="species" species="lianhua" status="sp. n.">
<emphasis id="7183EAF3FA76FFE3FCE2F8E28395C705" box="[796,902,1837,1861]" italics="true" pageId="2" pageNumber="542">L. lianhua</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA76FFE3FC70F8E283DBC706" bold="true" box="[910,968,1838,1862]" pageId="2" pageNumber="542">
<taxonomicNameLabel id="6AB05788FA76FFE3FC70F8E283DBC706" box="[910,968,1838,1862]" pageId="2" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♁ holotype, Jiangxi, SMNS.
<emphasis id="7183EAF3FA76FFE3FAFCF8E18507C705" bold="true" box="[1282,1300,1837,1861]" pageId="2" pageNumber="542">4.</emphasis>
<taxonomicName id="84F74D62FA76FFE3FAE2F8E28565C705" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[1308,1398,1837,1861]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="542" phylum="Arthropoda" rank="species" species="hajeki" status="sp. n.">
<emphasis id="7183EAF3FA76FFE3FAE2F8E28565C705" box="[1308,1398,1837,1861]" italics="true" pageId="2" pageNumber="542">L. hajeki</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA76FFE3FA80F8E280A1C72A" bold="true" pageId="2" pageNumber="542">
<taxonomicNameLabel id="6AB05788FA76FFE3FA80F8E280BFC72A" pageId="2" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
♁ paratype, Guangdong, SMNS. 5.
<taxonomicName id="84F74D62FA76FFE3FDD7F89E82B9C729" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[553,682,1873,1897]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="542" phylum="Arthropoda" rank="species" species="hainanica" status="sp. n.">
<emphasis id="7183EAF3FA76FFE3FDD7F89E82B9C729" box="[553,682,1873,1897]" italics="true" pageId="2" pageNumber="542">L. hainanica</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA76FFE3FD4FF89E82F8C72A" bold="true" box="[689,747,1874,1898]" pageId="2" pageNumber="542">
<taxonomicNameLabel id="6AB05788FA76FFE3FD4FF89E82F8C72A" box="[689,747,1874,1898]" pageId="2" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♁ holotype, Hainan, NMPC.
<emphasis id="7183EAF3FA76FFE3FBDBF89D842BC729" bold="true" box="[1061,1080,1873,1897]" pageId="2" pageNumber="542">6.</emphasis>
<taxonomicName id="84F74D62FA76FFE3FBBEF89E84C6C729" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[1088,1237,1873,1897]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="542" phylum="Arthropoda" rank="species" species="logshengica" status="sp. n.">
<emphasis id="7183EAF3FA76FFE3FBBEF89E84C6C729" box="[1088,1237,1873,1897]" italics="true" pageId="2" pageNumber="542">L. logshengica</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA76FFE3FB23F89E8504C72A" bold="true" box="[1245,1303,1874,1898]" pageId="2" pageNumber="542">
<taxonomicNameLabel id="6AB05788FA76FFE3FB23F89E8504C72A" box="[1245,1303,1874,1898]" pageId="2" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♁ paratype, Guangxi, NMPC.
<emphasis id="7183EAF3FA76FFE3FEAEF8BA8171C7CE" bold="true" box="[336,354,1910,1934]" pageId="2" pageNumber="542">7.</emphasis>
<taxonomicName id="84F74D62FA76FFE3FE97F8BA81D0C7CD" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[361,451,1909,1933]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="542" phylum="Arthropoda" rank="species" species="honba" status="sp. n.">
<emphasis id="7183EAF3FA76FFE3FE97F8BA81D0C7CD" box="[361,451,1909,1933]" italics="true" pageId="2" pageNumber="542">L. honba</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA76FFE3FE37F8BA8211C7CE" bold="true" box="[457,514,1910,1934]" pageId="2" pageNumber="542">
<taxonomicNameLabel id="6AB05788FA76FFE3FE37F8BA8211C7CE" box="[457,514,1910,1934]" pageId="2" pageNumber="542" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ♁ holotype, Vietnam, SMNS. Scale line: 3.0 mm.
</paragraph>
</caption>
<subSubSection id="0BED656AFA77FFE2FF39FF5A8445C101" pageId="3" pageNumber="543" type="diagnosis">
<paragraph id="434836E1FA77FFE2FF39FF5A8445C101" blockId="3.[151,1436,150,393]" pageId="3" pageNumber="543">
<emphasis id="7183EAF3FA77FFE2FF39FF5A8151C0F0" bold="true" box="[199,322,150,176]" pageId="3" pageNumber="543">Diagnosis.</emphasis>
<taxonomicName id="84F74D62FA77FFE2FEB5FF5B81CFC0F1" authority="Schawaller &amp; Bellersheim, 2023" authorityName="Schawaller &amp; Bellersheim" authorityYear="2023" box="[331,476,151,177]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="faillei" status="sp. n.">
<emphasis id="7183EAF3FA77FFE2FEB5FF5B81CFC0F1" box="[331,476,151,177]" italics="true" pageId="3" pageNumber="543">Laena faillei</emphasis>
</taxonomicName>
<emphasis id="7183EAF3FA77FFE2FE18FF5B823BC0F1" bold="true" box="[486,552,151,177]" pageId="3" pageNumber="543">
<taxonomicNameLabel id="6AB05788FA77FFE2FE18FF5B823BC0F1" box="[486,552,151,177]" pageId="3" pageNumber="543" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
shares with
<taxonomicName id="84F74D62FA77FFE2FD43FF5B8431C0F1" authority="Masumoto, 1995" authorityName="Masumoto" authorityYear="1995" box="[701,1058,150,177]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="7183EAF3FA77FFE2FD43FF5B834BC0F0" box="[701,856,151,176]" italics="true" pageId="3" pageNumber="543">L. vietnamica</emphasis>
Masumoto, 1995
</taxonomicName>
and
<taxonomicName id="84F74D62FA77FFE2FBA0FF5B858BC0F1" authority="Masumoto, 1996" authorityName="Masumoto" authorityYear="1996" box="[1118,1432,151,177]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="7183EAF3FA77FFE2FBA0FF5B84D8C0F1" box="[1118,1227,151,177]" italics="true" pageId="3" pageNumber="543">L. hystrix</emphasis>
Masumoto, 1996
</taxonomicName>
, both from
<collectingCountry id="3BE07671FA77FFE2FEF0FF778163C095" box="[270,368,187,213]" name="Vietnam" pageId="3" pageNumber="543">Vietnam</collectingCountry>
, the general body size and shape, the beaded lateral margins of the pronotum, and the unarmed femora. However, in
<taxonomicName id="84F74D62FA77FFE2FE73FF13823BC0B8" authorityName="Masumoto" authorityYear="1995" box="[397,552,223,248]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="vietnamica">
<emphasis id="7183EAF3FA77FFE2FE73FF13823BC0B8" box="[397,552,223,248]" italics="true" pageId="3" pageNumber="543">L. vietnamica</emphasis>
</taxonomicName>
and
<taxonomicName id="84F74D62FA77FFE2FD9DFF1382DCC0B9" authorityName="Masumoto" authorityYear="1996" box="[611,719,223,249]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="543" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="7183EAF3FA77FFE2FD9DFF1382DCC0B9" box="[611,719,223,249]" italics="true" pageId="3" pageNumber="543">L. hystrix</emphasis>
</taxonomicName>
the dorsal side is without metallic luster, the punctures of the pronotum and elytra are smaller and denser, nearly all dorsal punctures bear long erect setae, and the elytral intervals are nearly flat. Both mentioned species are described without figures of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="0BED656AFA77FFE2FF39FE8780CCC1C9" pageId="3" pageNumber="543" type="etymology">
<paragraph id="434836E1FA77FFE2FF39FE8780CCC1C9" blockId="3.[151,1436,150,393]" pageId="3" pageNumber="543">
<emphasis id="7183EAF3FA77FFE2FF39FE87815DC125" bold="true" box="[199,334,331,357]" pageId="3" pageNumber="543">Etymology.</emphasis>
Named in honour of my colleague Arnaud Faille (Stuttgart), one of the collectors of the
<typeStatus id="9C4C8843FA77FFE2FA92FE87858FC125" box="[1388,1436,331,357]" pageId="3" pageNumber="543">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>
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